UBC Theses and Dissertations

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UBC Theses and Dissertations

Biomass and nutrient distribution in two old growth forest ecosystems in south coastal British Columbia Krumlik, Jiri George 1974

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BIOMASS AND N U T R I E N T D I S T R I B U T I O N I N TWO OLD GROWTH FOREST ECOSYSTEMS I N SOUTH COASTAL B R I T I S H COLUMBIA b y J I R I GEORGE K R U M L I K D i p l . I n g . , P r a g u e U n i v e r s i t y o f A g r i c u l t u r e , 1 9 6 4 A T H E S I S SUBMITTED I N P A R T I A L F U L F I L L M E N T OF THE REQUIREMENTS FOR THE DEGREE OF MASTER OF S C I E N C E I n T h e D e p a r t m e n t o f F o r e s t r y We a c c e p t t h i s t h e s i s a s c o n f o r m i n g t o t h e r e q u i r e d s t a n d a r d : THE U N I V E R S I T Y OF B R I T I S H C OLUMBIA M a y , 1 9 7 4 ( i i ) I n p r e s e n t i n g t h i s t h e s i s i n p a r t i a l f u l f i l l m e n t o f t h e r e q u i r e -m e n t s f o r a n a d v a n c e d d e g r e e a t t h e U n i v e r s i t y o f B r i t i s h C o l u m b i a , I a g r e e t h a t t h e L i b r a r y s h a l l m a ke i t f r e e l y a v a i l a b l e f o r r e f e r e n c e a n d s t u d y . I f u r t h e r a g r e e t h a t p e r m i s s i o n f o r e x t e n s i v e c o p y i n g o f t h i s t h e s i s f o r s c h o l a r l y p u r p o s e s may b e g r a n t e d b y t h e H e a d o f my D e p a r t m e n t o r b y h i s r e p r e s e n t a t i v e s . I t i s u n d e r s t o o d t h a t c o p y i n g o r p u b l i c a t i o n o f t h i s t h e s i s f o r f i n a n c i a l g a i n s h a l l n o t b e a l l o w e d w i t h o u t my w r i t t e n p e r m i s s i o n . D e p a r t m e n t o f F o r e s t r y T h e U n i v e r s i t y o f B r i t i s h C o l u m b i a V a n c o u v e r 8, C a n a d a D a t e : May 3 , 1 9 7 4 ( i i i ) A B S T R ACT T h e d i s t r i b u t i o n o f t h e a b o v e - g r o u n d b i o m a s s a n d m a c r o n u t r i e n t c o n t e n t o f t h e t r e e s w e r e s t u d i e d o n t w o s a m p l e p l o t s i n s o u t h c o a s t a l B r i t i s h C o l u m b i a , n e a r V a n c o u v e r . T h e p l o t s d i f f e r e d i n e l e v a t i o n ( 4 , 6 0 0 a n d 2 , 2 0 0 f e e t ; 1,500 a n d 7 0 0 m ) , i n s o i l t y p e a n d d e p t h . T r e e a g e w a s s i m i l a r i n b o t h p l o t s , r a n g i n g f r o m 1 5 0 t o 5 3 0 y e a r s . T h e t r e e c o v e r o n t h e h i g h e l e v a t i o n p l o t c o n s i s t e d o f T s u g a m e r t e n s i a n a ( B o n g . ) C a r r . ( m o u n -t a i n h e m l o c k ) a n d A b i e s a m a b i l i s ( D o u g l . ) F o r b e s ( P a c i f i c s i l v e r f i r ) w h i l e t h e l o w e r e l e v a t i o n p l o t w a s o c c u p i e d b y T s u g a h e t e r o p h y l l a ( R a f . ) S a r g . ( w e s t e r n h e m l o c k ) , T h u j a p l i c a t a D. D o n ( w e s t e r n r e d c e d a r ) a n d C h a m a e c y - p a r i s n o o t k a t e n s i s ( D . D o n ) S p a c h ( y e l l o w c e d a r ) . T w e n t y - f o u r t r e e s w e r e s a m p l e d t o d e t e r m i n e t h e b i o m a s s a n d n u t r i e n t c o n t e n t o f w o o d , b a r k , b r a n c h e s , t w i g s , f o l i a g e a n d c o n e s . A n o t h e r n i n e t r e e s w e r e s a m p l e d f o r t h e b i o m a s s a n d n u t r i e n t c o n t e n t o f w o o d a n d b a r k o n l y . M u l t i p l e r e g r e s s i o n a n a l y s i s w a s u s e d t o e s t a b l i s h t h e r e l a t i o n -s h i p b e t w e e n d . b . h . , t r e e l e n g t h , c r o w n l e n g t h a n d b i o m a s s o f t h e v a r i o u s t r e e c o m p o n e n t s . T h e r e g r e s s i o n e q u a t i o n s o b t a i n e d w e r e u s e d t o e s t i m a t e t h e t o t a l b i o m a s s o f w o o d , b a r k , b r a n c h e s , t w i g s a n d f o l i a g e c o n t a i n e d i n t h e t r e e s o n e a c h o f t h e s a m p l e p l o t s . T h e d a t a t h u s o b t a i n e d w e r e com-b i n e d w i t h d a t a o n c h e m i c a l c o n c e n t r a t i o n a n d u s e d t o e s t i m a t e t h e d i s t r i -b u t i o n o f m a c r o n u t r i e n t e l e m e n t s i n d i f f e r e n t a b o v e - g r o u n d b i o m a s s c o m p o n -e n t s o f t h e s t a n d s . ( i v ) T A B L E OF CONTENTS P a g e T I T L E PAGE i ABSTRACT i i i T A B L E OF CONTENTS i v L I S T OF T A B L E S , v * L I S T OF F I G U R E S , i x L I S T OF A P P E N D I C E S t X ACKNOWLEDGEMENTS , . x i I INTRODUCTION 1 O b j e c t i v e s o f t h e s i s . . . . . . . 2 I I L I T E R A T U R E REVIEW 3 N u t r i e n t u p t a k e b y t r e e s a n d n u t r i e n t i m m o b i l i z a -t i o n i n t r e e b i o m a s s . . . . . 3 N u t r i e n t d i s t r i b u t i o n i n a b o v e - g r o u n d t r e e b i o m a s s 8 R e m o v a l o f n u t r i e n t s f r o m a s i t e a s t h e r e s u l t o f t r e e h a r v e s t i n g 17 I I I D E S C R I P T I O N OF SAMPLE P L O T S 19 L o c a t i o n a n d s i z e 19 C l i m a t e . . . . . . . . . . . . 24 S o i l s 2 6 V e g e t a t i o n . . . . . . . . . 26 I V METHODS 2 9 F i e l d s a m p l i n g . . . . . . . 29 L a b o r a t o r y a n a l y s e s 3 3 ( v ) P a g e I V METHODS ( c o n t i n u e d ) D a t a a n a l y s e s . . . . . . f . . . . . . . . . . . . 3 8 L i m i t a t i o n o n d a t a i n t e r p r e t a t i o n . . . . . . . . . 47 M i s s i n g d a t a a n d a d j u s t m e n t o f i n c o r r e c t d a t a . . . 48 V R E S U L T S AND D I S C U S S I O N 5 0 B i o m a s s d i s t r i b u t i o n 5 0 N u t r i e n t d i s t r i b u t i o n w i t h i n t r e e l a y e r 57 R e m o v a l o f m a c r o n u t r i e n t s f r o m t h e e c o s y s t e m s d u e t o d i f f e r e n t l e v e l i o f t r e e u t i l i z a t i o n 73 V I SUMMARY AND CONCLUSIONS 77 ( v i ) L I S T OF T A B L E S T A B L E P a g e 1 T o t a l q u a n t i t i e s o f n u t r i e n t s i m m o b i l i z e d w i t h i n a l l o r g a n s o f t h e m a i n g r o w i n g c r o p o f v a r i o u s t r e e s t a n d s a t a g e 1 0 0 y e a r s ( R e n n i e , 1 9 5 5 ) . 4 2 T h e t o t a l q u a n t i t i e s o f n u t r i e n t s t a k e n u p b y 3 t y p e s o f t i m b e r - p r o d u c i n g f o r e s t a t a g e 1 0 0 y e a r s . ( R e n n i e , 1 9 5 5 ) . . 5 3 T h e p e r c e n t a g e d i s t r i b u t i o n o f n u t r i e n t s i m m o b i l i z e d i n t h e c o m p o n e n t o r g a n s o f t h e m a i n g r o w i n g c r o p o f t h r e e t y p e s o f s t a n d a t a g e 1 0 0 y e a r s ( R e n n i e , 1 9 5 5 ) . 9 4 C o n c e n t r a t i o n o f m a c r o n u t r i e n t s i n f o l i a g e o f u p p e r w h o r l s o f t r e e c r o w n f o r t h r e e s p e c i e s i n B. C. ( B e a t o n e t a l . , 1 9 6 5 ) . 11 5 A v e r a g e N - c o n c e n t r a t i o n s i n v a r i o u s l o b l o l l y p i n e t i s s u e s a t v a r i o u s a g e s o n g o o d s i t e s i n p e r c e n t o f o v e n - d r y w e i g h t ( S w i t z e r a n d N e l s o n , 1 9 6 4 ) . 12 6 T h e r a n g e o f n u t r i e n t c o n c e n t r a t i o n s ( i n p e r c e n t ) i n d i f f e r e n t t r e e c o m p o n e n t s i n v a r i o u s N o r w a y s p r u c e s t a n d s i n C z e c h o s l o v a k i a ( M a t e r n a , 1 9 6 4 ) . 13 7 D i s t r i b u t i o n o f N, P, K a n d C a among t h e n a j o r c o m p o n -e n t s o f a 3 6 - y e a r o l d s e c o n d - g r o w t h D o u g l a s - f i r f o r e s t ( C o l e , 1 9 6 7 ) . 14 8 D i s t r i b u t i o n o f b i o m a s s a n d n u t r i e n t s w i t h i n t h e a b o v e - g r o u n d t r e e c o m p o n e n t s o f a 1 2 - y e a r o l d s t a n d o f r a d i a t a p i n e , W h a k a r e w a r e w a F o r e s t ( W i l l , 1 9 6 4 ) . 15 9 A v e r a g e p e r c e n t a g e d i s t r i b u t i o n o f N , P, K, C a a n d Mg i n a e r i a l c o m p o n e n t s o f 12 n a t u r a l j a c k p i n e s t a n d s i n N o r t h e r n O n t a r i o ( M o r r i s o n , 1973)., 16 1 0 Summary o f c l i m a t i c i n f o r m a t i o n ( K r a j i n a , 1 9 6 5 ) . 25 < 11 D i a m e t e r c l a s s d i s t r i b u t i o n o n t h e s a m p l e p l o t s . 3 0 12 B a s i c s t a t i s t i c s o f o r i g i n a l m e a s u r e m e n t s f o r a l l t r e e s o v e r 6 i n c h e s o f d . b . h . o n t h e t w o s a m p l e p l o t s . 31 ( v i i ) T A B L E P a g e 13 P e r c e n t l o s s o f w e i g h t i n s a m p l e s f o r c h e m i c a l a n a l y s e s u p o n r e - d r y i n g a f t e r 2^-6 m o n t h s o f s t o r a g e , 34 14 C o m p a r i s o n o f r e s u l t s o f c h e m i c a l a n a l y s e s f o r f i v e r a n d o m l y s e l e c t e d s a m p l e s . 3 8 15 A v e r a g e p e r c e n t d i f f e r e n c e b e t w e e n t r e e t r u n k v o l u m e s c a l c u l a t e d a c c o r d i n g t o B C F S v o l u m e e q u a t i o n s a n d t h o s e c a l c u l a t e d s e c t i o n - b y - s e c t i o n a c c o r d i n g t o N e w t o n ' s f o r m u l a , 4 0 16 B a s i c s t a t i s t i c s f o r s a m p l e d t r e e s , 4 1 17 B i o m a s s d i s t r i b u t i o n f o r f o u r t r e e s p e c i e s o n t h e S q u a m i s h a n d H a n e y s a m p l e p l o t s ( i n k g / h a ) . 51 18 A v e r a g e b i o m a s s p r o p o r t i o n s o f d i f f e r e n t t r e e c o m -p o n e n t s f o r f o u r t r e e s p e c i e s o n t h e S q u a m i s h a n d H a n e y s a m p l e p l o t s . 5 3 19 M e a n c o n c e n t r a t i o n ( p e r c e n t a g e o v e r - d r y w e i g h t ) o f N, P, K, C a a n d Mg i n c o m p o n e n t s a n d s u b - c o m p o n e n t s o f P a c i f i c s i l v e r f i r s a m p l e - t r e e s . 5 8 2 0 M e a n c o n c e n t r a t i o n ( p e r c e n t a g e o v e r - d r y w e i g h t ) o f N , P, K, C a a n d Mg i n c o m p o n e n t s a n d s u b - c o m p o n e n t s o f m o u n t a i n h e m l o c k s a m p l e t r e e s . 5 9 21 M e a n c o n c e n t r a t i o n ( p e r c e n t a g e o v e n - d r y w e i g h t ) o f N, P, K, C a a n d Mg i n c o m p o n e n t s a n d s u b - c o m p o n e n t s o f w e s t e r n h e m l o c k s a m p l e t r e e s , 6 0 22 M e a n c o n c e n t r a t i o n ( p e r c e n t a g e o v e n - d r y w e i g h t ) o f N, P, K, C a a n d Mg i n c o m p o n e n t s a n d s u b - c o m p o n e n t s o f y e l l o w c e d a r s a m p l e t r e e s , 6 1 23 C o n c e n t r a t i o n o f m a c r o n u t r i e n t s i n f o l i a g e w i t h i n u p p e r t h i r d o f t r e e c r o w n , 65 24 D i s t r i b u t i o n o f m a c r o n u t r i e n t s b y p a r t i c u l a r t r e e c o m p o n e n t s o n t h e S q u a m i s h s a m p l e p l o t . 67 25 D i s t r i b u t i o n o f m a c r o n u t r i e n t s b y p a r t i c u l a r t r e e c o m p o n e n t s o n t h e H a n e y s a m p l e p l o t . 6 8 ( v i i i ) T A B L E P a g e 26 A v e r a g e p r o p o r t i o n s o f f i y e m a c r o n u t r i e n t s i n d i f V f e r e n t t r e e c o m p o n e n t s o n t h e S q u a m i s h a n d H a n e y s a m p l e p l o t s . 69 27 T h e a m o u n t o f m a c r o n u t r i e n t s i m m o b i l i z e d i n a b o v e -g r o u n d t r e e b i o m a s s b y v a r i o u s s p e c i e s i n d i f f e r e n t c o u n t r i e s . 72 28 R e m o v a l o f b i o m a s s a n d m a c r o n u t r i e n t s a c c o m p a n y i n g d i f f e r e n t l e v e l s o f u t i l i z a t i o n i n t h e S q u a m i s h s a m p l e p l o t . 74 29 R e m o v a l o f b i o m a s s a n d m a c r o n u t r i e n t s a c c o m p a n y i n g d i f f e r e n t l e v e l s o f u t i l i z a t i o n i n t h e H a n e y s a m p l e p l o t . 75 L I S T OF F I G U R E S F I G U R E P a g e V a n c o u v e r a n d v i c i n i t y - l o c a t i o n o f t h e s a m p l e p l o t s . 20 L I S T OF A P P E N D I C E S P a g e S c i e n t i f i c a n d common nam e s o f t r e e s p e c i e s . 3 8 B a s i c s t a t i s t i c s f o r t h e i n d e p e n d e n t v a r i a b l e s . 90 S i g n i f i c a n t i n d e p e n d e n t v a r i a b l e s i n s i m p l e l i n e a r a n d l o g a r i t h m i c r e g r e s s i o n e q u a t i o n s f o r b i o m a s s p r e d i c t i o n o f p a r t i c u l a r t r e e c o m p o n e n t s . B i o m a s s d i s t r i b u t i o n f o r 2 0 s a m p l e t r e e s o n t h e S q u a m i s h s a m p l e p l o t , B i o m a s s d i s t r i b u t i o n f o r 13 s a m p l e t r e e s o n t h e H a n e y s a m p l e p l o t . 92 99 110 N u t r i e n t c o n c e n t r a t i o n a n d n u t r i e n t d i s t r i b u t i o n w i t h i n 2 0 s a m p l e t r e e s o n t h e S q u a m i s h s a m p l e p l o t . 1 1 8 N u t r i e n t c o n c e n t r a t i o n a n d n u t r i e n t d i s t r i b u t i o n w i t h i n 13 s a m p l e t r e e s o n t h e H a n e y s a m p l e p l o t . 139 P r o g r a m f o r c a l c u l a t i n g m o i s t u r e c o n t e n t o f s a m p l e s f o r c h e m i c a l a n a l y s e s . 1 5 3 S p e c i f i c g r a v i t y p r o g r a m . 157 P r o g r a m f o r c a l c u l a t i n g b i o m a s s d i s t r i b u t i o n f o r s a m p l e d t r e e s . 1 5 9 P r o g r a m f o r c a l c u l a t i n g n u t r i e n t d i s t r i b u t i o n . 1 6 3 171 P r o g r a m f o r c a l c u l a t i n g w e i g h t e d c o n c e n t r a t i o n s o f n u t r i e n t s w i t h i n d i f f e r e n t c o m p o n e n t s o f s a m p l e t r e e s . P r o g r a m f o r c a l c u l a t i n g b i o m a s s f o r 2 5 6 t r e e s o n t h e S q u a m i s h s a m p l e p l o t a c c o r d i n g t o l o g a r i t h m i c r e g r e s s i o n e q u a t i o n s . 1 7 7 ( x i ) ACKNOWLEDGEMENTS T h e w r i t e r w i s h e s t o e x p r e s s h i s s i n c e r e t h a n k s t o D r . J . P. K i m m i n s f o r p r o v i d i n g t h e o p p o r t u n i t y t o u n d e r t a k e t h i s p r o j e c t a n d f o r h i s g u i d a n c e , a d v i c e a n d c r i t i c a l r e v i e w ; t o D r . T. B a l l a r d f o r n u m e r o u s h o u r s o f d i s c u s s i o n , h e l p w i t h d a t a i n t e r p r e t a t i o n a n d d r a w i n g c o n c l u s i o n s ; t o D r s . A. K o z a k a n d J . D e m a e r s c h a l k f o r h e l p w i t h s t a t i s t i c a l a n a l y s i s ; t o D r s . P. H a d d o c k a n d J . W o r r a l l f o r c r i t i c a l r e v i e w o f t h e m a n u s c r i p t ; t o D r . R. C. B r o o k e o f S i m o n F r a s e r U n i v e r s i t y a n d M r . K a r e l K l i n k a f o r h e l p i n s i t e i d e n t i f i c a t i o n a n d s e l e c t i o n ; t o M e s s r s . J o h n P a r m i n t e r a n d G r a n t S c h e i r m a n f o r h e l p w i t h f i e l d w o r k ; t o M r . A u s t i n P h i r i f o r s o i l s a m p l i n g a n d l a b o r a t o r y a n a l y s e s ; t o M r . K e n T w a i t e s f o r h e l p w i t h l a b o r a -t o r y a n a l y s e s a n d c o m p u t e r p r o g r a m m i n g ; t o M r . M i n T s z e f o r h e l p w i t h l a b o r a t o r y a n a l y s e s ; t o M i s s L . C o w d e l l f o r h e l p w i t h c o m p u t e r p r o g r a m m i n g ; t o M e s s r s . D o n G r a n t a n d T a g N e i g h b o u r o f t h e B C F S f o r h e l p i n s i t e l o c a -t i o n a n d o t h e r p r a c t i c a l d e t a i l s ; t o t h e s t a f f o f t h e UBC R e s e a r c h F o r e s t a t H a n e y ; t o M a c M i l l a n B l o e d e l l , S q u a m i s h O f f i c e , f o r c o o p e r a t i o n i n a c c e s s t o t h e h i g h e l e v a t i o n s i t e a n d h e l p i n f a l l i n g t r e e s . T h e h e l p o f t h e s e p e o p l e a n d o r g a n i z a t i o n s , t o g e t h e r w i t h f i n a n -c i a l s u p p o r t o f t h e B C F S P r o d u c t i v i t y C o m m i t t e e made t h i s t h e s i s p o s s i b l e . 1 I INTRODUCTION M u c h o f t h e l o g g i n g i n c o a s t a l B r i t i s h C o l u m b i a i n t h e p a s t w a s i n t h e m o r e a c c e s s i b l e l o w e l e v a t i o n s i t e s . M o r e r e c e n t l y , t h e l o g g e r ' s a t t e n t i o n h a s b e e n t u r n e d t o w a r d s h i g h e r e l e v a t i o n f o r e s t s , a n d b o t h o n t h e c o a s t a n d i n t h e i n t e r i o r , s u b a l p i n e f o r e s t s a r e b e i n g c u t o n a n i n -c r e a s i n g s c a l e . B e c a u s e o f t h e i r r e l a t i v e i n a c c e s s i b i l i t y a n d t h e i r r e l a -t i v e l y r e c e n t u t i l i z a t i o n , m u c h l e s s i s k n o w n a b o u t t h e m t h a n t h e l o w e r e l e v a t i o n f o r e s t s . L i t t l e i s k n o w n a b o u t t h e i r s o i l s , t h e i r h u m u s , t h e i r c l i m a t e , t h e i r r e s p o n s e t o h a r v e s t i n g , t h e s u s t a i n a b i l i t y o f t h e f o r e s t s , t h e s u i t a b i l i t y o f t h e c l e a r c u t h a r v e s t i n g m e t h o d s , a n d t h e s o l u t i o n t o t h e p r o b l e m s o f r e g e n e r a t i n g h i g h - e l e v a t i o n c u t o v e r a r e a s . S i n c e s u b a l p i n e f o r e s t s r e p r e s e n t a v e r y s u b s t a n t i a l a r e a o f c o a s t a l f o r e s t s ( 1 1 . 3 m i l l i o n a c r e s c o m p a r e d w i t h 2 6 . 3 m i l l i o n o f o t h e r f o r e s t t y p e s o n t h e c o a s t ( S m i t h , 1 9 7 4 ) ) t h e r e i s a n u r g e n t n e e d f o r a l a r g e v a r i e t y o f s t u d i e s t o b e u n d e r -t a k e n i n t h e s e f o r e s t s a s a f o u n d a t i o n f o r t h e i r r a t i o n a l m a n a g e m e n t . A l t h o u g h r e l a t i v e l y l i t t l e i s k n o w n a b o u t t h e s e f o r e s t s , i n t e r e s t i n t h e m h a s g r o w n o v e r t h e p a s t d e c a d e . A s u b s t a n t i a l c o n t r i b u t i o n t o o u r u n d e r s t a n d i n g o f t h e e c o l o g y o f t h e c o a s t a l s u b a l p i n e f o r e s t e x i s t s i n t h e f o r m o f t h e s t u d y b y B r o o k e , P e t e r s o n a n d K r a j i n a ( 1 9 7 0 ) , T h e B r i t i s h C o l u m b i a F o r e s t S e r v i c e h a s u n d e r -t a k e n a r e v i e w o f h i g h e l e v a t i o n r e g e n e r a t i o n p r o b l e m s , a n d i s c u r r e n t l y i n v e s t i g a t i n g a n u m b e r o f a s p e c t s o f h i g h e l e v a t i o n l o g g i n g . T h e w o r k r e -p o r t e d h e r e i n w a s f u n d e d b y t h e B, C, F o r e s t S e r v i c e P r o d u c t i v i t y C o m m i t t e e 2 a n d r e f l e c t s i t s c o n c e r n f o r a b e t t e r u n d e r s t a n d i n g o f t h e e c o l o g i c a l a n d p r a c t i c a l a s p e c t s o f h i g h - e l e v a t i o n c o a s t a l l o g g i n g . T h i s s t u d y w a s a p a r t o f a n i n v e s t i g a t i o n o f t h e p o t e n t i a l f o r s i t e d e g r a d a t i o n t h r o u g h n u t r i e n t d e p l e t i o n b y h a r v e s t i n g . T o i n v e s t i g a t e t h e f u l l s i g n i f i c a n c e o f n u t r i e n t l o s s e s a c c o m p a n y i n g f o r e s t h a r v e s t i n g i s b e y o n d t h e s c o p e o f t h i s s t u d y . T h i s t h e s i s w a s r e s t r i c t e d t o t h e f o l l o w i n g o b j e c t i v e s : 1. T o q u a n t i f y t h e a b o v e - g r o u n d t r e e b i o m a s s b y d i f f e r e n t s t e m a n d c r o w n b i o m a s s c o m p o n e n t s . 2. T o q u a n t i f y t h e m a c r o n u t r i e n t ( N , P, K, C a , Mg) c o n t e n t o f t h e s e b i o m a s s c o m p o n e n t s . 3. To c a l c u l a t e t h e q u a n t i t y o f n u t r i e n t s r e m o v e d f r o m t h e s i t e i n h a r v e s t e d m a t e r i a l s a t d i f f e r e n t l e v e l s o f u t i l i z a t i o n . T h e s t u d y t h u s a d o p t s t h e s o - c a l l e d " s u b s t i t u t i o n p r i n c i p l e " (Tamm, 1 9 6 4 ) m e t h o d o f a s s e s s i n g s i t e n u t r i e n t d e p l e t i o n a c c o m p a n y i n g t h e r e m o v a l o f f o r e s t p r o d u c t s . 3 I I L I T E R A T U R E REVIEW T h e r e v i e w e f t h e l i t e r a t u r e h a s b e e n o r g a n i z e d i n t o t h r e e m a j o r t o p i c s : 1. N u t r i e n t u p t a k e b y t r e e s a n d n u t r i e n t i m m o b i l i z a t i o n i n t r e e b i o m a s s . 2. N u t r i e n t d i s t r i b u t i o n i n a b o v e - g r o u n d t r e e b i o m a s s . 3. R e m o v a l o f n u t r i e n t s f r o m a s i t e a s t h e r e s u l t o f h a r -v e s t i n g . 2 . 1 : N u t r i e n t u p t a k e b y t r e e s a n d n u t r i e n t i m m o b i l i z a t i o n i n t r e e  b i o m a s s . S i n c e t h e m i d d l e o f t h e l a s t c e n t u r y t h e r e h a v e b e e n s e v e r a l a t -t e m p t s t o e s t i m a t e t h e a m o u n t o f n u t r i e n t s r e q u i r e d f o r o p t i m a l t r e e g r o w t h ( E b e r m a y e r , 1 8 7 6 ) . M o r e r e c e n t l y e f f o r t s h a v e b e e n made t o d e t e r m i n e t h e t o t a l n u t r i e n t d e m a n d o n a f o r e s t s i t e r e s u l t i n g f r o m n e t n u t r i e n t u p t a k e b y t r e e s w h i c h a r e s u b s e q u e n t l y h a r v e s t e d . A v e r y c o m p r e h e n s i v e r e v i e w o f t h e l i t e r a t u r e o n t h i s t o p i c , i n c l u d i n g n u t r i e n t u p t a k e a n d i m m o b i l i z a t i o n , w a s made b y R e n n i e ( 1 9 5 5 ) . He r e v i e w e d p u b l i c a t i o n s f r o m t h e s e c o n d h a l f o f t h e l a s t c e n t u r y a n d f r o m t h e f i r s t h a l f o f t h i s c e n t u r y . B a s e d u p o n t h i s l i t e r a t u r e t h e t o t a l q u a n t i t i e s o f n u t r i e n t s t a k e n u p a n d i m m o b i l i z e d w i t h i n a l l o r g a n s o f t h e m a i n g r o w i n g c r o p o f t h r e e t y p e s o f f o r e s t s t a n d w e r e e s t i m a t e d a n d a r e p r e s e n t e d i n T a b l e 1. 4 T A B L E 1: T o t a l q u a n t i t i e s o f n u t r i e n t s i m m o b i l i z e d w i t h i n a l l o r g a n s o f t h e m a i n g r o w i n g c r o p o f v a r i o u s t r e e s t a n d s a t a g e 1 0 0 y e a r s ( R e n r i i e , 1 9 5 5 ) . T o t a l n u t r i e n t s i m m o b i l i z e d w i t h i n t h e m a i n g r o w i n g c r o p , k g e l e m e n t p e r a c r e . S p e c i e s C r o p A f t e r 5 0 y e a r s ' g r o w t h A f t e r 1 0 0 y e a r s ' g r o w t h C a K P C a K P P i n e s 98 5 7 1 3 1 1 5 5 6 1 2 O t h e r c o n i f e r s 2 0 8 1 1 3 2 5 2 7 4 1 5 2 28 H a r d w o o d s 3 0 2 8 2 19 5 2 0 1 2 9 2 8 C o m m e n t i n g o n t h e v a l i d i t y o f t h e s e n u t r i e n t e s t i m a t e s , R e n n i e s t a t e d : " P a r t i c u l a r e s t i m a t e s m i g h t s h o w a p p r e c i a b l e c u m u l a t i v e e r r o r . B u t f o r many i n d i v i d u a l e s t i m a t e s a n d f o r t h e m e a n e s t i m a t e s r e p r e s e n t i n g t h e t h r e e s p e c i e s g r o u p s t h e v a r i a t i o n i n u p t a k e s e e n w o u l d , i n t h e m a i n , b e g e n u i n e a n d d u e t o d i f f e r e n c e s i n s p e c i e s g r o w t h - r a t e a n d s i t e . " T h u s , a l -t h o u g h R e n n i e ' s c o n c l u s i o n s w e r e b a s e d o n o l d a n d s c a t t e r e d d a t a , h e b e -l i e v e d t h e y w e r e a t l e a s t a p p r o x i m a t e l y c o r r e c t . C o n s i d e r a b l e q u a n t i t i e s o f n u t r i e n t s , p a r t i c u l a r l y C a , may b e t a k e n u p a n d i m m o b i l i z e d w i t h i n t h e v a r i o u s t r e e o r g a n s o f a g r o w i n g f o r e s t . T h e r e l a t i v e l y s m a l l i n c r e a s e i n n u t r i e n t i m m o b i l i z a t i o n f r e q u e n t l y o c c u r -r i n g f r o m 5 0 t o 1 0 0 y e a r s ( s e e T a b l e 1 ) s u g g e s t s t h a t n u t r i e n t i m m o b i l i z a -t i o n may r e a c h a n u p p e r l i m i t a n d t h a t t h e s o l e n u t r i e n t m o v e m e n t t h e r e -a f t e r i s a c o m p a r a t i v e l y s m a l l n u t r i e n t c y c l e a s s o c i a t e d w i t h f o l i a r f l u s h a n d f a l l . T h e same c o n c l u s i o n w a s r e a c h e d b y S w i t z e r a n d N e l s o n ( 1 9 6 4 ) . 5 T h e y f o u n d t h a t t h e r a t e s o f u p t a k e o f a l l n u t r i e n t s a r e g r e a t e s t d u r i n g t h e e a r l y l i f e o f t h e s t a n d a n d t h a t b y a g e 4 5 y e a r s n u t r i t i o n i s m o s t l y a m a t t e r o f c y c l i n g . T a b l e 1 o n l y r e f e r s t o n u t r i e n t s i m m o b i l i z e d i n t h e f i n a l c r o p , h o w e v e r . I f we i n c l u d e t h e a m o u n t o f n u t r i e n t s r e m o v e d i n t h i n n i n g s , t h e t o t a l a m o u n t o f n u t r i e n t s i m m o b i l i z e d o v e r t h e r o t a t i o n i n c r e a s e s c o n s i d e r -a b l y . T h i s c a n b e s e e n b y a c o m p a r i s o n b e t w e e n T a b l e 1 a n d T a b l e 2, T h e l a t t e r s h o w s t h e t o t a l q u a n t i t i e s o f n u t r i e n t s t a k e n u p b y t h e s t a n d t y p e s r e f e r r e d t o i n T a b l e 1, T h u s , t h e q u a n t i t y o f n u t r i e n t s r e m o v e d i n h a r -v e s t i n g i s l i k e l y t o i n c r e a s e a s u t i l i z a t i o n s t a n d a r d s a n d i n t e n s i t y o f m a n a g e m e n t i n c r e a s e , a n d r o t a t i o n l e n g t h d e c r e a s e . T A B L E 2: T h e t o t a l q u a n t i t i e s o f n u t r i e n t s t a k e n u p b y 3 t y p e s o f t i m b e r -p r o d u c i n g f o r e s t a t a g e 1 0 0 y e a r s ( R e n n i e , 1 9 5 5 ) . T o t a l n u t r i e n t u p t a k e ( e x c l . f o l i a g e a n d r o o t s o f t h i n n i n g s a n d l i t t e r ) , k g . e l e m e n t p e r a c r e . S p e c i e s C r o p C o m p o n e n t A f t e r 5 0 y e a r s ' g r o w t h A f t e r 1 0 0 y e a r s ' g r o w t l C a K P C a K P t i m b e r t h i n n i n g s 35 15 4 88 3 5 9 P i n e s m a i n c r o p 98 57 1 3 1 1 5 56 12 t o t a l 1 3 3 72 17 2 0 3 9 1 2 1 t i m b e r O t h e r t h i n n i n g s 56 26 5 1 6 4 8 2 1 3 c o n i f e r s m a i n c r o p 2 0 8 1 1 3 25 2 7 4 1 5 2 28 t o t a l 2 6 4 1 3 9 3 0 4 3 8 2 3 4 4 1 t i m b e r t h i n n i n g s 1 6 1 44 1 0 3 5 9 96 22 H a r d w o o d s m a i n c r o p 3 0 2 82 1 9 5 2 0 1 2 9 2 8 t o t a l 4 6 3 1 2 6 29 8 7 9 2 2 5 5 0 6 T h e r e a r e many o t h e r s t u d i e s i n w h i c h n u t r i e n t u p t a k e a n d i m -m o b i l i z a t i o n o f n u t r i e n t s b y f o r e s t h a v e b e e n d o c u m e n t e d . V i r o ( 1 9 5 6 ) c a l c u l a t e d t h e a v e r a g e a n n u a l u p t a k e f o r v a r i o u s p i n e s t a n d s i n F i n l a n d a v e r a g i n g a b o u t 1 0 0 y e a r s i n a g e t o b e 7.36 k g o f N, 0.58 k g o f P, 0.60 k g o f K, 8.4 k g o f C a a n d 0.87 k g o f Mg p e r h e c t a r e . T h e g r e a t e r p a r t o f t h e s e n u t r i e n t s i s r e t u r n e d t o t h e f o r e s t f l o o r i n t h e f o r m o f l i t t e r a n d t h r o u g h f a l l , h o w e v e r , a n d o n l y p a r t o f t h e t o t a l u p t a k e i s i m m o b i l i z e d i n t h e t r e e b i o m a s s . W r i g h t a n d W i l l ( 1 9 5 8 ) c a l c u l a t e d t h e a m o u n t o f n u t r i e n t s b e i n g i m m o b i l i z e d i n a 6 4 - y e a r o l d s t a n d o f S c o t s p i n e . * T o t a l d r y w e i g h t o f t h e b i o m a s s w a s 1 1 9 , 0 0 0 k g / h a , c o n t a i n i n g 2 0 5 k g o f N, 2 3 k g o f P, 1 0 0 k g o f K, 2 0 2 k g o f C a a n d 3 5 k g o f Mg p e r h e c t a r e . W i l l ( 1 9 6 4 ) . c a l c u l a t e d t h e t o t a l n u t r i e n t r e m o v a l d u e t o t h i n n i n g a n d c l e a r - f e l l i n g f o r a 3 5 - y e a r o l d s t a n d o f r a d i a t a p i n e . T h e a m o u n t o f d r y b i o m a s s h a r v e s t e d w a s 3 6 2 , 0 0 0 k g / h a c o n t a i n i n g 2 3 0 k g o f N, 3 2 k g o f P, 2 8 3 k g o f K a n d 1 8 8 k g o f C a p e r h e c t a r e . I n c o n t r a s t , a 3 5 - y e a r o l d s t a n d o f C o r s i c a n p i n e i n E n g l a n d ( C u l b i n f o r e s t , M o r a y s h i r e ) w a s f o u n d t o c o n -t a i n c o n s i d e r a b l y m o r e n u t r i e n t s t h a n t h i s i n s p i t e o f i t s s m a l l e r a m o u n t o f d r y b i o m a s s ( C r a i g a n d M i l l e r , 1 9 6 6 ) . T h e c a l c u l a t e d a m o u n t o f d r y b i o -m a s s w a s o n l y 1 9 9 , 5 0 0 k g / h a b u t c o n t a i n e d 5 8 6 k g o f N, 75 k g o f P, 2 0 7 k g o f K, 2 3 0 k g o f C a a n d 9 3 k g o f Mg p e r h e c t a r e . * L a t i n n a m e s o f t r e e s p e c i e s a r e p r e s e n t e d i n A p p e n d i x 1. 7 T h e n u t r i e n t a c c u m u l a t i o n w i t h i n a l o b l o l l y p i n e p l a n t a t i o n i n s o u t h - e a s t e r n U.S.A. w a s s t u d i e d b y S w i t z e r a n d N e l s o n ( 1 9 7 2 ) . A 2 0 - y e a r o l d p l a n t a t i o n w a s f o u n d t o c o n t a i n 1 7 4 k g o f N, 1 9 k g o f P, 9 8 k g o f K, 9 0 k g o f C a a n d 23 k g o f Mg p e r h e c t a r e . I n s t u d y i n g a 9 3 - y e a r o l d s p r u c e f o r e s t i n R u s s i a , S m i r n o v a ( 1 9 5 1 ) f o u n d 4 6 8 k g o f N, 85 k g o f P, 2 7 8 k g o f K, 5 5 2 k g o f C a a n d 9 0 k g o f Mg p e r h e c t a r e i n a n u n d e f i n e d q u a n t i t y o f b i o m a s s . T h e d i s t r i b u t i o n o f n u t r i e n t s w i t h i n a 3 6 - y e a r o l d s e c o n d * - g r o w t h D o u g l a s - f i r e c o s y s t e m w a s s t u d i e d b y C o l e e t a l , ( 1 9 6 7 ) . T h e d r y w e i g h t o f t r e e o r g a n i c m a t t e r ( i n c l u d i n g r o o t s ) w a s 2 0 4 , 5 2 9 k g / h a c o n t a i n i n g 3 2 0 k g o f N, 66 k g o f P, 2 2 0 k g o f K, 3 3 3 k g o f C a p e r h e c t a r e . T h e a n n u a l u p -t a k e o f t h e f o r e s t s t a n d w a s : 3 8 . 8 , 7 , 2 , 2 9 . 4 , 2 4 . 4 k g p e r h e c t a r e f o r n i t r o g e n , p h o s p h o r o u s , p o t a s s i u m a n d c a l c i u m , r e s p e c t i v e l y . M o r r i s o n ( 1 9 7 2 ) m e a s u r e d t h e d i s t r i b u t i o n o f b i o m a s s a n d m a c r o -n u t r i e n t s o f t w e l v e j a c k p i n e s t a n d s i n N o r t h e r n O n t a r i o . T h e a v e r a g e a m o u n t o f d r y a b o v e - g r o u n d o r g a n i c m a t t e r w a s 1 0 5 , 7 8 3 k g / h a . ( S t a n d s 6 5 y e a r s o l d , s i t e c l a s s I , h i g h s t o c k i n g d e n s i t y . ) T h e a m o u n t o f n u t r i e n t s i m m o b i l i z e d w i t h i n t h e a b o v e - g r o u n d t r e e b i o m a s s w a s 1 6 1 . 1 , 1 2 . 5 , 8 1 . 0 , 1 1 5 . 6 a n d 1 7 . 5 k g p e r h e c t a r e o f n i t r o g e n , p h o s p h o r o u s , p o t a s s i u m , c a l c i u m a n d m a g n e s i u m , r e s p e c t i v e l y . L j a m e b o r s a j ( 1 9 6 8 ) s t u d i e d i m m o b i l i z a t i o n o f n i t r o g e n a n d a s h e l e m e n t s i n a n u m b e r o f t r e e s p e c i e s . I n v e s t i g a t i o n s o f t h e b i o m a s s w e r e 8 made o n 3 0 0 s a m p l e t r e e s o n 35 p l o t s i n s t a n d s o f 7 s p e c i e s a t t h e a g e o f m aximum c u r r e n t a n n u a l i n c r e m e n t . F r o m t h e d a t a o b t a i n e d i t w a s c a l c u l a t e d t h a t S c o t s p i n e r e q u i r e d 1.7 k g o f N , 0.6 k g o f P, 1.1 k g o f K a n d 1.9 k g o f C a t o p r o d u c e o n e c u b i c m e t e r o f m e r c h a n t a b l e w o o d ; c o r r e s p o n d i n g f i g -u r e s f o r N o r w a y s p r u c e w e r e 2 . 1 , 1 . 1 , 1.8 a n d 2.5 k g ; f o r b i r c h 3 . 8 , 1 . 0 , 1.5 a n d 2.8 k g ; f o r o a k 6.7, 1.4, 4.4 a n d 1 3 . 4 k g ; f o r a s p e n 3 . 2 , 0 . 8 , 1.8 a n d 5.3 k g ; f o r l i m e 5 . 9 , 0 . 8 , 3.4 a n d 7.2 k g ; a n d f o r l a r c h 1 . 3 , 1 . 1 , 0.8 a n d 8.1 k g ; r e s p e c t i v e l y . T h e a m o u n t o f n u t r i e n t s i m m o b i l i z e d w i t h i n t h e t r e e s t a n d d e p e n d s m a i n l y o n t h e w e i g h t o f d r y t r e e b i o m a s s a n d s e c o n d l y o n t h e c o n c e n t r a t i o n o f n u t r i e n t s i n p a r t i c u l a r t r e e s p e c i e s . T h e w e i g h t o f i m m o b i l i z e d e l e m e n t s a c c o r d i n g t o t h e a b o v e - p r e s e n t e d l i t e r a t u r e c a n b e a s h i g h a s 5 8 0 k g N, 85 k g P, 2 8 0 k g K, 5 5 0 k g C a a n d 9 0 k g Mg p e r h e c t a r e . T h e a n n u a l u p t a k e o f n u t r i e n t s d u r i n g t h e e a r l y s t a g e o f s t a n d g r o w t h ( f i r s t 3 0 - 4 0 y e a r s ) c a n b e a s h i g h a s 4 0 k g N , 8 k g P, 3 0 k g K a n d 25 k g C a p e r h e c t a r e ( C o l e e t a l . , 1 9 6 7 ; W i l l 1 9 6 4 ) . 2 , 2 : N u t r i e n t d i s t r i b u t i o n i n a b o v e - g r o u n d t r e e b i o m a s s . T h e d i s t r i b u t i o n o f n u t r i e n t s among t h e c o m p o n e n t o r g a n s o f t h e m a i n g r o w i n g c r o p ( T a b l e 1 ) i s s h o w n i n T a b l e 3. T h e p e r c e n t a g e d i s t r i b u t i o n s o f n u t r i e n t s among t h e f i v e c o m p o n -e n t s f o r e a c h s p e c i e s g r o u p r e p r e s e n t m e a n s o f t h e p e r c e n t a g e d i s t r i b u t i o n s o f t h e s e v e r a l s p e c i e s w i t h i n t h a t g r o u p . T h e s e d a t a a r e i n a g r e e m e n t w i t h 9 T A B L E 3: T h e p e r c e n t a g e d i s t r i b u t i o n o f n u t r i e n t s i m m o b i l i z e d i n t h e c o m -p o n e n t o r g a n s o f t h e m a i n g r o w i n g c r o p o f t h r e e t y p e s o f s t a n d a t a g e 1 0 0 y e a r s ( R e n n i e , 1 9 5 5 ) . M a i n M e a n s o f t h e p e r c e n t a g e d i s t r i b u t i o n o f n u t r i -G r o w i n g e n t s a mong t h e m a i n c r o p c o m p o n e n t s S p e c i e s G r o u p C r o p C o m p o n e n t A f t e r 5 0 y e a r s ' g r o w t h A f t e r 1 0 0 y e a r s ' g r o w t h C a K P C a K P n e e d l e s 2 3 3 9 4 3 14 2 8 32 b r u s h 25 1 8 2 1 1 8 14 1 8 P i n e s s t e m b a r k 17 8 9 2 5 1 2 14 s t e m w o o d 22 22 1 4 3 0 3 3 2 3 r o o t s 1 3 1 3 1 3 1 3 1 3 13 n e e d l e s 26 3 0 4 0 1 5 17 27 b r u s h 3 0 27 25 2 1 2 0 2 2 O t h e r s t e m b a r k 2 0 14 1 3 3 2 23 2 3 c o n i f e r s s t e m w o o d 1 1 16 9 1 9 27 1 5 r o o t s 1 3 1 3 1 3 13 1 3 13 l e a v e s 1 1 2 0 22 5 1 0 1 2 b r u s h 22 2 5 28 16 2 1 24 H a r d w o o d s * s t e m b a r k 3 8 14 17 46 18 2 3 s t e m w o o d 16 2 8 2 0 2 0 3 8 28 r o o t s 1 3 1 3 1 3 1 3 13 1 3 * e x c l u d i n g t h o s e w i t h o u t l e a v e s s a m p l e d i n w i n t e r . d a t a r e v i e w e d b y O v i n g t o n ( 1 9 6 2 ) f o r a l a r g e n u m b e r o f e c o s y s t e m s i n d i f -f e r e n t p a r t s o f t h e w o r l d . D u v i g n e a u d a n d D e n a e y e r - D e S m e t ( 1 9 6 4 ) a n a l y s e d t h e v a r i o u s o r g a n s o f t h r e e d e c i d u o u s s p e c i e s a n d f o u n d t h e f o l l o w i n g s e -q u e n c e w h e n r e s u l t s a r e e x p r e s s e d a s t h e c o n t e n t o f e a c h n u t r i e n t i n p e r -c e n t a g e o f t h e t o t a l t r e e c o n t e n t o f t h a t e l e m e n t : 1 0 N - f o l i a g e > b o l e w o o d > b o l e b a r k = b r a n c h b a r k > -b r a n c h w o o d t w i g s C a - b o l e b a r k > b r a n c h b a r k > f o l i a g e > b o l e w o o d t w i g s > b r a n c h w o o d K - f o l i a g e > b o l e w o o d > b r a n c h w o o d ; > t w i g s > b o l e b a r k = b r a n c h b a r k S l i g h t c h a n g e s i n t h i s s e q u e n c e w e r e n o t e d a mong some s p e c i e s d u e t o i n -h e r e n t m o r p h o l o g i c a l d i f f e r e n c e s . F o r e x a m p l e , r e p r o d u c t i v e o r g a n s a r e n o t a b l y r i c h i n some e l e m e n t s . D u v i g n e a u d a n d D e n a e y e r - D e S m e t s u m m a r i z e d t h e l i t e r a t u r e o n m i n e r a l c o n t e n t o f p o l l e n a n d e s t a b l i s h e d t h a t t h e e l e -m e n t s a r e p r e s e n t i n t h e o r d e r o f N > K J > P > - C a o n a p e r c e n t a g e b a s i s . T h e c o n c e n t r a t i o n s o f N, P, K, C a , Mg a n d S i n n e e d l e s o f P a c i f i c s i l v e r f i r , a l p i n e f i r , w e s t e r n r e d c e d a r , D o u g l a s - f i r , w e s t e r n h e m l o c k , l o d g e p o l e p i n e , E n g e l m a n n s p r u c e a n d S i t k a s p r u c e o f d i f f e r e n t a g e s a n d l o c a t i o n s i n B r i t i s h C o l u m b i a w e r e s t u d i e d b y B e a t o n e_t a l . ( 1 9 6 5 ) . H i s r e s u l t s f o r w e s t e r n h e m l o c k , a m a b i l i s f i r a n d r e d c e d a r a r e s u m m a r i z e d i n T a b l e 4. N u t r i e n t c o n c e n t r a t i o n s i n f o l i a g e o f D o u g l a s - f i r i n W a s h i n g t o n S t a t e w e r e s t u d i e d b y G e s s e l eX a l . ( 1 9 6 0 ) f o r s t a n d s o f d i f f e r e n t g r o w t h v i g o r . H i s r e s u l t s c a n b e s u m m a r i z e d : N - 0.6 t o 2.3 p e r c e n t ; P - 0.1 t o 0.25 p e r c e n t ; K - 0.3 t o 1.0 p e r c e n t : C a - 0.2 t o 0.75 p e r c e n t , a n d 1 1 T A B L E 4: C o n c e n t r a t i o n o f m a c r o n u t r i e n t s i n f o l i a g e o f t h e u p p e r w h o r l s o f t r e e c r o w n f o r t h r e e t r e e s p e c i e s i n B. C. ( B e a t o n e t •. ml.'» 1 9 6 5 ) . T y p e , l o c a t i o n D a t e N o . o f Y e a r P l a n t n u t r i e n t c o n c e n t r a t i o n & a g e o f s t a n d o f T r e e s o f p e r c e n t i n d r y m a t t e r S a m p l e S a m p l e d G r o w t h N P K C a Mg A m a b i l i s f i r A u g u s t 5 1 9 6 3 0.99 0.16 1.20 0.30 0.08 T e r r a c e ( 8 y r . ) 1 9 6 3 1 9 6 2 0 . 9 1 0.12 1.10 0 . 4 1 0.07 W e s t e r n r e d A u g u s t c e d a r , T e r r a c e 1 9 6 3 5 1 9 6 3 0 .73 0.13 0.52 1.16 0.10 ( 5 8 y e a r s ) W e s t e r n h e m l o c k F e b . 1 9 6 3 1.17 0.16 0.43 0.20 0.08 Q u a d r a I s l a n d 1 9 6 4 1 2 1 9 6 2 1 . 2 1 0.18 0.40 0.24 0.10 ( 6 0 y e a r s ) 1 9 6 1 1.22 0.18 0.38 0.30 0.08 G o l d R i v e r A p r i l ( 1 4 0 - 1 5 0 y e a r s ) 1 9 6 4 5 1 9 6 3 0 . 9 1 0 . 1 1 0.28 0.18 0.13 T e r r a c e ( 8 , 5 7 , A u g u s t 1 9 6 3 0.86 0.19 0.57 0.27 0.12 7 3 , 1 1 5 , 1 5 3 , 1 9 6 3 2 0 1 9 6 2 0.84 0 . 2 1 0 . 5 1 0.32 0.14 1 8 5 y e a r s ) 1 9 6 1 0.88 0.22 0.47 0.35 0.14 Mg - 0.05 t o 0.15 p e r c e n t . T h e i m p o r t a n c e o f h a v i n g a c o r r e c t b a l a n c e b e t w e e n n u t r i e n t s i n t h e f o l i a g e a n d e s p e c i a l l y o f t h e r a t i o s N /P, N/K, a r i d K/P h a s b e e n s t r e s -s e d b y a n u m b e r o f i n v e s t i g a t o r s . F o r e x a m p l e , L e y t o n ( 1 9 5 7 ) c a l c u l a t e d N/P r a t i o s f o r J a p a n e s e l a r c h i n B r i t a i n a n d f o u n d t h a t t h e y g e n e r a l l y l i e b e t w e e n 1 2 a n d 1 5 . He c i t e s o t h e r a u t h o r s a s f i n d i n g N/P r a t i o s o f b e t w e e n 9 t o 1 2 . T h e t o t a l a m o u n t o f n i t r o g e n c o n t a i n e d i n a t r e e i n c r e a s e s w i t h 1 2 a g e a s a f u n c t i o n o f b i o m a s s a c c u m u l a t i o n a n d n i t r o g e n c o n c e n t r a t i o n w i t h i n a t r e e . S w i t z e r a n d N e l s o n ( 1 9 6 4 ) g i v e a v e r a g e N c o n c e n t r a t i o n s i n v a r i o u s l o b l o l l y p i n e t i s s u e s a t v a r i o u s a g e s o n g o o d s i t e s ( i n p e r c e n t o f o v e n -d r y w e i g h t ) , ( T a b l e 5).. T A B L E 5: A v e r a g e N - c o n c e n t r a t i o n s i n v a r i o u s l o b l o l l y p i n e t i s s u e s a t v a r i o u s a g e s o n g o o d s i t e s i n p e r c e n t o f o v e n d r y w e i g h t ( S w i t z e r a n d N e l s o n , 1 9 6 4 ) . T r e e P L A N T T I S S U E  A g e C u r r e n t f o l i a g e O l d e r b r a n c h e s S t e m b a r k S t e m w o o d 4 1.00 0 . 4 1 0.42 ' 0.16 8 0.95 0.24 0.24 0.06 18 1,08 0.23 0.23 0.06 3 0 1.22 0.22 0.19 0.04 56 1.16 0 . 2 1 0.17 0.03 A v e r a g e c o n c e n t r a t i o n s o f m a c r o n u t r i e n t s i n t h e a b o v e - g r o u n d o r g a n s o f N o r w a y s p r u c e s t a n d s i n C z e c h o s l o v a k i a w e r e c a l c u l a t e d b y M a t e r n a ( 1 9 6 4 ) , T h e r e s u l t s a r e s u m m a r i z e d i n T a b l e 6, T h e d i s t r i b u t i o n o f m a c r o n u t r i e n t s b e t w e e n t h e m a j o r c o m p o n e n t s o f t h e 3 6 - y e a r o l d s e c o n d - g r o w t h D o u g l a s - f i r f o r e s t w a s c a l c u l a t e d b y C o l e e t a l , ( 1 9 6 7 ) . T h e r e s u l t s a r e s u m m a r i z e d i n T a b l e 7. 1 3 T A B L E 6: T h e r a n g e o f n u t r i e n t c o n c e n t r a t i o n s ( i n p e r c e n t ) i n t h e d i f -f e r e n t t r e e c o m p o n e n t s i n v a r i o u s N o r w a y s p r u c e s t a n d s i n C z e c h o s l o v a k i a ( s i t e c l a s s c l a s s i f i c a t i o n a f t e r m o d i f i e d S c h w a p p a c h t a b l e s f o r C z e c h o s l o v a k i a ) , ( M a t e r n a , 1 9 6 4 ) . S i t e C l a s s N P K C a Mg S WOOD 1 0 , 1 3 3 - 0 , 0 1 5 - 0 , 0 3 0 - 0 , 0 6 6 - t r a c e 0 , 0 1 3 -0,217 0 , 0 3 0 0 , 0 5 1 0 , 0 8 6 0 , 0 3 5 5 0 , 1 4 7 - 0 , 0 2 4 - 0 , 0 3 4 - 0 , 0 4 4 - 0 , 0 0 7 - 0 , 0 1 3 -0 , 1 8 9 0 , 0 2 6 0 , 0 5 1 0 , 0 9 0 0 , 0 1 0 0 , 0 3 1 9 0 , 1 4 0 - 0 , 0 2 6 - 0 , 0 1 2 - 0 , 0 3 2 - 0 , 0 1 0 - 0 , 0 1 1 -0 , 2 1 0 0 , 0 4 8 0 , 0 4 2 0 , 0 6 6 0 , 0 1 8 0 , 0 2 9 BARK 1 0 , 3 6 7 - 0 , 0 3 8 - 0 , 2 0 2 - 0 , 5 0 4 - 0 , 0 1 4 - 0 , 0 4 3 -0 , 5 9 5 0 , 0 7 6 0 , 4 0 4 1 , 0 6 4 0 , 0 3 9 0 , 0 6 1 5 0 , 2 4 5 - 0 , 0 4 0 - 0 , 1 1 7 - 0 , 4 5 0 - 0 , 0 0 8 - 0 , 0 2 7 -0 , 4 8 3 0,087 0,216 1 , 0 2 0 0 , 0 2 9 0 , 0 6 0 9 0 , 2 9 4 - 0 , 0 3 3 - 0 , 0 5 2 - 0 , 4 9 4 - 0 , 0 3 5 - 0 , 0 1 5 -0 , 5 6 0 0 , 0 6 1 0,247 1 , 0 6 8 0 , 0 8 8 0 , 0 4 5 F O L I A G E 1 1 , 3 0 - 2 , 1 0 0 , 1 3 - 0 , 2 7 0 , 4 8 - 0 , 9 8 0 , 2 1 - 0 , 5 8 0 , 0 4 - 0 , 1 1 0 , 1 1 - 0 , 1 4 5 1 , 0 5 - 1 , 4 1 0 , 1 1 - 0 , 1 8 0 , 1 6 - 0 , 6 6 0 , 1 7 - 0 , 4 6 0 , 0 4 - 0 , 1 9 0 , 1 0 - 0 , 1 5 9 0 , 8 3 - 1 , 3 1 0 , 1 0 - 0 , 1 6 0 , 1 1 - 0 , 7 9 0 , 0 8 - 0 , 4 2 0 , 0 4 - 0 , 0 9 0 , 1 0 - 0 , 1 5 TWIGS 0 , 2 3 - 0 , 8 6 0 , 0 3 - 0 , 1 4 0 , 1 1 - 0 , 6 4 0 , 2 9 - 0 , 4 1 0 , 0 4 - 0 , 0 8 0 , 0 5 - 0 , 0 9 1 4 T A B L E 7: D i s t r i b u t i o n o f N, P, K a n d C a among t h e m a j o r c o m p o n e n t s o f a 3 6 - y e a r - o l d s e c o n d - g r o w t h D o u g l a s - f i r f o r e s t ( C o l e e t a l . , 1 9 6 7 ) E c o s y s t e m N P K C a C o m p o n e n t k g / h a % o f k g / h a % o f k g / h a % o f k g / h a % o f t o t a l t o t a l t o t a l t o t a l F o l i a g e 1 0 2 3 1 . 9 29 4 3 . 9 6 2 2 8 . 2 73 2 1 . 9 B r a n c h e s 6 1 1 9 . 1 1 2 1 8 . 2 3 8 1 7 . 3 1 0 6 3 1 . 8 Wood 77 2 4 . 0 9 1 3 . 6 5 2 2 3 . 6 47 1 4 . 1 B a r k 4 8 1 5 . 0 1 0 1 5 . 2 4 4 2 0 . 0 70 2 1 . 0 R o o t s 3 2 1 0 . 0 6 9.1 24 1 0 . 9 37 1 1 . 2 T o t a l 3 2 0 66 2 2 0 3 3 3 A s i m i l a r s t u d y w a s u n d e r t a k e n b y W i l l ( 1 9 6 4 ) i n a 1 2 - y e a r o l d s t a n d o f r a d i a t a p i n e o n t h e N o r t h I s l a n d o f New Z e a l a n d ( T a b l e 8 ) . T h e p e r c e n t a g e d i s t r i b u t i o n o f m a c r o n u t r i e n t s i n t h e a e r i a l c o m -p o n e n t s o f j a c k p i n e w a s c a l c u l a t e d b y M o r r i s o n ( 1 9 7 3 ) f o r t w e l v e n a t u r a l s t a n d s i n N o r t h e r n O n t a r i o . T a b l e 9 s u m m a r i z e s h i s r e s u l t s f o r t h r e e a g e c l a s s e s . A s a g e n e r a l r u l e , o n e c a n s a y t h a t t h e c o n c e n t r a t i o n o f n u t r i e n t s i s t h e h i g h e s t i n r e p r o d u c t i v e o r g a n s a n d s e e d s b e i n g f o l l o w e d b y f o l i a g e , t w i g s , b a r k o r b r a n c h e s a n d t r e e b o l e , s a p w o o d a n d h e a r t w o o d i n t h a t o r d e r . T h e n u t r i e n t c o n c e n t r a t i o n o f a p a r t i c u l a r t r e e c o m p o n e n t i s v e r y T A B L E 8: D i s t r i b u t i o n o f b i o m a s s a n d n u t r i e n t s w i t h i n a b o v e - g r o u n d t r e e c o m p o n e n t s o f 1 2 - y e a r - o l d s t a n d o f r a d i a t a p i n e , W h a k a r e w a r e w a F o r e s t ( W i l l , 1 9 6 4 ) . D r y N P K C a Mg_ M a t t e r % l b / % l b / % l b / % l b / % l b / l b / a c D.M.* a c D.M. a c D.M. a c D.M. a c D.M. a c S t a n d i n g t r e e s F o l i a g e 8 , 0 0 0 1.47 1 1 8 0.15 1 2 . 0 0.99 79 0.33 26 0.12 1 0 T w i g s 2 , 0 0 0 0.90 1 8 0.10 2.0 0.79 16 0 . 2 1 4 0.13 3 D e a d B r . 6,000 0.28 17 0 . 0 1 0.6 0.08 5 0.18 1 1 0.08 5 L i v e B r . 7 , 0 0 0 0.38 27 0.03 2.1 0.27 19 0.19 13 0.07 5 D e a d S t . 7 , 3 0 0 0.12 9 0 . 0 1 0.7 0.16 12 0 . 0 8 5 6 0 . 0 3 5 3 S m a l l L i v e S t e m s 9 , 3 0 0 0.12 1 1 0 . 0 1 0.9 0.13 1 2 0.09 8 0 . 0 3 5 3 L a r g e L i v e S t e m s 1 0 9 , 2 0 0 0.10 1 0 9 0 . 0 0 9 9.8 0.11 1 2 0 0.08 89 0 . 0 3 5 37 T o t a l 1 4 8 , 0 0 0 3 0 9 2 8 . 1 2 6 3 1 5 7 6 6 * D.M. - d r y m a t t e r . 16 T A B L E 9: A v e r a g e p e r c e n t a g e d i s t r i b u t i o n o f N, P, K, C a a n d Mg i n a e r i a l c o m p o n e n t s o f 12 n a t u r a l j a c k p i n e s t a n d s i n N o r t h e r n O n t a r i o ( M o r r i s o n , 1 9 7 3 ) . A g e L i v e D e a d S t e m S t e m L i v e C l a s s F o l i a g e B r a n c h e s B r a n c h e s B a r k Wood C o n e s C r o w n N I T R O G E N 20 6 1 . 8 a 1 9 . 9 a 1.8 a 5.2 a 1 0 . 9 a 0.4 a 8 1 . 7 a 3 0 3 6 . 5 b 1 9 . 4 a 1 1 . 8 b 9.5* b 2 2 . 4 b 0.4 a 5 5 . 9 b 65 2 8 . 0 c 2 1 . 0 a 2.7 a 9.3 b 3 6 . 3 c 2.7 b 4 9 . 0 b PHOSPHOROUS 20 6 6 . 6 a 1 5 . 0 a 0.6 a 5.4 a 1 1 . 9 a 0.5 a 8 1 . 6 a 3 0 4 2 . 8 b 1 7 . 8 a 4.1 b 9.9 a 2 4 . 7 b 0.7 a 6 0 . 6 b 65 4 2 . 6 b 1 9 . 0 a 1.6 a b 8.8 a 2 4 . 2 b 3.8 b 6 1 . 6 b P O T A S S I U M 20 4 2 . 0 a 3 1 . 6 a 0.7 a 6.7 a 1 8 . 6 a 0.4 a 7 3 . 6 a 3 0 2 3 . 3 b 2 6 . 8 b 3.9 b 1 0 . 0 a 3 5 . 7 b 0.3 a 5 0 . 1 b 6 5 1 5 . 8 c 2 5 . 4 b 1.3 a 8.2 a 4 7 . 9 c 1.4 b 4 1 . 2 b C A L C I U M 20 3 7 . 3 a 3 3 . 7 a 2.2 a 1 1 . 0 a 1 5 . 7 a 0.1 a 7 1 . 0 a 30 1 6 . 1 b 2 4 . 2 b 7.8 b 2 1 . 1 b 3 0 . 7 b 0.1 a 4 0 . 3 b 6 5 9.4 c 1 9 . 4 b 2.3 a 2 3 . 3 b 4 5 . 2 c 0.4 b 2 8 . 8 c MAGNESIUM 20 4 0 . 6 a 2 9 . 4 a 1.0 a 1 0 . 5 a 1 8 . 2 a 0.3 a 7 0 . 0 a 3 0 2 0 . 4 b 2 3 . 9 a b 3.0 a 1 3 . 3 a 3 9 . 2 b 0.2 a 4 4 . 3 b 6 5 1 5 . 6 b 2 0 . 0 b 2.2 a 9.0 a 5 1 . 1 c 2.1 b 3 5 . 6 b N o t e : C o r r e s p o n d i n g s u p e r s c r i p t l e t t e r s i n v e r t i c a l c o l u m n s i n d i c a t e n o s i g n i f i c a n t ( 5 % ) d i f f e r e n c e s b e t w e e n v a l u e s , a s c a l c u l a t e d b y a t - t e s t . 17 v a r i a b l e a n d d e p e n d s o n t h e e l e m e n t , s p e c i e s , a g e a n d n u t r i e n t a v a i l a b i l i t y i n a p a r t i c u l a r e c o s y s t e m . D i s t r i b u t i o n o f n u t r i e n t s b e t w e e n t h e v a r i o u s a b o v e - g r o u n d t r e e c o m p o n e n t s o f a s t a n d v a r i e s f o r d i f f e r e n t n u t r i e n t e l e -m e n t s , t r e e s p e c i e s a n d t r e e a g e s . M o s t o f t e n t h e p r o p o r t i o n s o f n u t r i e n t s o f y o u n g s t a n d s a p p e a r i n t h e s e q u e n c e : f o l i a g e a n d t w i g s > b r a n c h e s > s t e m b a r k > s t e m w o o d . T h e p r o p o r t i o n s o f n u t r i e n t s o f o l d s t a n d s a r e r e -v e r s e d : s t e m w o o d > s t e m b a r k > f o l i a g e a n d t w i g s >* b r a n c h e s , T h i s i s o n l y a g e n e r a l r u l e , h o w e v e r , a n d t h e a c t u a l d i s t r i b u t i o n o f p a r t i c u l a r n u t r i -t i o n a l e l e m e n t s w i t h i n a p a r t i c u l a r t r e e s t a n d may v a r y c o n s i d e r a b l y f r o m t h i s b a s i c p a t t e r n . 2 . 3 : R e m o v a l o f n u t r i e n t s f r o m a s i t e a s a r e s u l t o f t r e e h a r v e s t i n g . T h e a m o u n t o f n u t r i e n t s r e m o v e d f r o m a n e c o s y s t e m d u e t o l o g g i n g I s e q u a l t o t h e a m o u n t o f n u t r i e n t s i m m o b i l i z e d w i t h i n t h o s e a b o v e - g r o u n d c o m p o n e n t s t h a t a r e p h y s i c a l l y r e m o v e d f r o m t h e s i t e p l u s t h e a m o u n t o f n u t r i e n t s l o s t t h r o u g h t h e i n c r e a s e d s o i l l e a c h i n g w h i c h may a c c o m p a n y t h e i n c r e a s e i n t h e r a t e o f o r g a n i c m a t t e r d e c o m p o s i t i o n a n d t h e d i s r u p t i o n o f t h e n u t r i e n t c y c l e . I n m o u n t a i n t e r r a i n c o n s i d e r a b l e a m o u n t s o f n u t r i e n t s may a l s o b e l o s t a s t h e r e s u l t o f s o i l e r o s i o n ( C u r r y , 1 9 7 3 ) . T h e r e i s a c o n t i n u o u s i n p u t o f n u t r i e n t s t o a n e c o s y s t e m , h o w e v e r , i n t h e f o r m o f p r e c i p i t a t i o n , d u s t f a l l a n d b e d r o c k w e a t h e r i n g . T h e c a p i t a l o f a v a i l a b l e n u t r i e n t s r e s u l t s f r o m t h e s e i n p u t s a n d o u t p u t s a n d f r o m e x c h a n g e s b e t w e e n t h e a v a i l a b l e a n d u n a v a i l a b l e p o o l s i n t h e s o i l . T h e s i g n i f i c a n c e o f a n y 1 8 n u t r i e n t r e m o v a l i n h a r v e s t e d m a t e r i a l s m u s t b e v i e w e d t o g e t h e r w i t h o t h e r l o s s e s a n d t h e v a r i o u s i n p u t s b y w h i c h t h e l o s s e s may b e r e p l a c e d . T h e f o l l o w i n g d i s c u s s i o n w i l l b e l i m i t e d t o o u t p u t s i n l o g g e d m a t e r i a l s h o w e v e r , s i n c e t h i s p a r a m e t e r w a s , m e a s u r e d i n t h e r e s e a r c h f o r t h i s t h e s i s . T h e a m o u n t o f n u t r i e n t s r e m o v e d f r o m a n e c o s y s t e m i n t r e e b i o m a s s i s d e t e r m i n e d b y t h e l e v e l o f t r e e u t i l i z a t i o n . D i s t r i b u t i o n o f n u t r i e n t s b e t w e e n d i f f e r e n t c o m p o n e n t s o f t h e t r e e b i o m a s s h a s a l r e a d y b e e n d i s c u s s e d i n S e c t i o n 2 . 2 . F r o m t h i s r e v i e w i t i s a p p a r e n t t h a t f o r t h e m a j o r i t y o f t r e e s p e c i e s , a p p r o x i m a t e l y 5 0 p e r c e n t o r e v e n m o r e o f t h e t o t a l n u t r i e n t s i m m o b i l i z e d i n t h e a b o v e - g r o u n d t r e e b i o m a s s i s r e t a i n e d w i t h i n c o m p o n e n t s o f t h e t r e e c r o w n . T h e e x a c t p r o p o r t i o n v a r i e s f o r d i f -f e r e n t t r e e s p e c i e s a n d d i f f e r e n t n u t r i e n t e l e m e n t s , b u t i t i s g e n e r a l l y a s u b s t a n t i a l p r o p o r t i o n o f t h e t o t a l . T r e e b a r k i s a n o t h e r n u t r i t i o n a l l y r i c h c o m p o n e n t o f t r e e b i o m a s s . A s a g e n e r a l r u l e , o n e c a n s a y t h a t a p p r o x -i m a t e l y h a l f o f t h e n u t r i e n t s i n t h e a b o v e - g r o u n d t r e e b i o m a s s c a n b e r e -t a i n e d o n t h e s i t e a n d b e a v a i l a b l e f o r t h e g r o w t h o f t h e n e x t t r e e r o t a t i o n i f o n l y t h e t r e e b o l e s a r e h a r v e s t e d . Up t o 6 0 - 8 0 p e r c e n t o f t r e e n u t r i -e n t s c a n b e r e t a i n e d i f t h e l o g s a r e d e - b a r k e d a t s t u m p a n d o n l y t h e t r e e b o l e w o o d i s h a r v e s t e d . W e e t m a n a n d W e b e r c a l c u l a t e d t h a t h a r v e s t i n g o f t r e e b o l e s o n l y , a s c o m p a r e d w i t h w h o l e t r e e l o g g i n g , c a n r e d u c e r e m o v a l o f m a c r o n u t r i e n t s f r o m a n e c o s y s t e m b y a s m u c h a s 75 p e r c e n t o f N a n d P, a n d 6 0 p e r c e n t o f K, C a a n d Mg. 19 I I I D E S C R I P T I O N OF SAMPLE P L O T S L o c a t i o n a n d s i z e : T h e s t u d y w a s c o n d u c t e d a t t w o s i t e s . T h e f i r s t w a s i n i t i a l l y s e l e c t e d a s a p i l o t s t u d y f o r t h e s e c o n d s i t e , b u t , o w i n g t o t h e e x c e p t i o n -a l l y l a t e s n o w m e l t w h i c h r e s t r i c t e d a c c e s s t o t h e h i g h e r e l e v a t i o n s i t e u n t i l m i d - s u m m e r , a r e a s o n a b l y c o m p l e t e s t u d y w a s a c c o m p l i s h e d o n b o t h s i t e s . T h e r e w a s n o i n t e n t i o n t o m a k e a c o m p a r a t i v e s t u d y , h o w e v e r , a n d t h e s a m p l e p l o t s w e r e n o t s e l e c t e d t o b e c o m p a r a b l e . T h e f i r s t p l o t w a s l o c a t e d o n a n o r t h - f a c i n g a s p e c t o n a r o c k y r i d g e a t 7 2 0 m ( 2 , 2 0 0 f t . ) e l e v a t i o n i n t h e n o r t h - e a s t e r n c o r n e r o f t h e U n i v e r s i t y o f B r i t i s h C o l u m b i a R e s e a r c h F o r e s t a t H a n e y , B. C. I t l a y a p -p r o x i m a t e l y h a l f a m i l e e a s t o f t h e n o r t h e r n e n d o f C a t h e r i n e L a k e . T h e p l o t w a s 0.79 h a ( a b o u t 2 a c r e s ) i n s i z e , e l o n g a t e i n s h a p e , a n d i n c l u d e d 1 1 4 t r e e s . T h e s e c o n d p l o t w a s l o c a t e d o n a s o u t h - f a c i n g a s p e c t a t 1 , 5 0 0 m ( 4 , 5 0 0 f t . ) e l e v a t i o n o n t h e n o r t h b a n k o f t h e Mamquam R i v e r b e t w e e n S k o o k u m C r e e k a n d C r a w f o r d C r e e k . T h e p l o t w a s l o c a t e d o n a g e n t l y ( 2 5 p e r c e n t ) s l o p i n g b e n c h , w i t h s t e e p e r s l o p e s i m m e d i a t e l y b e l o w a n d s t e e p , b r o -k e n , r o c k y t e r r a i n a b o v e . T h e p l o t w a s 1.16 h a ( a b o u t 3 a c r e s ) i n s i z e , t r i a n g u l a r i n s h a p e , a n d i n c l u d e d 2 5 6 t r e e s . T h e l o c a t i o n o f t h e t w o p l o t s may b e s e e n i n m o r e d e t a i l i n „ F i g u r e 1. FIGURE 1: Location of the sample plots. l i t ' 21 V i e w o f t h e l o g g e d a r e a n e a r t h e S q u a m i s h s a m p l e p l o t . A r r o w i n d i c a t e s l o c a t i o n o f t h e p l o t . 2 3 l 24 A c c o r d i n g t o H a l l i d a y ' s c l a s s i f i c a t i o n ( H a l l i d a y , 1 9 3 7 ) b o t h p l o t s a r e l o c a t e d i n t h e S o u t h e r n C o a s t S e c t i o n . R o w e ' s ( 1 9 5 9 ) c l a s s i f i -c a t i o n , w h i c h i s b a s i c a l l y a n e l a b o r a t i o n u p o n t h a t o f H a l l i d a y , p l a c e s t h e S q u a m i s h p l o t i n t h e C o a s t a l S u b a l p i n e S e c t i o n a n d t h e H a n e y p l o t i n t h e S o u t h e r n P a c i f i c C o a s t S e c t i o n . A c c o r d i n g t o K r a j i n a ' s c l a s s i f i c a t i o n ( 1 9 6 9 ) , t h e S q u a m i s h p l o t i s i n t h e s u b a l p i n e f o r e s t s u b z o n e o f t h e M o u n -t a i n H e m l o c k Z o n e w h i l e t h e H a n e y p l o t i s i n t h e u p p e r e l e v a t i o n o f t h e w e t s u b z o n e o f t h e C o a s t a l W e s t e r n H e m l o c k Z o n e . A c c o r d i n g t o t h e F o r e s t P r o d u c t i v i t y Z o n e s o f t h e B C F S , t h e S q u a m i s h p l o t w o u l d b e d e s c r i b e d a s i n t e r m e d i a t e b e t w e e n C o a s t R a n g e S q u a m i s h - F r a s e r a n d S u b a l p i n e - A l p i n e T u n d r a - G l a c i e r s , w h i l e t h e H a n e y p l o t w o u l d b e C o a s t R a n g e S q u a m i s h -F r a s e r . C l i m a t e : No c l i m a t i c d a t a w e r e c o l l e c t e d o n t h e s a m p l e p l o t s . T a b l e 1 0 s u m m a r i z e s c l i m a t i c d a t a r e p r e s e n t a t i v e o f t h e a r e a o f b o t h s a m p l e p l o t s a c c o r d i n g t o K r a j i n a ( 1 9 6 5 ) , C l i m a t e i n t h e f o r e s t s u b z o n e o f t h e M o u n -t a i n H e m l o c k Z o n e i s r a t h e r u n f a v o u r a b l e t o t r e e g r o w t h . T h e n u m b e r o f f r o s t - f r e e d a y s r a n g e s f r o m 4 0 t o 1 2 0 p e r annum, w h i l e t h e mean d a i l y t e m p -e r a t u r e i s a b o v e 50 ° F f o r o n l y 2 o r 3 m o n t h s i n d i c a t i n g a v e r y s h o r t g r o w i n g s e a s o n . Snow p e r s i s t s f o r 8 t o 9 m o n t h s e v e r y y e a r , r e t r e a t i n g i n t h e m i d d l e o f J u l y a n d r e t u r n i n g i n t h e m i d d l e o f O c t o b e r . A n n u a l t o t a l p r e c i p i t a t i o n i s v e r y h i g h , r a n g i n g f r o m 2 , 2 0 0 t o 4 , 3 0 0 mm m a i n l y i n t h e T A B L E 1 0 : P l o t S q u a m i s h H a n e y Summary o f c l i m a t i c i n f o r m a t i o n ( K r a j i n a , 1 9 6 5 ) . D a t a r e p r e s e n t c h a r a c t e r i s t i c r a n g e s f o r t h e M o u n t a i n H e m l o c k a n d t h e W e s t e r n H e m l o c k ( w e t s u b z o n e ) B i o g e o c l i m a t l c z o n e s . No c l i m a t i c d a t a w e r e o b t a i n e d o n t h e p l o t s . M e a n A n n u a l 3 8 - 4 4 4 1 - 4 9 M e a n M o n t h l y J A N J U L 1 6 - 3 2 5 0 - 5 5 2 4 - 4 1 5 5 - 6 4 TEMPERATURE (°F) N u m b e r o f M o n t h s A n n u a l A b o v e B e l o w R a n g e 50°F 32°F 2-3 (-4) 1-6 3 0 - 4 0 4-6 0-3 (-4) 1 6 - 3 7 N u m b e r o f F r o s t - f r e e D a y s 4 0 - 1 2 0 ( 1 2 0 - ) 1 5 0 - 2 5 0 A b s o l u t e M a x . M i n . 7 5 - 8 6 - 3 1 - 1 0 7 8 - 1 0 4 - 2 2 - 1 9 P R E C I P I T A T I O N P l o t S q u a m i s h H a n e y A n n u a l T o t a l (mm) 2 2 0 0 -4 3 0 0 1 7 8 0 -6 6 5 0 A n n u a l S n o w f a l l (mm) 2 5 8 0 -2 0 0 0 0 1 2 6 -7 5 0 0 D r i e s t M o n t h (mm) 3 3 - 8 4 3 2 - 1 6 5 W e t t e s t M o n t h (mm) 3 0 0 -4 0 0 2 8 0 -1 1 6 0 C l i m a t e C l a s s i f i c a t i o n ( K o p p e n ) D f b - D f c C f b ( a n d t h e m i l d e s t D f b ) D a t a o r i g i n a l l y e x p r e s s e d i n i n c h e s . 26 f o r m o f s n o w . Summer i s u s u a l l y v e r y s h o r t (.6-8 w e e k s ) a n d i s h o t a n d d r y . I n t h e w e t s u b z o n e o f t h e W e s t e r n H e m l o c k Z o n e t h e c l i m a t e i s m o r e f a v o u r a b l e . T h e n u m b e r o f f r o s t - f r e e d a y s r a n g e s f r o m 1 5 0 t o 2 5 0 d a y s p e r a n n u m a n d t h e m e a n d a i l y t e m p e r a t u r e i s a b o v e 5 0 ° F f o r 4 t o 6 m o n t h s , p r o v i d i n g a r e a s o n a b l y l o n g p e r i o d f o r g r o w t h . Snow c o v e r r e m a i n s f o r 3 t o 4 m o n t h s e v e r y y e a r . A n n u a l t o t a l p r e c i p i t a t i o n r a n g e s b e t w e e n 1 , 7 0 0 - 2 , 2 0 0 mm, m a i n l y i n t h e f o r m o f r a i n . S o i l s ; B e c a u s e o f t i m e a n d f i n a n c i a l c o n s t r a i n t s s o i l a n a l y s e s w e r e n o t c o n d u c t e d a s p a r t o f t h i s t h e s i s s t u d y . S o i l c a t e g o r i e s f o r t h e S u b a l p i n e M o u n t a i n H e m l o c k Z o n e i n g e n e r a l h a v e b e e n d e s c r i b e d b y B r o o k e e t a l . ( 1 9 7 0 ) . S o i l s o n t h e S q u a m i s h a n d H a n e y s a m p l e p l o t s w e r e d e s c r i b e d a n d a n a l y s e d b y P h i r i ( 1 9 7 3 ) . V e g e t a t i o n : * A c c o r d i n g t o t h e c l a s s i f i c a t i o n o f B r o o k e e t a l . ( 1 9 7 0 ) , ( w h i c h i s b a s e d o n t h e Z u r i c h - M o n t p e l l i e r s y n e c o l o g i c a l c l a s s i f i c a t i o n s c h e m e ) f o r t h e M o u n t a i n H e m l o c k z o n e , t h e S q u a m i s h p l o t i s l o c a t e d i n t h e a b i e t o -t s u g o s u m m e r t e n s i a n a e v a r i a n t o f t h e A b i e t o - T s u g e t u m m e r t e n s i a n a e a s s o c i a -* L a t i n n ames o f t r e e s a r e t o b e f o u n d i n A p p e n d i x 1. 27 t i o n . T h e t r e e l a y e r i s f o r m e d . b y t w o s p e c i e s , P a c i f i c s i l v e r f i r a n d m o u n t a i n h e m l o c k . Y e l l o w c e d a r w h i c h i s a r e p r e s e n t a t i v e s p e c i e s f o r t h i s a s s o c i a t i o n ( B r o o k e , e t a l . , 1 9 7 0 ) w a s n o t p r e s e n t o n t h e p l o t . P a c i f i c s i l v e r f i r c o m p r i s e d 69 p e r c e n t o f t h e s t a n d w i t h m o u n t a i n h e m l o c k m a k i n g up t h e r e m a i n i n g 3 1 p e r c e n t . L i s t e d i n o r d e r o f a b u n d a n c e , t h e u n d e r s t o r e y v e g e t a t i o n o n t h e S q u a m i s h p l o t c o n s i s t e d o f V e r a t r u m e s c h s c h o l t z i i A . G r a y , V a c c i n i u m mem- b r a n a e e u m D o u g l . e x H o o k , a l a s k a e n s e H o w e l l , V_. o v a l i f o l i u m S m i t h , L u e t k e a  p e c t i n a t a ( P u r s h ) K u n t z e , R u b u s p e d a t u s J . E. S m i t h , a n d L u z u l a s p p . V a c c i n i u m  o v a l i f o l i u m a n d L u e t k e a p e c t i n a t a w e r e o b s e r v e d o n l y i n t h e d e p r e s s i o n a t t h e l o w e r e n d o f t h e p l o t . T h e V a c c i n i u m s p p . a n d R u b u s p e d a t u s a r e i n d i c a -t i v e o f e x t r e m e l y a c i d s o i l c o n d i t i o n s ( B r o o k e , p e r s o n a l c o m m u n i c a t i o n ) , w h i l e V e r a t r u m e s c h s c h o l t z i i i s i n d i c a t i v e o f t e m p o r a r y s o i l s e e p a g e ( K r a j i n a , p e r s o n a l c o m m u n i c a t i o n ) . T h e m o s s l a y e r w h i c h w a s p r e s e n t i n s c a t t e r e d p a t -c h e s w a s d o m i n a t e d b y D i c r a n u m f u s c e s c e n s T u r n , a n d D_. s c o p a r i u m Hedw. r e -f l e c t i n g t h e p r e s e n c e o f a c i d r a w h u m u s . U s i n g t h e same c l a s s i f i c a t i o n s c h e m e a s f o r t h e S q u a m i s h p l o t , t h e H a n e y p l o t i s a s s i g n e d t o t h e o r t h i c V a c c i n i u m - P l a g i o t h e c i u m e c o s y s t e m t y p e o f t h e A b i e t o - T s u g e t u m h e t e r o p h y l l a e f o r e s t p l a n t a s s o c i a t i o n ( O r l o c i , 1 9 6 5 ) . T h e t r e e s p e c i e s o n t h e p l o t w e r e w e s t e r n r e d c e d a r , w e s t e r n hem-l o c k a n d y e l l o w c e d a r c o m p r i s i n g 2 8 % , 5 9 % a n d 1 3 % o f t h e s t a n d , r e s p e c t i v e l y . S i z e c l a s s f r e q u e n c i e s a r e s h o w n b y s p e c i e s i n T a b l e 1 1 . 28 L i s t e d i n o r d e r o f a b u n d a n c e t h e u n d e r s t o r e y v e g e t a t i o n o n t h e H a n e y p l o t c o n s i s t e d o f M e n z i e s i a f e r r u g i n e a S m i t h , V a c c i n i u m p a r v i f o l i u m S m i t h , V_. a l a s k a e n s e H o w e l l , V . o v a l i f o l i u m S m i t h , G o o d y e r a o b l o n g i f o l i a R a f . , a n d G a u l t h e r i a s h a l l o n P u r s h , t h e l a t t e r b e i n g a s s o c i a t e d w i t h t h i c k a c c u m u l a t i o n s o f o r g a n i c m a t t e r o n t h e l i t h i c f o l i s o l s . T h e m o s s l a y e r w a s a l m o s t e x c l u s i v e l y R h y t i d i o p s i s r o b u s t a ( H o o k . ) B r o t h . 29 I V METHODS F i e l d S a m p l i n g : * A l l t r e e s o n t h e s a m p l e p l o t s w e r e n u m b e r e d a n d m e a s u r e d f o r d . b . h . , t o t a l t r e e l e n g t h , c r o w n l e n g t h , a n d c r o w n d i a m e t e r ( T a b l e 1 2 ) . T h e d . b . h . d a t a w e r e g r o u p e d i n t o 4 - i n c h ( 1 0 . 1 6 cm) s i z e c l a s s e s b y s p e c i e s , r e s u l t i n g i n s i x s u c h s i z e c l a s s e s ( 6 " t o 3 0 " ; 1 5 . 2 4 cm t o 7 6 . 2 0 cm) f o r t h e H a n e y p l o t a n d e l e v e n f o r t h e S q u a m i s h p l o t ( 6 " t o 5 0 " ; 1 5 . 2 4 cm t o 1 2 7 c m ) , ( T a b l e 1 1 ) . T r e e s f o r s a m p l i n g w e r e s e l e c t e d f r o m m o s t o f t h e s i z e c l a s s e s , t h e s e l e c t i o n b e i n g m ade a t r a n d o m w i t h i n e a c h s i z e c l a s s s a m p l e d . A t o t a l o f 13 t r e e s w e r e s a m p l e d a t t h e H a n e y p l o t a n d 2 0 a t t h e S q u a m i s h p l o t . B e -c a u s e o f t i m e l i m i t a t i o n s , e i g h t o f t h o s e a t t h e S q u a m i s h p l o t w e r e s a m p l e d f o r s t e m b i o m a s s o n l y , b r a n c h e s a n d f o l i a g e b e i n g e x c l u d e d . S a m p l e t r e e s w e r e f e l l e d a n d t h e f o l l o w i n g d a t a r e c o r d e d : a g e a t s t u m p ( a b o u t 1 f t . h i g h ; 3 0 . 5 c m ) , t o t a l l e n g t h o f s t e m , l e n g t h t o s i x i n c h d i a m e t e r , l e n g t h t o o n e i n c h d i a m e t e r , d . b . h . , c r o w n l e n g t h , a n d t h e a v e r a g e l e n g t h o f t h e t w o l a r g e s t b r a n c h e s . B r a n c h e s w e r e s e v e r e d f r o m t h e s t e m a n d d i v i d e d m a n u a l l y i n t o t h r e e g r o u p s : b i g b r a n c h e s ( > 1 i n c h d i a m e t e r ; 2.54 c m ) , s m a l l b r a n c h e s (1-H i n . d i a m e t e r ) , a n d t w i g s (<Ch i n , d i a m e t e r ; 0.64 c m ) , w i t h f o l i a g e . S e p a r a t i n g s m a l l b r a n c h e s f r o m t w i g s w i t h f o l i a g e w a s t h e m o s t t i m e - c o n s u m i n g * F i e l d w o r k r e q u i r e d 1 3 m a n - m o n t h s . 3 0 T A B L E 1 1 : D i a m e t e r c l a s s d i s t r i b u t i o n o n t h e s a m p l e p l o t s , F r e q u e n c y o f t r e e s S q u a m i s h H a n e y DBH * C l a s s ( i n . ) M o u n t a i n h e m l o c k P a c i f i c s i l v e r f i r W e s t e r n h e m l o c k W e s t e r n r e d c e d a r Y e l l o w c e d a r 6-10 4 3 25 ( 3 ) 2 6 ( 2 ) 1 0 - 1 4 6 ( 0 + 1 ) 19 ( 1 + 1 ) 12 ( 2 ) 7 2 1 4 - 1 8 1 0 ( 1 ) 2 3 ( 1 + 1 ) 18 ( 2 ) 7 5 ( 1 ) 1 8 - 2 2 1 2 ( 1 ) 2 9 ( 1 + 1 ) 1 0 ( 1 ) 1 1 ( 0 + 1 ) -2 2 - 2 6 1 2 ( 1 ) 27 ( 1 ) - 5 2 ( 1 ) 2 6 - 3 0 15 ( 1 + 2 ) 27 ( 1 ) 2 - -3 0 - 3 4 6 ( 1 ) 3 1 ( 1 + 1 ) - - -3 4 - 3 8 3 15 (1+D - - -3 8 - 4 2 2 4 - - -4 2 - 4 6 1 3 - - -> 4 6 3 - - - -F i r s t n u m b e r i n b r a c k e t s s t a t e s n u m b e r o f c o m p l e t e l y s a m p l e d t r e e s . W h e r e a s e c o n d n u m b e r i s s h o w n i t i n d i c a t e s t h e n u m b e r o f t r e e s s a m p l e d f o r t r e e b o l e o n l y . * ( d i a m e t e r b r e a t h e i g h t m e a s u r e d 4,5 f e e t a b o v e g r o u n d . ) T A B L E 1 2 : B a s i c s t a t i s t i c s o f o r i g i n a l m e a s u r e m e n t s f o r a l l t r e e s o v e r 6 i n c h e s o f d . b . h . o n t h e t w o s a m p l e p l o t s , n ; m o u n t a i n hem-l o c k = 7 4 ; P a c i f i c s i l v e r f i r = 1 8 2 ; w e s t e r n h e m l o c k = 6 7 ; y e l l o w c e d a r = 1 5 ; w e s t e r n r e d c e d a r = 3 2 . T r e e T r e e M e a n S t a n d . M i n i m u m M a x i m u m P a r a m e t e r S p e c i e s U n i t V a l u e D e v . V a l u e V a l u e T r e e M.h. m 0 . 6 2 1 0 . 2 4 6 0 . 2 3 1 1 . 4 1 0 d i a m e t e r P . s . f . 0 . 6 2 1 0 . 2 1 0 0 . 1 8 8 1 . 1 3 8 W.h. 0 . 3 3 2 0 . 1 3 3 0.157 0 . 7 0 6 Y . c . 0 . 3 4 8 0 . 1 3 6 0 . 1 8 0 0 . 6 3 5 W . r . c . 0 . 4 3 9 0 . 1 2 1 0 . 1 5 5 0 . 6 5 5 T r e e M.h. m 2 8 . 2 1 0 . 2 7.0 4 5 . 0 h e i g h t P . s . f . 3 0 . 8 8.9 9.0 4 6 . 0 W.h. 2 0 . 8 4.8 1 2 . 0 3 1 . 0 Y . c . 2 1 . 1 4.7 1 2 . 5 2 8 . 0 W . r . c . 2 0 . 9 4.8 7.0 3 0 . 0 C r o w n M.h. m 1 9 . 8 8.9 1.5 3 6 . 0 l e n g t h P . s . f . 2 0 . 8 7.8 1.5 3 8 . 0 W.h. 1 4 . 4 4.2 5.0 2 4 . 5 Y . c . 1 4 . 0 3.8 7.0 2 0 . 0 W . r . c . 1 4 . 4 3.8 4.0 2 5 . 0 C r o w n M.h. m 6.5 1.8 3.2 1 1 . 0 d i a m e t e r P . s . f . 6.2 1.5 1.8 1 0 . 2 W.h. 6.5 1.9 2.9 1 1 . 7 Y . c . 6.6 1.6 3.7 9.1 W . r . c . 5.0 2.5 0.5 8.5 V o l u m e o f M.h. m 3 4 . 0 4 8 — — — t r e e b o l e P . s . f . 4 . 6 8 2 - - -W.h. 0 . 8 9 6 - - -Y . c . 1 . 0 3 5 - - -W . r . c . 1 . 3 0 3 - - -j o b , a n d f o r t r e e s w i t h a l a r g e a m o u n t o f b r a n c h e s a s u b - s a m p l e ( a p p r o x i -m a t e l y 1 0 p e r c e n t ) w a s t a k e n f r o m w h i c h t h e p r o p o r t i o n s o f t h e s e t w o 32 b i o m a s s c o m p o n e n t s w a s d e t e r m i n e d , T w i g s a n d f o l i a g e w e r e s a m p l e d s e p a r -a t e l y f o r t h e t o p , m i d d l e a n d l o w e r p a r t o f t h e t r e e c r o w n . T h e f r e s h w e i g h t o f e a c h b r a n c h g r o u p w a s t a k e n a s q u i c k l y a s p o s s i b l e t o m i n i m i z e l o s s o f m o i s t u r e . Two r e p r e s e n t a t i v e s a m p l e s w e r e t a k e n f r o m e a c h g r o u p : ( a ) t o d e t e r m i n e o v e n d r y w e i g h t ( d r i e d a t 1 5 0 ° C) a n d m o i s t u r e c o n t e n t , ( b ) t o d e t e r m i n e c o n c e n t r a t i o n o f m a c r o n u t r i e n t s ( d r i e d a t 70° C ) , T h e s a m p l e s w e r e t a k e n a c c o r d i n g t o a t a b l e o f r a n d o m n u m b e r s w h i c h s e l e c t e d t h e p o s i t i o n o f t h e s a m p l e a l o n g t h e b r a n c h . T h e s a m p l e s c o n s t i t u t e d b e t w e e n 0.5 p e r c e n t a n d 5 p e r c e n t o f t h e t o t a l b r a n c h b i o m a s s , a n d w e i g h e d 2.0 - 4.0 k g f o r b i o m a s s d e t e r m i n a t i o n , a n d 1.0 - 1.5 k g f o r c h e m i c a l a n a l y s i s . T h e s a m p l e s o f b r a n c h e s u n d e r % - i n c h d i a m e t e r ( w h i c h i n c l u d e d f o l i a g e ) w e r e d r i e d a t 70° C t h e d a y t h e y w e r e c u t i n o r d e r t o p r e v e n t c h a n g e o f n u t r i e n t c o n t e n t i n t h e n e e d l e s . H e m l o c k f o l i a g e w a s e a s i l y s e p -a r a t e d f r o m t w i g s f o l l o w i n g d r y i n g . T h e f o l i a g e a n d t w i g s o f P a c i f i c s i l -v e r f i r , o n t h e o t h e r h a n d , d i d n o t s e p a r a t e s o e a s i l y a n d t h e r e f o r e t h e s e t w o b i o m a s s c a t e g o r i e s w e r e p o o l e d . F o l i a g e o f y e l l o w c e d a r w a s s e p a r a t e d m a n u a l l y f r o m t h e b r a n c h e s a t t h e p o i n t a t w h i c h t h e b r a n c h e s t u r n e d b r o w n . T h e t w i g s a n d f o l i a g e b i o m a s s s a m p l e s w e i g h e d b e t w e e n 2,0 - 4,0 k g . T h e w e i g h t o f c h e m i c a l s a m p l e s w a s b e t w e e n 1.0 - 1.5 k g . T r e e b o l e s w e r e t r i m m e d a t 1 - i n c h ( 2 , 5 4 cm) d i a m e t e r , a n d t h e n d i v i d e d u p i n t o s e c t i o n s e q u a l t o 1 0 p e r c e n t o f t h e t r i m m e d l e n g t h . Two 3 3 m e a s u r e m e n t s w e r e made o f b o t h o u t s i d e a n d i n s i d e b a r k d i a m e t e r a t r i g h t a n g l e s t o e a c h o t h e r a t t h e m i d d l e a n d e a c h e n d o f t h e s e c t i o n s . Two d i s c s , a p p r o x i m a t e l y 5 cm i n t h i c k n e s s , w e r e c u t f r o m t h e l a r g e e n d o f e a c h s t e m s e c t i o n , a n d t h e f r e s h w e i g h t o f t h e b a r k a n d w o o d f r o m o n e o f t h e m d e t e r -m i n e d . One d i s c w a s f o r b i o m a s s d e t e r m i n a t i o n a n d t h e o t h e r f o r c h e m i c a l a n a l y s i s . D i s c s s a t u r a t e d w i t h w a t e r ( b y i m m e r s i o n i n c r e e k f o r 24 h o u r s ) f o r b i o m a s s d e t e r m i n a t i o n w e r e p l a c e d i n p l a s t i c b a g s p a r t i a l l y f i l l e d w i t h w a t e r t o p r e v e n t m o i s t u r e l o s s a n d t h e a c c o m p a n y i n g s h r i n k a g e . L a b o r a t o r y A n a l y s e s : * B i o m a s s T h e v o l u m e o f b a r k a n d w o o d w a s d e t e r m i n e d o n o n e o f e a c h o f t h e p a i r s o f s t e m d i s c s b y t h e d i s p l a c e m e n t m e t h o d ( H e i n r i c h s 1 9 5 4 ) . E a c h s a m -p l e w a s c o m p l e t e l y i m m e r s e d i n w a t e r , ( b u t w i t h o u t t o u c h i n g t h e b o t t o m o r w a l l s o f t h e c o n t a i n e r ) a n d i t s v o l u m e d e t e r m i n e d b y s u b t r a c t i n g t h e w e i g h t o f w a t e r f r o m t h e w e i g h t o f w a t e r w i t h t h e s a m p l e f u l l y i m m e r s e d . T h e s a m -p l e s w e r e t h e n d r i e d a t 105° C f o r 4 8 h o u r s a n d w e i g h e d . S p e c i f i c g r a v i t y w a s c a l c u l a t e d f r o m t h e v o l u m e a n d w e i g h t o f e a c h d i s c ( A p p e n d i x 9 ) . B r a n c h b i o m a s s s a m p l e s w e r e d r i e d a t 1 0 5 ° C f o r 48 h o u r s a n d w e i g h e d . M o i s t u r e c o n t e n t w a s c a l c u l a t e d a s t h e d i f f e r e n c e b e t w e e n f r e s h w e i g h t a n d o v e n d r y w e i g h t . * L a b o r a t o r y a n a l y s e s r e q u i r e d 7 m a n - m o n t h s . 34 S a m p l e s f o r c h e m i c a l a n a l y s i s w e r e d r i e d a t 70° C ± 5 ° C f o r 4 8 h o u r s . I m m e d i a t e l y f o l l o w i n g d r y i n g , s a m p l e s w e r e g r o u n d i n a W i l e y m i l l t o p a s s a 2-mm m e s h a n d s e a l e d i n p o l y t h e n e b a g s t o a w a i t d r y - a s h i n g f o r t h e d e t e r m i n a t i o n o f K, C a a n d Mg. P o r t i o n s o f t h i s m a t e r i a l w e r e r e -g r o u n d t o p a s s a #40 m e s h a n d s e a l e d i n b a g s a s a b o v e t o a w a i t K j e l d a h l d i g e s t i o n f o r t h e d e t e r m i n a t i o n o f N a n d P. S a m p l e s w e r e s t o r e d f o r b e -t w e e n 2 a n d 1 0 m o n t h s p r i o r t o a n a l y s e s . S i n c e i t w a s s u s p e c t e d t h a t t h e s a m p l e s m i g h t h a v e a b s o r b e d m o i s t u r e w h i l s t b e i n g g r o u n d , 1 1 6 s a m p l e s c o v e r i n g a l l t y p e s o f m a t e r i a l w e r e s e l e c t e d a t r a n d o m i n O c t o b e r 1 9 7 2 ( a b o u t 2 m o n t h s a f t e r g r i n d i n g ) a n d t h e m o i s t u r e c o n t e n t d e t e r m i n e d b y r e - d r y i n g a t b o t h 70° C a n d 1 0 5 ° C. T h e r e s u l t s ( T a b l e 1 3 ) c o n f i r m t h a t m o i s t u r e h a d i n d e e d b e e n a b s o r b e d b e f o r e t h e s a m p l e s w e r e s e a l e d i n t h e b a g s . A p p r o p r i a t e c o r r e c t i o n s w e r e made t o t h e n u t r i e n t c o n c e n t r a t i o n d a t a . T A B L E 1 3 : P e r c e n t l o s s o f w e i g h t i n s a m p l e s f o r c h e m i c a l a n a l y s e s u p o n -r e - d r y i n g a f t e r 2-6 m o n t h s o f s t o r a g e . N o . o f 70°C D r y i n g 105°C D r y i n g S a m p l e S p e c i f i c a t i o n S a m p l e s T e m p e r a t u r e T e m p e r a t u r e A n a l y s e d M e a n SD M e a n SD Wood 20 5.06 1.01 5.63 0.88 B a r k 2 0 4.77 0.67 5.84 0.66 B i g B r a n c h e s 18 5 . 5 0 2.24 6.32 2.30 S m a l l B r a n c h e s 2 0 4.93 1.64 5.86 1.64 T w i g s 20 4.83 0,85 5.94 0 . 8 1 F o l i a g e 1 8 2.98 0.86 4 . 3 1 0.89 A v e r a g e f r o m a l l g r o u p s 4.66 0 . 0 2 3 5.65 0.03 3 5 P a n d N A n a l y s i s T h e m e t h o d u s e d w a s a m o d i f i e d K j e l d a h l d i g e s t i o n ( T w i n e a n d W i l l i a m s , 1 9 7 1 ) u s i n g s e l e n i u m a s a c a t a l y s t w i t h s u b s e q u e n t a n a l y s i s f o r N a n d P u s i n g a T e c h n i c o n A u t o ^ - A n a l y s e r . B e t w e e n 0.15 a n d 0.25 gm o f s a m -p l e w a s a d d e d t o a b o i l i n g t u b e c o n t a i n i n g 5 m l o f d i g e s t i o n m i x t u r e a n d s e l e n i z e d H e n g a r g r a n u l e s t o f a c i l i t a t e e v e n b o i l i n g . T h e t u b e s w e r e p l a c e d i n a 7 7 - p l a c e h e a t i n g b l o c k a n d d i g e s t e d a t 8 0 t o 1 0 0 ° C f o r 1 2 h o u r s , t h e n s l o w l y r a i s e d t o t h e b o i l i n g t e m p e r a t u r e o f t h e d i g e s t i o n m i x t u r e a n d h e l d t h e r e u n t i l c o m p l e t e l y c o l o u r l e s s a n d t r a n s p a r e n t . G l a s s b a l l s p l a c e d o v e r t h e m o u t h s o f t h e t u b e s a c t e d a s r e f l u x c o n d e n s e r s , w a s h i n g u n d i g e s t e d m a t e r i a l s p a t t e r e d o n t o t h e w a l l s o f t h e t u b e b a c k down i n t o t h e d i g e s t i o n m i x t u r e . When t h e d i g e s t s w e r e c o o l t h e y w e r e d i l u t e d t o 1 0 0 m l w i t h d e -i o n i z e d w a t e r , a n d l e f t t o s t a n d o v e r n i g h t t o p e r m i t s e t t l i n g o f a n y u n d i -g e s t e d s o l i d s . T h e s u p e r n a t a n t d i g e s t a t e w a s d e c a n t e d i n t o p l a s t i c b o t t l e s t o a w a i t a n a l y s i s . N i t r o g e n w a s d e t e r m i n e d u s i n g a n a m m o n i a c a r t r i d g e o f a T e c h n i c o n A u t o - A n a l y s e r w h i c h m a k e s u s e o f t h e B e r t h e l o t r e a c t i o n ( r e a c t i o n o f a m m o n i a w i t h s o d i u m p h e n a t e a n d s o d i u m h y p o c h l o r i t e t o y i e l d a b l u e i n d o p h e n o l c om-p l e x w h i c h i s q u a n t i f i e d i n a c o l o r i m e t e r ) . P h o s p h o r u s w a s d e t e r m i n e d u s i n g a n o r t h o - p h o s p h a t e c a r t r i d g e o f a T e c h n i c o n A u t o - A n a l y s e r , T h i s i n v o l v e s f o r m a t i o n o f a r e d u c e d p h o s p h o -m o l y b d a t e c o m p l e x , w h i c h i s q u a n t i f i e d i n a c o l o r i m e t e r . 36 C a t i o n A n a l y s i s A n a l y s i s f o r K, C a a n d Mg w a s c o n d u c t e d o n a V a r i a n T e c h t r o n A t o m i c A b s o r p t i o n S p e c t r o p h o t o m e t e r ( M o d e l A A 5 ) f o l l o w i n g d r y - a s h i n g . One gm o f s a m p l e w a s w e i g h e d i n t o a s i l i c a c r u c i b l e a n d a s h e d i n a m u f f l e f u r -n a c e a t 200° C f o r 1 % - 2 h o u r s , f o l l o w e d b y 4 7 5 ° C ± 5 ° C f o r 1 2 h o u r s . T h e w e i g h t o f a s h f o r e a c h s a m p l e w a s d e t e r m i n e d a n d t h e n t a k e n up i n 5 m l o f 2 0 % H C 1 . A f u r t h e r 1 5 m l o f h y d r o c h l o r i c a c i d w a s t h e n a d d e d a n d t h e c r u c i b l e h e a t e d o n a s a n d b a t h t o f a c i l i t a t e s o l u t i o n . T h e s o l u t i o n w a s t h e n t r a n s f e r r e d t o a v o l u m e t r i c f l a s k , made up t o 1 0 0 m l , a n d t r a n s f e r r e d t o a p l a s t i c b o t t l e f o r s t o r a g e p r i o r t o a n a l y s i s . A n a i r - a c e t y l e n e f l a m e w a s u s e d f o r a l l d e t e r m i n a t i o n s . B e c a u s e t h e r e a r e a n u m b e r o f i n t e r f e r e n c e s p o s s i b l e i n t h e m e t h o d u s e d , t h e f o l l o w i n g i n v e s t i g a t i o n s w e r e u n d e r t a k e n : 1. T o c h e c k t h e i n f l u e n c e o f p y r o p h o s p h a t e s a n d h y d r a t e d s i l i c a o n d e t e r m i n a t i o n o f C a , Mg a n d K, 2 8 s a m p l e s w e r e p r o c e s s e d a s f o l l o w s ( A l l a n , 1969).. T h e a s h o b t a i n e d f r o m t h e m u f f l e f u r n a c e w a s d i s s o l v e d i n 5 m l o f 2 0 % HC1 a n d s l o w l y t a k e n t o d r y n e s s o n a s a n d h e a t e r ( t o h y d r o l y z e t h e p y r o p h o s p h a t e s a n d t o d e h y d r a t e t h e s i l i c a ) . T h i s o p e r a t i o n w a s r e p e a -t e d w i t h a f u r t h e r 5 m l o f 2 0 % H C 1 . T h e r e s i d u e w a s t h e n d i s s o l v e d i n 2 0 m l o f 2 0 % H C 1 , a n d a n a l y s e d o n a n A t o m i c A b s o r p t i o n S p e c t r o p h o t o m e t e r . No s i g -n i f i c a n t d i f f e r e n c e s w e r e f o u n d b e t w e e n t h e c o n c e n t r a t i o n s o f C a , Mg a n d K o b t a i n e d b y t h i s m e t h o d a n d t h o s e o b t a i n e d b y t h e b a s i c m e t h o d w h i c h vras t h e r e f o r e u s e d t h r o u g h o u t t h e s t u d y . 3 7 2, T o d e t e r m i n e t h e c h e m i c a l i n t e r f e r e n c e o f p h o s p h o r o u s , t w o r e l e a s i n g a g e n t s w e r e t e s t e d : s t r o n t i u m c h l o r i d e a n d l a n t h a n u m c h l o r i d e , t h e f o r m e r a t 3 , 0 0 0 ppm a n d t h e l a t t e r a t 1 0 , 0 0 0 ppm ( D a v i d , I 9 6 0 ) , No s i g n i f i c a n t d i f f e r e n c e w a s f o u n d i n t h e c o n c e n t r a t i o n o f C a a n d Mg u s i n g e i t h e r o f t h e s e r e l e a s i n g a g e n t s a s c o m p a r e d t o t h e b a s i c m e t h o d w h i c h w a s t h e r e f o r e u s e d t h r o u g h o u t t h e s t u d y . C h e c k o n t h e a c c u r a c y o f t h e c h e m i c a l a n a l y s i s T o v e r i f y t h e r e l i a b i l i t y o f t h e c h e m i c a l a n a l y s e s , some s a m p l e s w e r e s e n t t o o t h e r l a b o r a t o r i e s f o r a n a l y s i s . F o u r s a m p l e s w e r e s e n t t o t h e P a c i f i c F o r e s t R e s e a r c h C e n t r e i n V i c t o r i a , B. C., a n d f i v e s a m p l e s w e r e s e n t t o t h e R e s e a r c h D i v i s i o n o f t h e B r i t i s h C o l u m b i a F o r e s t S e r v i c e i n V i c t o r i a , B. C. T h e r e s u l t s f r o m a l l t h r e e l a b o r a t o r i e s a r e s u m m a r i z e d i n T a b l e 1 4 , T h e r e s u l t s o f t h i s s t u d y s h o w c o n s i s t e n t l y l o v e r f i g u r e s t h a n t h o s e f r o m t h e o t h e r l a b o r a t o r i e s , s u g g e s t i n g a g e n e r a l u n d e r - e s t i m a t i o n i n my d a t a . O n l y o n e s a m p l e ( t r e e n u m b e r 6 8 ) s h o w s s e r i o u s ( 4 0 p e r c e n t ) u n d e r -e s t i m a t i o n h o w e v e r . F r o m t h i s t a b l e i t a p p e a r s t h a t t h e d a t a o n b i o m a s s c h e m i c a l c o n t e n t s may b e u n d e r - e s t i m a t e d b u t i n g e n e r a l , my r e s u l t s a r e r e a s o n a b l y a c c u r a t e . 3 8 T A B L E 1 4 : C o m p a r i s o n o f r e s u l t s o f c h e m i c a l a n a l y s e s f o r f i v e r a n d o m l y s e l e c t e d s a m p l e s ( r e s u l t s a r e i n p e r c e n t ) , T r e e R e s u l t S a m p l e S p e c i f i c a t i o n N o , o f P. s i l , f i r , f o l i a g e 22 UBC 1 ( l o w t h i r d o f c r o w n ) B C F S 2 P F R C 3 M o u n . h e m l o c k , t w i g s 6 8 UBC ( l o w t h i r d o f c r o w n ) B C F S P F R C M o u n , h e m l o c k , t w i g s 1 1 9 UBC ( l o w t h i r d o f c r o w n ) B C F S P F R C P. s i l . f i r , b a r k 1 7 8 UBC ( d i s c N o . 7) B C F S P F R C P. s i l , f i r , b a r k 2 1 5 UBC ( d i s c N o . 5 ) B C F S P F R C 1 - U n i v e r s i t y o f B r i t i s h C o l u m b i a , L a b o r a t o r y . 2 - B r i t i s h C o l u m b i a F o r e s t S e r v i c e , 3 - C a n a d i a n F o r e s t r y S e r v i c e , P a c i f N P K C a Mg 0.76 0.77 0.75 0.107 0 . 1 0 8 0 . 0 8 6 0 . 3 1 0.39 0.22 0,56 0.69 0.66 0 . 0 7 3 0 . 0 7 6 0 . 0 7 6 0.24 0.55 0.46 0.034 0 . 0 7 1 0 . 0 6 0 0.17 0.24 0.14 0.12 0.20 0.24 0 . 0 4 5 0 . 0 5 3 0 . 0 5 6 0,40 0.42 0.43 0 . 0 6 1 0 . 0 6 5 0 . 0 6 0 0.14 0.23 0.13 0.19 0.28 0.23 0 . 0 4 6 0 . 0 5 5 0 . 0 5 6 0.18 0.22 0.17 0 . 0 3 5 0 . 0 3 6 0 . 0 3 0 0 . 1 1 0.15 0.10 0 . 3 1 0.36 0.30 0 . 0 3 1 0.034 0 . 0 2 2 0 . 2 1 0.22 0.037 0.037 0.08 0.10 0.29 0.35 0 . 0 2 2 0. 0 2 5 F a c u l t y o f F o r e s t r y , F o r e s t E c o l o g y R e s e a r c h D i v i s i o n , V i c t o r i a , i c F o r e s t R e s e a r c h C e n t r e , V i c t o r i a . D a t a A n a l y s i s : * B i o m a s s D a t a D r y b i o m a s s o f w o o d a n d b a r k w a s c a l c u l a t e d f r o m s p e c i f i c g r a v i t y a n d v o l u m e d a t a f o r i n d i v i d u a l t r e e b o l e s e c t i o n s , V o l u m e o f e a c h s e c t i o n * D a t a a n a l y s i s r e q u i r e d 5 m a n - m o n t h s . 3 9 ( w o o d a n d b a r k ) w a s c a l c u l a t e d a c c o r d i n g t o N e w t o n ' s f o r m u l a : ft Ik ( d 2 + 4 d | + d 2 ) w h e r e d ^ , d ^ a n d d 2 a r e d i a m e t e r s a t b o t h e n d s a n d t h e m i d d l e o f t h e s e c t i o n , r e s p e c t i v e l y ; S i s t h e l e n g t h o f t h e s e c t i o n . V o l u m e o f w o o d w a s c a l c u l a t e d i n t h e same m a n n e r . V o l u m e o f b a r k w a s o b t a i n e d b y s u b t r a c t i n g t h e v o l u m e o f w o o d f r o m t h e v o l u m e o f t h e w h o l e s e c t i o n . T h e v o l u m e s o f t r e e b o l e s c a l c u l a t e d a s t h e sum o f t h e v o l u m e s o f t h e t e n s e c t i o n s u s i n g N e w t o n ' s f o r m u l a w e r e c o m p a r e d w i t h v o l u m e s o b t a i n e d u s i n g B. C. F o r e s t S e r v i c e v o l -ume e q u a t i o n s ( T a b l e 1 5 ) . V o l u m e s c a l c u l a t e d b y t h e f o r m e r m e t h o d w e r e f o u n d t o b e s m a l l e r t h a n v o l u m e s o b t a i n e d u s i n g t h e B, C. F o r e s t S e r v i c e v o l u m e e q u a t i o n s . T h e d i f f e r e n c e m i g h t b e e x p l a i n e d b y a g r e a t e r t a p e r t h a n i s a s s u m e d i n t h e F o r e s t S e r v i c e e q u a t i o n s o r i t c a n b e t h e r e s u l t o f s a m -p l i n g a s m a l l n u m b e r , o f t r e e s . D r y b i o m a s s o f w o o d f o r a n y s e c t i o n w a s o b t a i n e d b y m u l t i p l y i n g t h e v o l u m e o f w o o d f o r t h a t s e c t i o n b y t h e m e a n o f t h e v a l u e s o f s p e c i f i c g r a v i t y o f w o o d f o r t h a t s e c t i o n a n d t h e n e x t s e c t i o n u p t h e s t e m . B a r k b i o m a s s w a s e s t i m a t e d i n t h e same w a y . B ( ± ) „ sgU) Y S s C i + i) . v ( i ) w h e r e B i s b i o m a s s , S g i s s p e c i f i c g r a v i t y a n d V i s v o l u m e o f t h e i - t h s e c -t i o n . O n l y o n e s p e c i f i c g r a v i t y v a l u e w a s u s e d f o r t h e u p p e r m o s t s e c t i o n . 4 0 T A B L E 1 5 : A v e r a g e p e r c e n t d i f f e r e n c e b e t w e e n t r e e t r u n k v o l u m e s c a l c u l a -t e d a c c o r d i n g t o B C F S v o l u m e e q u a t i o n s a n d t h o s e c a l c u l a t e d s e c t i o n - b y - s e c t i o n a c c o r d i n g t o N e w t o n ' s f o r m u l a . N o , o f o b s e r -v a t i o n s : M o u n t a i n h e m l o c k = 8; P a c i f i c s i l v e r f i r = 1 2 ; W e s t e r n h e m l o c k = 8; Y e l l o w c e d a r = 4, P E R CENT D I F F E R E N C E = V S N - V B F S # 1 0 0 VSN VSN = v o l u m e o f t r e e b o l e a c c o r d i n g t o N e w t o n ' s f o r m u l a . V B C F S = v o l u m e o f t r e e b o l e a c c o r d i n g t o B C F S v o l u m e e q u a t i o n . T r e e S p e c i e s P e r C e n t D i f f e r e n c e * M o u n t a i n h e m l o c k - 8.304 P a c i f i c s i l v e r f i r - 1 0 . 8 7 8 W e s t e r n h e m l o c k - 8 . 1 8 3 Y e l l o w c e d a r - 1 4 . 0 0 6 1 - R e a l v o l u m e ( s e c t i o n - b y - s e c t i o n ) i s s m a l l e r t h a n t a b u l a r v o l u m e . B i o m a s s o f b i g b r a n c h e s , s m a l l b r a n c h e s a n d t w i g s a n d n e e d l e s w a s c a l c u l a t e d b y m u l t i p l y i n g t h e f r e s h w e i g h t o f t h e p a r t i c u l a r c o m p o n e n t b y t h e a p p r o p r i a t e f i g u r e f o r p e r c e n t a g e d r y w e i g h t ( A p p e n d i x 1 0 ) . B a s i c s t a t i s t i c s f o r s a m p l e d t r e e s a r e s u m m a r i z e d i n T a b l e 1 6 . B i o m a s s d i s t r i b u -t i o n f o r a l l s a m p l e d t r e e s i s p r e s e n t e d i n A p p e n d i c e s 4 a n d 5. R e g r e s s i o n e q u a t i o n s w e r e d e v e l o p e d t o c a l c u l a t e t h e b i o m a s s b y c o m p o n e n t s f o r a l l t r e e s o n t h e s a m p l e p l o t s w i t h t h e e x c e p t i o n o f y e l l o w T A B L E 1 6 : B a s i c s t a t i s t i c s f o r s a m p l e d t r e e s , n ; M o u n t a i n h e m l o c k = 5; P a c i f i c s i l v e r f i r = 7; W e s t e r n h e m l o c k = 8; Y e l l o w c e d a r = 4. T r e e T r e e M e a n S t a n d a r d M i n i m u m M a x i m u m P a r a m e t e r S p e c . U n i t V a l u e D e v i a t i o n V a l u e V a l u e d . b . h . M.h. m 0 . 6 0 2 0 . 1 3 3 0.437 0 . 7 6 2 P . s . f . 0.587 0 . 2 2 1 0 . 3 1 0 0 . 9 0 4 W.h. 0 . 3 1 4 0.116 0 . 1 6 0 0 . 4 9 0 Y . c . 0 . 3 5 3 0 . 1 9 1 0 . 1 8 0 0 . 6 0 2 T r e e M.h. m 3 2 . 1 4.7 2 5 . 1 3 8 . 0 l e n g t h P . s . f . 3 0 . 9 8.7 1 7 . 8 4 1 . 2 W.h. 2 0 . 8 4.3 1 5 . 5 2 7 . 7 Y . c . 2 1 . 6 5.8 1 4 . 8 2 7 . 2 C r o w n M.h. m 2 3 . 9 2.0 2 0 . 6 2 5 . 8 l e n g t h P . s . f . 2 2 . 4 6.4 1 1 . 5 2 8 . 9 W.h. 1 5 . 3 4.3 1 0 . 3 2 3 . 2 Y . c . 1 3 . 2 2.9 9.1 1 5 . 4 C r o w n M.h. m 6.6 2.1 4.7 9.4 d i a m e t e r P . s . f . 7.1 1.3 5.5 8.5 W.h. 5.0 1.6 2.9 7.3 Y . c . 7.4 1.7 5.8 9.1 W e i g h t o f M.h. k g 1 3 5 2 . 6 5 7 5 . 6 7 0 2 . 1 9 9 4 , w o o d P . s . f . 1 3 3 2 . 5 1 0 7 0 . 9 1 7 5 3 0 6 6 W.h. 2 8 0 . 6 1 9 5 . 0 66 6 0 1 Y . c . 4 2 1 . 3 - 4 0 1 . 8 6 8 9 3 8 W e i g h t o f M.h. k g 3 6 4 . 1 1 6 3 . 6 1 7 8 0 575. b a r k P . s . f . 2 7 7 . 3 2 2 6 . 0 37„ 6 8 6 W.h. 6 6 . 2 5 2 . 5 1 1 , 1 5 3 Y . c . 3 5 . 7 3 2 . 8 6 79 W e i g h t o f M.h. k g 1 5 0 . 4 9 9 . 5 53; 2 9 0 b i g P . s . f . 1 6 1 . 9 1 5 3 . 8 33 4 1 1 b r a n c h e s W.h. 4 0 . 4 4 3 . 4 lo 1 2 2 Y . c . 4 6 . 1 5 7 . 4 4 126. W e i g h t o f M.h. k g 3 6 . 5 1 1 . 1 2 4 5 4 s m a l l P . s . f . 5 3 . 3 2 4 . 7 1 8 86 b r a n c h e s W.h. 4 5 . 0 2 5 . 8 •16 74-Y . c . 3 3 . 8 1 8 . 2 14 53-T A B L E 16 ( c o n t i n u e d ) : T r e e P a r a m e t e r T r e e S p e c . U n i t M e a n V a l u e S t a n d a r d D e v i a t i o n M i n i m u m V a l u e M a x i m u m V a l u e W e i g h t o f t w i g s M.h. P . s . f . W.h. Y . c . k g 4 9 . 6 1 2 . 6 1 8 . 9 8.1 24 , 5, 70 24 W e i g h t o f f o l i a g e M.h. P . s . f . W.h. Y . c . k g 7 0 . 6 2 3 . 2 2 1 . 3 1 7 . 4 35. 3, 90 , 4 8 W e i g h t o f t w i g s a n d f o l i a g e M.h. P . s . f . W.h. Y . c . k g 1 2 0 . 2 1 2 0 . 9 3 5 . 8 5 9 . 4 3 9 . 5 7 3 . 4 2 5 . 2 4 0 . 2 5 9 c 3 0 9 15 1 6 1 1 9 8 72. 1 1 0 W e i g h t o f w h o l e t r e e M.h. P . s . f . W.h. Y . c . k g 2 0 2 4 . 2 1 9 4 9 . 2 4 6 8 . 0 5 7 5 . 0 8 7 5 . 2 1 5 1 9 . 0 3 3 4 . 0 5 5 3 . 0 1 0 1 6 r 2 9 5 , 1 0 9 1 0 7 c 3 0 0 7 4 3 5 4 1 0 2 1 1 3 0 5 A g e M.h. P . s . f . W.h. Y . c . Y e a r 4 3 7 4 0 5 2 8 3 1 7 9 3 2 5 3 0 0 1 1 5 1 1 5 5 3 0 5 3 0 4 1 0 2 2 1 4 3 c e d a r a t t h e H a n e y p l o t , Two r e g r e s s i o n m o d e l s w e r e t e s t e d ( F r e e s e , 1 9 6 4 ) . A ) L i n e a r m o d e l : Y = a + b x ( 1 ) B) E x p o n e n t i a l m o d e l : Y = a X b C X ( 2 ) T r a n s f o r m e d t o l i n e a r l o g a r i t h m i c f o r m u s i n g l o g a r i t h m i c t r a n s f o r m a t i o n l o g Y = b Q + b1 l o g X + b 2 X ( 3 ) w h e n C = 1 , t h e n ( 2 ) b e c o m e s : l o g Y = b 0 + b L l o g X ( 4 ) w h e n b = 0, t h e n ( 2 ) b e c o m e s : l o g Y = b Q + b 2 X ( 5 ) T w e n t y - f i v e i n d e p e n d e n t v a r i a b l e s w e r e t e s t e d i n t h e s e r e g r e s s i o n m o d e l s : D D.CL + ( D . C L ) 2 BA.H.CL H C L 2 BA.H + C L C L D.CW 2 D 2 , H + C L D 2.H D.CW.CL 2 BA BA.II D . C L . B T L CW 2 D.CL D 2 . H . C L CW 2.CL D . C L 2 D 2.H.CW D,CW 2.CL D . H . C L 2 ( B A . H ) + (CW.CL) C r o w n v o l u m e C r o w n s u r f a c e 44 w h e r e D = d . b . h , H = t r e e l e n g t h C L = c r o w n l e n g t h CW = c r o w n w i d t h BA = c r o s s s e c t i o n a l a r e a a t b r e a s t h e i g h t B T L = t r e e l e n g t h m i n u s c r o w n l e n g t h C r o w n v o l u m e = CW 2,CL C r o w n s u r f a c e = 7V . CW . "1 4 C L + CW: O f t h e s e i n d e p e n d e n t v a r i a b l e s , 14 w e r e f o u n d s i g n i f i c a n t a t 1 p e r c e n t p r o b a b i l i t y l e v e l f o r d e s c r i b i n g b i o m a s s o f d i f f e r e n t t r e e c o m p o n e n t s i n l i n e a r , l o g a r i t h m i c o r i n b o t h m o d e l s . 12 4 S i g n i f i c a n t v a r i a b l e s w e r e : D 2 . H CR.VOL CW 2 B A C R .SURF D.CW 2 BA.H D D.CL D 2.H.CW H D.CL 2 ( BA.H). + (CL.CW). C L B a s i c s t a t i s t i c s f o r i n d e p e n d e n t v a r i a b l e s a r e p r e s e n t e d i n A p p e n d i x 2. I n d e p e n d e n t v a r i a b l e s , a s t h e y a p p e a r i n p a r t i c u l a r m o d e l s f o r d i f f e r e n t t r e e c o m p o n e n t s a r e p r e s e n t e d i n A p p e n d i x 3, 4 5 L o g a r i t h m i c r e g r e s s i o n e q u a t i o n s w e r e u s e d t o e s t i m a t e b i o m a s s f o r p a r t i c u l a r t r e e c o m p o n e n t s a n d t h e e n t i r e t r e e l a y e r b i o m a s s o n b o t h s a m p l e p l o t s ( A p p e n d i x 1 3 ) , R e s u l t s a r e s u m m a r i z e d i n T a b l e 17, R e s i d u a l s w e r e p l o t t e d f o r a l l r e g r e s s i o n e q u a t i o n s u s e d f o r c a l c u l a t i n g b i o m a s s o n t h e s a m p l e p l o t s i n o r d e r t o t e s t f o r l a c k o f f i t . N u t r i e n t C o n c e n t r a t i o n D a t a : C o n c e n t r a t i o n o f e l e m e n t s w a s r e a d i n ppm ( p a r t s p e r m i l l i o n ) e i t h e r o n t h e T e c h n i c o n A u t o - A n a l y s e r ( f o r N a n d P ) . o r o n t h e A t o m i c A b s o r p -t i o n S p e c t r o p h o t o m e t e r ( f o r C a , Mg, K ) . R e s u l t s w e r e t r a n s f e r r e d i n t o p e r -c e n t a g e c o n c e n t r a t i o n . T h e a m o u n t o f n u t r i e n t s w i t h i n t h e s a m p l e d t r e e s was o b t a i n e d b y m u l t i p l y i n g t h e b i o m a s s o f t h e v a r i o u s t r e e c o m p o n e n t s b y t h e a p p r o p r i a t e f i g u r e f o r c o n c e n t r a t i o n o f n u t r i e n t e l e m e n t s a n d s u m m i n g t h e p r o d u c t s . T h e d i s t r i b u t i o n o f n u t r i e n t s w i t h i n t h e s a m p l e d t r e e s a n d c o n c e n t r a t i o n o f n u t r i e n t s w i t h i n d i f f e r e n t t r e e c o m p o n e n t s a r e l i s t e d i n A p p e n d i c e s 6 a n d 7. T h e t o t a l a m o u n t o f n u t r i e n t s c o n t a i n e d w i t h i n t h e a b o v e - g r o u n d p o r t i o n o f t h e t r e e l a y e r o n t h e s a m p l e p l o t s w a s c a l c u l a t e d b y m u l t i p l y i n g t h e w e i g h t o f i n d i v i d u a l b i o m a s s c o m p o n e n t s o f e a c h t r e e b y t h e n u t r i e n t c o n c e n t r a t i o n s f o r t h e s e c o m p o n e n t s m e a s u r e d o n a s a m p l e d t r e e o f t h e same d . b . h . c l a s s . F o r t h o s e t r e e s i n d . b . h . c l a s s e s f o r w h i c h n o t r e e w a s s a m p l e d , n u t r i e n t c o n c e n t r a t i o n d a t a f r o m t h e n e a r e s t s a m p l e d s i z e c l a s s w a s u s e d . 46 B y s u m m a t i o n , t h e t o t a l a b o v e - g r o u n d b i o m a s s n u t r i e n t c o n t e n t w a s o b t a i n e d f o r t h e p l o t s , w i t h a b r e a k d o w n b y i n d i v i d u a l b i o m a s s c o m p o n e n t s . No s i g n i f i c a n t r e g r e s s i o n e q u a t i o n s w e r e f o u n d f o r t h e b i o m a s s o f s m a l l b r a n c h e s 0-~H i n c h d i a m e t e r ) f o r m o u n t a i n h e m l o c k , p o s s i b l y b e -c a u s e o f t h e s m a l l n u m b e r o f s a m p l e d t r e e s ( 5 ) . T h e a m o u n t o f b i o m a s s a n d t h e a m o u n t o f i n d i v i d u a l n u t r i e n t s i n p a r t i c u l a r d i a m e t e r c l a s s e s w e r e t h e r e f o r e c a l c u l a t e d i n t h e f o l l o w i n g m a n n e r : c l a s s e s 1 a n d 2 w e r e c a l c u -l a t e d w i t h c l a s s t h r e e ( T a b l e 1 1 ) u s i n g b i o m a s s a n d n u t r i e n t c o n t e n t d a t a o f c l a s s t h r e e , B i o m a s s a n d n u t r i e n t c o n t e n t f o r c l a s s e s 4, 5 a n d 6 w e r e o b t a i n e d b y m u l t i p l y i n g n u m b e r s o f t r e e s i n e a c h . c l a s s b y c o r r e s p o n d i n g b i o m a s s a n d n u t r i e n t c o n t e n t o f t h e s a m p l e d t r e e f o r e a c h p a r t i c u l a r c l a s s . C l a s s e s 8, 9, 1 0 a n d 1 1 w e r e c a l c u l a t e d w i t h c l a s s 7, u s i n g b i o m a s s a n d n u t r i e n t c o n t e n t d a t a o f c l a s s 7. I t i s o b v i o u s t h a t b i o m a s s a n d n u t r i e n t c o n t e n t w e r e o v e r - -e s t i m a t e d f o r c l a s s e s 1 a n d 2 a n d w e r e u n d e r - e s t i m a t e d f o r c l a s s e s 8, 9, 1 0 a n d 1 1 , A l l t o g e t h e r , t h e b i o m a s s a n d n u t r i e n t c o n t e n t f o r s m a l l b r a n c h e s o f m o u n t a i n h e m l o c k a r e p r o b a b l y s l i g h t l y u n d e r - e s t i m a t e d b e c a u s e a l t h o u g h t h e n u m b e r o f t r e e s w h i c h w e r e u n d e r - e s t i m a t e d a n d o v e r - e s t i m a t e d i s a p -p r o x i m a t e l y t h e s a me ( 1 0 a n d 9, r e s p e c t i v e l y ) , t h e u n d e r - e s t i m a t i o n w a s f o r a m u c h , w i d e r c l a s s r a n g e t h a n t h e o v e r - e s t i m a t i o n . No s i g n i f i c a n t r e g r e s s i o n e q u a t i o n s w e r e f o u n d f o r b i o m a s s o f y e l l o w c e d a r . T h e n u m b e r o f s a m p l e d t r e e s ( 4 ) w a s n o t s u f f i c i e n t t o f i n d s i g n i f i c a n t c o r r e l a t i o n s b e t w e e n m e n s u r a t i o n a l p a r a m e t e r s a n d t r e e b i o m a s s . 47 T h e a m o u n t o f b i o m a s s a n d t h e a m o u n t o f i n d i v i d u a l n u t r i e n t s i n p a r t i c u l a r d i a m e t e r c l a s s e s w e r e , t h e r e f o r e , c a l c u l a t e d i n t h e f o l l o w i n g m a n n e r : f o r c l a s s 1 ( T a b l e 1 1 ) t h e a v e r a g e a m o u n t o f b i o m a s s a n d n u t r i e n t s f o r p a r t i c u -l a r t r e e c o m p o n e n t s w a s c a l c u l a t e d f r o m t w o s a m p l e d t r e e s f o r t h i s c l a s s . A v e r a g e v a l u e s w e r e m u l t i p l i e d b y t h e n u m b e r o f t r e e s i n c l a s s 1. F o r c l a s s 2 a v e r a g e a m o u n t o f b i o m a s s a n d n u t r i e n t f o r i n d i v i d u a l t r e e c o m p o n e n t s w e r e c a l c u l a t e d f r o m t h e a v e r a g e o f t h e l a r g e s t t r e e i n s i z e c l a s s 1 a n d t h e t r e e i n s i z e c l a s s 3. A v e r a g e v a l u e s w e r e a g a i n m u l t i p l i e d b y t h e n u m b e r o f t r e e s i n c l a s s 2, F o r c l a s s 3 , b i o m a s s a n d n u t r i e n t v a l u e s f o r t h e s a m p l e t r e e w e r e s i m p l y m u l t i p l i e d b y t h e n u m b e r o f t r e e s i n c l a s s 3, T h e same wa s d o n e f o r c l a s s 5. L i m i t a t i o n o n D a t a I n t e r p r e t a t i o n : T h e n u m b e r o f o b s e r v a t i o n s u s e d f o r d e v e l o p i n g t h e r e g r e s s i o n e q u a t i o n s w a s v e r y l i m i t e d ( 5 t o 1 2 o b s e r v a t i o n s p e r s p e c i e s ) d u e t o t h e l a r g e s i z e o f many o f t h e t r e e s s a m p l e d a n d t h e s h o r t p e r i o d o f f i e l d w o r k i m p o s e d b y s n o w c o n d i t i o n s . T h e r e g r e s s i o n e q u a t i o n s a r e , t h e r e f o r e , n o t a s p r e c i s e a s t h e y w o u l d h a v e b e e n h a d i t b e e n p o s s i b l e t o s a m p l e a g r e a t e r n u m b e r o f t r e e s ( 3 0 o r m o r e r e s i d u a l d e g r e e s o f f r e e d o m a r e u s u a l l y s u g -g e s t e d t o o b t a i n a s t a b l e r e g r e s s i o n m o d e l ) . R e g r e s s i o n e q u a t i o n s w e r e e x t r a p o l a t e d t o e s t i m a t e t h e b i o m a s s o f t h e f e w t r e e s w h i c h o c c u r r e d o u t s i d e o f t h e r a n g e o f t r e e s i z e s t h a t w e r e s a m p l e d . I am a w a r e o f t h e d a n g e r s o f u s i n g r e g r e s s i o n s t o e s t i m a t e v a l u e s o u t s i d e o f t h e r a n g e o f s a m p l i n g , b u t b e c a u s e o f t h e l i m i t a t i o n s i m p o s e d b y t h e f i e l d p r o b l e m s t h e r e w a s n o o t h e r 4 8 r e c o u r s e . Due t o t h e c o m p l e x i t y o f b i o m a s s c a l c u l a t i o n a n d i t s c o m b i n a t i o n w i t h n u t r i e n t c o n c e n t r a t i o n d a t a , t h e a m o u n t o f m a c r o n u t r i e n t s i m m o b i l i z e d i n t r e e b i o m a s s p e r h e c t a r e s h o u l d b e c o n s i d e r e d m o r e a s a n a p p r o x i m a t i o n t h a n a s a n e x a c t n u m b e r . M i s s i n g D a t a a n d A d j u s t m e n t o f I n c o r r e c t D a t a : D u r i n g t r a n s p o r t a t i o n f r o m t h e s a m p l e p l o t s t o t h e l a b o r a t o r y o n e b i o m a s s s a m p l e a n d t h r e e c h e m i c a l a n a l y s i s ''samples w e r e l o s t . B e c a u s e o f t h e n a t u r e o f t h e c o m p u t e r p r o g r a m f o r t h e c a l c u l a t i o n a d r y b i o m a s s a n d n u t r i e n t c o n t e n t ( A p p e n d i c e s 10 a n d 1 1 ) a c o m p l e t e s e t o f s p e c i f i c g r a v i t y d a t a a n d n u t r i e n t c o n c e n t r a t i o n d a t a w a s r e q u i r e d . T h e l o s t b i o m a s s s a m p l e w a s d i s c #10, t r e e # 4 1 , S p e c i f i c g r a v i t y o f d i s c #9 o f t h e same t r e e w a s u s e d t o r e p l a c e t h e m i s s i n g d a t a . T h e l o s t c h e m i c a l s a m p l e s w e r e a d i s c f r o m a t r e e b o l e , a b r a n c h s a m p l e a n d a f o l i a g e s a m p l e . D a t a w e r e r e p l a c e d b y c o r r e s p o n d i n g d a t a f r o m t h e s a m p l e t r e e o f t h e same s p e c i e s w i t h t h e m o s t s i m i l a r b i o m a s s . A p p r o x i m a t e l y 1 0 p e r c e n t o f t h e s p e c i f i c g r a v i t y d a t a w e r e o u t o f t h e r a n g e c i t e d i n t h e l i t e r a t u r e f o r t h e s e p a r t i c u l a r s p e c i e s . Wood s p e c i f i c g r a v i t y d a t a w i t h i n 9 5 p e r c e n t c o n f i d e n c e i n t e r v a l f o r P a c i f i c w i l v e r f i r i s 0.36 ± 0 . 0 7 2 g / c m 3 , f o r m o u n t a i n h e m l o c k i t i s 0 , 4 1 ± 0 . 0 8 2 g/cm^ ( F o r e s t r y H a n d b o o k ) . D a t a w h i c h w e r e t h r e e t i m e s s t a n d a r d d e v i a t i o n o u t o f t h i s r a n g e w e r e c o r r e c t e d t o t h e m e a n o f t h e v a l u e s f o r t h e t w o n e i g h b o u r i n g d i s c s w i t h i n t h e same t r e e . T h i s m e t h o d w a s s e l e c t e d , e v e n t h o u g h f r o m s t a t i s t i c a l s t a n d p o i n t i t i s n o t t h e m o s t a p p r o p r i a t e o n e b e c a u s e t h e s p e c i f i c g r a v i t y d a t a d i d n o t s h o w a n y t r e n d a n d b e c a u s e t h e s i m p l i c i t y o f t h e c o r r e c t i o n . 5 0 V R E S U L T S AND D I S C U S S I O N B i o m a s s D 1 s t r i b u t i o n : ^ B o t h s t a n d s c a n b e c h a r a c t e r i z e d a s m a t u r e o r o v e r - m a t u r e ( m e a n a g e : S q u a m i s h - 4 1 7 y e a r s ; H a n e y - 2 4 8 y e a r s ) a n d t h e r e f o r e t h e y a r e b o t h i n a c o m p a r a b l e p h y s i o l o g i c a l s t a g e . T h e r e l a t i v e d i s t r i b u t i o n o f b i o m a s s among d i f f e r e n t t r e e c o m p o n e n t s b e t w e e n b o t h s t a n d s s h o u l d t h e r e f o r e b e m o r e a f u n c t i o n o f s i t e a n d e l e v a t i o n t h a n d i f f e r e n c e i n s t a n d a g e . P e r h a p s t h e m o s t d r a m a t i c d i f f e r e n c e b e t w e e n t h e t w o p l o t s i s t h e f a c t t h a t t h e 3 q u a m i s h s a m p l e p l o t h a s a b o u t e i g h t t i m e s a s m u c h a b o v e -g r o u n d t r e e b i o m a s s p e r h e c t a r e a s t h e H a n e y p l o t . T h e d i s t r i b u t i o n o f t h i s b i o m a s s b e t w e e n d i f f e r e n t c o m p o n e n t s a l s o v a r i e s b e t w e e n t h e t w o s i t e s ( T a b l e 1 7 ) . T h e p r o p o r t i o n o f b i o m a s s o n t h e H a n e y p l o t t o t h e b i o m a s s o n t h e S q u a m i s h p l o t e x h i b i t s t h e r a t i o 1 t o 9.3 f o r s t e m w o o d , 1 t o 1 0 . 8 f o r s t e m b a r k , 1 t o 6.6 f o r b i g b r a n c h e s , 1 t o 1.8 f o r s m a l l b r a n c h e s , 1 t o 6.4 f o r t w i g s a n d f o l i a g e , 1 t o 9.5 f o r t r e e b o l e s , 1 t o 4.7 f o r t r e e c r o w n s , a n d 1 t o 8.2 f o r t o t a l a b o v e - g r o u n d t r e e b i o m a s s . T h e g r e a t e s t d i f f e r e n c e i s i n t h e b i o m a s s o f s t e m b a r k a n d s t e m w o o d , a n d t h e l e a s t i s i n t h e b i o m a s s o f s m a l l b r a n c h e s . T h e b i o m a s s o f t r e e c r o w n s ( 1 : 4.7) s h o w s c o n s i d e r a b l y l e s s d i f f e r e n c e t h a n t h e b i o m a s s o f t r e e b o l e s ( 1 : 9 . 5 ) . T h e d i s t r i b u t i o n o f b i o m a s s among d i f f e r e n t t r e e c o m p o n e n t s i s 1 - Some o f t h e m a t e r i a l i n t h i s s e c t i o n w a s p r e s e n t e d a t t h e I U FRO m e e t i n g i n V a n c o u v e r , B. C. i n A u g u s t , 1 9 7 3 . T A B L E 1 7 : B i o m a s s d i s t r i b u t i o n f o r f o u r t r e e s p e c i e s i M o u n t a i n h e m l o c k = 7 4 ; P a c i f i c s i l v e r f i r = T r e e S p e c i e s Wood B a r k B i g B r a n c h S m a l l B r a n c h M o u n t a i n h e m l o c k 9 1 3 7 8 2 7 8 2 6 1 0 4 8 6 2 2 7 1 P a c i f i c s i l v e r f i r 2 2 7 5 6 1 5 0 1 6 7 2 5 0 8 4 8 1 7 2 W e s t e r n h e m l o c k 2 7 3 4 8 6 5 9 5 4 6 6 8 5 2 7 3 Y e l l o w c e d a r 6 9 3 4 5 9 8 6 7 5 4 1 1 S q u a m i s h P l o t T o t a l 3 1 8 9 3 9 7 7 9 9 3 3 5 5 7 0 1 0 4 4 3 H a n e y P l o t T o t a l 3 4 2 8 2 7 1 9 3 5 3 4 3 5 6 8 4 n t h e S q u a m i s h a n d H a n e y s a m p l e p l o t s ( i n k g / h a ) . n 1 8 2 ; W e s t e r n h e m l o c k = 6 7 ; Y e l l o w c e d a r = 1 5 . T w i g F o l i a g e T w i g & F o l . T r e e B o l e C r o w n W h o l e T r e e 3 3 2 7 3 3 4 5 5 9 0 5 1 1 9 2 0 3 1 8 6 6 2 1 3 7 8 6 5 — — 1 8 4 5 5 2 7 7 7 2 8 5 1 7 1 1 3 2 9 4 4 0 1 4 5 4 1 9 7 5 3 0 8 4 3 3 9 4 4 1 3 0 2 5 4 6 9 7 0 — — 7 1 3 7 5 2 3 1 7 9 9 9 3 2 2 — — 2 4 3 6 0 3 9 6 9 3 1 7 0 3 7 3 4 6 7 3 0 5 3 7 9 7 4 1 4 6 7 1 4 8 2 4 5 6 2 9 2 5 2 r a t h e r s i m i l a r f o r a l l f o u r s p e c i e s w i t h t h e e x c e p t i o n o f b i o m a s s o f s m a l l b r a n c h e s ( T a b l e 18)_, T h e p r o p o r t i o n o f s t e m w o o d v a r i e s f r o m 6 0 , 7 p e r c e n t o f t o t a l a b o v e - g r o u n d t r e e b i o m a s s f o r m o u n t a i n h e m l o c k t o 7 1 . 8 p e r c e n t f o r y e l l o w c e d a r . T h e p r o p o r t i o n o f b a r k v a r i e s f r o m 6,4 p e r c e n t f o r y e l l o w c e d a r t o 1 7 . 9 p e r c e n t f o r m o u n t a i n h e m l o c k . T h e p r o p o r t i o n o f s m a l l b r a n c h e s i s a b o u t s i x t i m e s g r e a t e r f o r w e s t e r n h e m l o c k ( 1 1 . 2 p e r c e n t ) t h a n f o r m o u n t a i n h e m l o c k ( 1 . 9 p e r c e n t ) . T h e r e m a i n i n g t w o s p e c i e s h a v e i n t e r m e d i a t e p r o p o r t i o n s o f s m a l l b r a n c h e s . T r e e b o l e a c c o u n t s f o r 74 t o 85 p e r c e n t o f t h e w h o l e t r e e b i o m a s s ; t r e e c r o w n f o r 1 5 t o 26 p e r c e n t . T h e b i o m a s s o f p a r t i c u l a r t r e e c o m p o n e n t s a s w e l l a s t o t a l a b o v e -g r o u n d t r e e b i o m a s s c a n b e d e s c r i b e d b y s i m p l e l o g a r i t h m i c r e g r e s s i o n e q u a -t i o n s o r s i m p l e l i n e a r r e g r e s s i o n e q u a t i o n s u s i n g m e n s u r a t i o n a l t r e e p a r a -m e t e r s a s s i m p l e ( e . g . D) o r c o m b i n e d ( e . g . D 2.H) i n d e p e n d e n t v a r i a b l e s ( A p p e n d i x 3 ) . I n t h i s s t u d y l o g a r i t h m i c r e g r e s s i o n e q u a t i o n s w i t h t h e h i g h e s t c o e f f i c i e n t o f d e t e r m i n a t i o n ( r 2 . 1 0 0 ) w e r e u s e d f o r p r e d i c t i o n o f b i o m a s s f o r a l l t r e e s o n t h e s a m p l e p l o t s . I n t h e f o l l o w i n g d i s c u s s i o n b o t h l o g a r i t h m i c a n d l i n e a r e q u a t i o n s a r e d i s c u s s e d , b u t t h e y w e r e n o t s t a -t i s t i c a l l y c o m p a r e d . T h e d e c i s i o n t o u s e l o g a r i t h m i c e q u a t i o n s w a s made b e c a u s e o f t h e i r b e t t e r f i t f o r v e r y s m a l l a n d v e r y b i g t r e e s . W h o l e t r e e b i o m a s s a n d w o o d b i o m a s s : T h e b e s t e s t i m a t o r o f t h e o t r e e b i o m a s s a n d w o o d b i o m a s s i s t h e c o m b i n e d v a r i a b l e D ,H o r D a l o n e , i n b o t h t h e l i n e a r a n d l o g a r i t h m i c e q u a t i o n . T A B L E 1 8 : A v e r a g e b i o m a s s p r o p o r t i o n s o f d i f f e r e n t t r e e c o m p o n e n t s f o r f o u r t r e e s p e c i e s o n S q u a m i s h a n d H a n e y s a m p l e p l o t s ( i n p e r c e n t ) . n ; M o u n t a i n h e m l o c k = 5 ; P a c i f i c s i l v e r f i r = 7; W e s t e r n h e m l o c k = 8; Y e l l o w c e d a r = 4. S q u a m i s h S q u a m i s h H a n e y H a n e y T r e e C o m p o n e n t P a c i f i c s i l v e r f i r M o u n t a i n h e m l o c k W e s t e r n h e m l o c k Y e l l o w c e d a r S t e m w o o d R a n g e 5 9 . 6 - 7 0 . 5 6 4 . 1 - 6 9 . 2 5 4 . 8 - 6 8 . 1 6 3 . 5 - 7 7 . 3 M e a n 6 6 . 3 6 7 . 2 6 0 . 8 7 1 . 7 S t e m b a r k R a n g e 9 . 5 - 1 8 . 8 1 7 . 0 - 1 9 . 1 9 . 8 - 1 6 . 4 5 . 4 - 8 . 2 M e a n 1 3 . 9 1 7 . 9 1 3 . 3 6.4 B i g b r a n c h e s R a n g e 4 . 4 - 1 2 . 4 5 . 2 - 9 . 6 0 . 6 - 1 3 . 2 2 . 3 - 9 . 7 M e a n 8.1 6.9 6.6 5.9 S m a l l b r a n c h e s R a n g e 1 . 6 ^ 6 . 5 1.1-2.4 7 . 0 - 1 5 . 9 3 . 4 - 1 0 . 4 M e a n 3.9 1.9 1 1 . 3 6.2 T w i g s R a n g e 2 . 2 - 3 . 2 1 . 7 - 6 . 5 M e a n 2.5 3.3 F o l i a g e R a n g e 2 . 9 - 4 . 5 2 . 7 - 7 . 0 M e a n 3.6 4.7 T w i g & F o l i a g e R a n g e 4 . 5 - 1 1 . 9 5 . 1 - 7 . 6 5 . 9 - 1 0 . 0 6 . 2 - 1 6 . 4 M e a n 7.8 6.1 8.0 9.8 B o l e s M e a n 8 0 . 2 8 5 . 1 7 4 . 1 7 8 . 1 C r o w n s M e a n 1 9 . 8 1 4 . 9 2 5 . 9 2 1 . 9 54 B a r k b i o m a s s ; B a r k b i o m a s s i s m a i n l y d e p e n d e n t o n t h e b o l e 2 d i a m e t e r a n d t h e r e f o r e B A , D o r D ,H a r e t h e b e s t i n d e p e n d e n t v a r i a b l e s i n b o t h t h e l i n e a r a n d l o g a r i t h m i c e q u a t i o n s . B i g b r a n c h e s b i o m a s s : On t h e S q u a m i s h p l o t t h e b i o m a s s o f b i g b r a n c h e s i s d e p e n d e n t o n t r e e d i a m e t e r , t r e e l e n g t h , c r o w n d i a m e t e r a n d i n t h e l o g a r i t h m i c e q u a t i o n , o n c r o w n l e n g t h . I n b o t h t h e l i n e a r a n d l o g a r -i t h m i c e q u a t i o n f o r h e m l o c k , t h e m o s t i m p o r t a n t i n d e p e n d e n t v a r i a b l e i s D 2.H,CW, w h i l e f o r P a c i f i c s i l v e r f i r I t i s D.CL i n t h e l o g a r i t h m i c e q u a -t i o n , a n d BA o r D i n t h e l i n e a r e q u a t i o n . On t h e H a n e y p l o t , t h e b e s t i n d e p e n d e n t v a r i a b l e f o r h e m l o c k i n t h e l i n e a r e q u a t i o n i s D.CL, w h i l e i n t h e l o g a r i t h m i c e q u a t i o n i t i s D a l o n e . S m a l l b r a n c h e s b i o m a s s : B i o m a s s o f s m a l l b r a n c h e s f o r h e m l o c k o n t h e S q u a m i s h p l o t d o e s n o t s h o w a n y r e l a t i o n s h i p w i t h t h e i n d e p e n d e n t v a r i -a b l e s t e s t e d i n e i t h e r t h e l i n e a r o r t h e l o g a r i t h m i c e q u a t i o n s . T h e b i o m a s s o f s m a l l b r a n c h e s f o r P a c i f i c s i l v e r f i r o n t h e S q u a m i s h p l o t i s b e s t d e -s c r i b e d i n t h e l i n e a r e q u a t i o n b y e i t h e r t r e e l e n g t h H o r t r e e d i a m e t e r D, o r b y t h e c o m b i n a t i o n o f t r e e d i a m e t e r a n d c r o w n l e n g t h (D + C L ) . I n t h e l o g a r i t h m i c e q u a t i o n b i o m a s s o f s m a l l b r a n c h e s i s b e s t d e s c r i b e d b y t h e l o g -a r i t h m o f t r e e d i a m e t e r a n d c r o w n l e n g t h b y e i t h e r l o g D.CL o r l o g D . C L 2 . B i o m a s s o f s m a l l b r a n c h e s f o r h e m l o c k o n t h e H a n e y p l o t i s b e s t d e s c r i b e d i n t h e l i n e a r e q u a t i o n b y t r e e l e n g t h ( H ) a l o n e o r b y t h e c o m b i n a t i o n o f d . b . h . a n d c r o w n l e n g t h (D + C L ) . I n t h e l o g a r i t h m i c e q u a t i o n s t h e m o s t s i g n i f i c a n t i n d e p e n d e n t v a r i a b l e i s t h e c o m b i n a t i o n o f d . b . h , , t r e e l e n g t h 5 5 a n d c r o w n w i d t h (D Z.H,CW)> T w i g s a n d f o l i a g e - b i o m a s s : T h e b i o m a s s o f t w i g s a n d f o l i a g e f o r b o t h s p e c i e s i n S q u a m i s h i s b e s t d e s c r i b e d i n t h e l i n e a r e q u a t i o n b y t r e e d i a m e t e r i n c o m b i n a t i o n w i t h c r o w n l e n g t h ( D . C L ) o r b y t r e e d i a m e t e r ( D ) a l o n e . I n t h e l o g a r i t h m i c e q u a t i o n , t h e m o s t s i g n i f i c a n t i n d e p e n d e n t v a r -i a b l e i s t h e c o m b i n a t i o n o f t r e e d i a m e t e r w i t h t r e e l e n g t h ( D 2 , H ) f o r m o u n t a i n h e m l o c k , a n d t r e e d i a m e t e r w i t h c r o w n l e n g t h ( D . C L ) f o r P a c i f i c s i l v e r f i r . F o r h e m l o c k i n H a n e y , t r e e l e n g t h a l o n e ( H ) o r t h e c o m b i n a t i o n o f t r e e d i a m e t e r a n d c r o w n l e n g t h ( D . C L ) a r e t h e b e s t i n d e p e n d e n t v a r i a b l e s i n l i n e a r e q u a t i o n s . I n t h e l o g a r i t h m i c e q u a t i o n , t h e c o m b i n a t i o n o f t r e e d i a m e t e r w i t h c r o w n l e n g t h ( D . C L ) i s t h e m o s t s i g n i f i c a n t i n d e p e n d e n t v a r i -a b l e . T w i g b i o m a s s ' • F o r m o u n t a i n h e m l o c k o n t h e S q u a m i s h p l o t , t h e c o m b i n a t i o n o f t r e e d i a m e t e r a n d c r o w n l e n g t h ( D . C L ) o r t r e e d i a m e t e r a l o n e (D) i s t h e b e s t i n d e p e n d e n t v a r i a b l e i n t h e l i n e a r e q u a t i o n . I n t h e l o g a r i t h m i c e q u a t i o n , t r e e d i a m e t e r w i t h c r o w n l e n g t h o r t r e e d i a m e t e r w i t h c r o w n l e n g t h s q u a r e d ( D . C L o r D . C L 2 ) i s t h e b e s t i n d e p e n d e n t v a r i a b l e . F o r h e m l o c k o n t h e H a n e y p l o t , c r o w n w i d t h s q u a r e d ( C W 2 ) i s t h e b e s t i n d e p e n -d e n t v a r i a b l e i n t h e l i n e a r e q u a t i o n , w h i l e t h e c o m b i n a t i o n o f t r e e b a s a l a r e a a n d t r e e l e n g t h w i t h c r o w n d i a m e t e r a n d l e n g t h ( B A . H ) + (CL.CW) i s t h e b e s t i n d e p e n d e n t v a r i a b l e i n t h e l o g a r i t h m i c e q u a t i o n . F o l i a g e b i o m a s s : F o r m o u n t a i n h e m l o c k o n t h e S q u a m i s h p l o t e i t h e r 56 t h e c o m b i n a t i o n o f t r e e d i a m e t e r w i t h c r o w n l e n g t h ( D . C L ) o r t r e e d i a m e t e r a l o n e i s t h e b e s t i n d e p e n d e n t v a r i a b l e i n t h e l i n e a r e q u a t i o n . I n t h e l o g a r i t h m i c e q u a t i o n , t r e e d i a m e t e r w i t h t r e e l e n g t h (D ,H) a p p e a r s t o b e t h e m o s t s i g n i f i c a n t i n d e p e n d e n t v a r i a b l e . F o r h e m l o c k o n t h e H a n e y p l o t , t r e e l e n g t h a l o n e (II) o r t r e e d i a m e t e r w i t h c r o w n l e n g t h ( D . C L ) a r e t h e m o s t s i g n i f i c a n t i n d e p e n d e n t v a r i a b l e s i n t h e l i n e a r r e g r e s s i o n e q u a t i o n . I n t h e l o g a r t h m i c e q u a t i o n t r e e d i a m e t e r i n c o m b i n a t i o n w i t h c r o w n l e n g t h ( D . C L ) i s t h e b e s t i n d e p e n d e n t v a r i a b l e . I n s u m m a r y , f r o m t h e a b o v e d i s c u s s i o n , i t w o u l d a p p e a r t h a t t h e e q u a t i o n l o g Y = b + b1 l o g D i s a m o d e l w h i c h c a n b e u s e d f o r p r e d i c t i o n o f t h e w h o l e t r e e b i o m a s s a s w e l l a s t h e b i o m a s s o f a n y i n d i v i d u a l t r e e c o m p o n e n t . T h i s i s i n a g r e e m e n t w i t h a n u m b e r o f o t h e r p u b l i c a t i o n s l i s t e d b y M a d g w i c k ( 1 9 6 3 ) . T h e m o d e l f i t s w e l l f o r b o t h s p e c i e s o n t h e S q u a m i s h p l o t ( w i t h t h e e x c e p t i o n o f s m a l l b r a n c h e s f o r m o u n t a i n h e m l o c k ) , a n d f o r h e m l o c k o n t h e H a n e y p l o t . A l o g a r i t h m i c e q u a t i o n i n t h e f o r m l o g ( w e i g h t ) = a + b ( l o g D ) w a s u s e d b y M o r r i s o n ( 1 9 7 2 ) . f o r j a c k p i n e s t a n d s i n N o r t h e r n O n t a r i o . On t h e o t h e r h a n d , K u r u c z ( 1 9 6 9 ) . u s e d l i n e a r r e g r e s s i o n e q u a t i o n s f o r b i o m a s s p r e d i c t i o n o f D o u g l a s - f i r , w e s t e r n h e m l o c k a n d w e s t e r n r e d c e d a r i n U.B.C. r e s e a r c h f o r e s t i n H a n e y . 57 N u t r i e n t D i s t r l h u t l d i K W i t h i n - t h e - T r e e L a y e r ; A v e r a g e n u t r i e n t c o n c e n t r a t i o n s f o r p a r t i c u l a r t r e e c o m p o n e n t s f o r t h e f o u r t r e e s p e c i e s a r e s u m m a r i z e d i n T a b l e s 1 9 , 2 0 , 2 1 a n d 2 2 . N u t r i e n t c o n c e n t r a t i o n s f o r i n d i v i d u a l s a m p l e t r e e s a r e p r e s e n t e d i n A p p e n -d i c e s 6 a n d 7. I w o u l d l i k e t o d i s c u s s f i r s t t h e c o n c e n t r a t i o n o f s p e c i f i c e l e m e n t s i n p a r t i c u l a r t r e e c o m p o n e n t s f o r e a c h t r e e s p e c i e s s e p a r a t e l y a n d t h e n c o m p a r e n u t r i e n t c o n c e n t r a t i o n s b e t w e e n s p e c i e s . C o n c e n t r a t i o n s o f n u t r i e n t s w i t h i n d i f f e r e n t b i o m a s s c o m p o n e n t s o f P a c i f i c s i l v e r f i r w e r e f o u n d t o b e i n t h e s e q u e n c e stemwood<C b i g b r a n c h e s <C s m a l l b r a n c h e s <C stembark<C t w i g s a n d f o l i a g e f o r a l l n u t r i e n t s w i t h t h e e x c e p t i o n o f c a l c i u m . C o n c e n t r a t i o n o f c a l c i u m i s h i g h e r i n b a r k t h a n i n t w i g s a n d f o l i a g e . N u t r i e n t c o n c e n t r a t i o n s i n p a r t i c u l a r t r e e c o m -p o n e n t s w e r e f o u n d t o b e i n t h e s e q u e n c e : w o o d C a ^ N £ = K > M g > P b a r k C a > N > K > P > M g b i g b r a n c h e s C a > N > K > - Mg £z P s m a l l b r a n c h e s C a > N > K > M g « r P t w i g s a n d f o l i a g e N ;> C a >• K >• P >• Mg c o n e s N > K > P > M g > C a N u t r i e n t c o n c e n t r a t i o n s w i t h i n m o u n t a i n h e m l o c k w e r e f o u n d t o b e i n t h e s e q u e n c e : s t e m w o o d < ^ b i g b r a n c h e s <^ s m a l l b r a n c h e s <^ s t e m b a r k < ^ twigs<C f o l i a g e f o r a l l n u t r i e n t s e x c e p t p h o s p h o r o u s a n d c a l c i u m . T h e T A B L E 1 9 : M e a n c o n c e n t r a t i o n ( p e r c e n t a g e o v e n d r y w e i g h t ) . , o n N, P, K, C a a n d Mg i n c o m p o n e n t s a n d s u b - c o m p o n e n t s o f P a c i f i c s i l v e r f i r s a m p l e t r e e s . C o m p o n e n t N P K C a Mg Wood 1 0.05 0 , 0 1 0.06 0,05 0 , 0 1 ( d i s c n o , f r o m 3 0.04 0 . 0 1 0.05 0.07 0 . 0 1 t h e b o t t o m ) 5 0.05 0 . 0 1 0.06 0,06 0 . 0 1 7 0,05 0 . 0 1 0.06 0,06 0 . 0 1 9 0.06 0 . 0 1 0.07 0.06 0 . 0 1 1 0 0.07 0.02 0.07 0.05 0 . 0 1 B a r k 1 0.25 0.04 0 . 1 1 0.49 0.02 ( d i s c n o , f r o m 3 0 . 2 1 0.04 0.12 0.48 0.03 t h e b o t t o m ) 5 0.22 0.04 0.13 0.47 0.03 7 0.24 0.05 0.14 0.46 0.04 9 0 . 3 1 0.06 0.18 , 0.50 0.04 1 0 0.40 0.08 0.24 0.49 0.04 B i g b r a n c h e s 0.15 0.02 0.09 0 . 3 1 0.02 S m a l l b r a n c h e s 0.22 0.03 0.12 0.29 0.03 T w i g s & F o l i a g e 1 0.73 0.10 0.36 0.40 0.08 ( u p p e r , m i d d l e , 2 0.72 0.10 0 . 3 1 0.46 0.07 l o w e r c r o w n ) 3 0.76 0.10 0.32 0.46 0.06 C o n e s 1.35 0.24 0.70 0.04 ' 0.08 D a t a f o r w o o d a n d b a r k b a s e d o n 1 2 s a m p l e d t r e e s . D a t a f o r c r o w n c o m p o n e n t s b a s e d o n 7 s a m p l e d t r e e s . D a t a f o r c o n e s b a s e d o n 6 s a m p l e d t r e e s . T A B L E 2 0 : M e a n c o n c e n t r a t i o n ( p e r c e n t a g e o v e n d r y w e i g h t ) , o f N, P, K, a n d Mg i n c o m p o n e n t s a n d s u b - c o m p o n e n t s o f m o u n t a i n h e m l o c k s a m p l e t r e e s . C o m p o n e n t N P K. C a Mg Wood 1 0,06 0.02 0.07 0,06 0,02 ( d i s c n o , f r o m 3 0.05 0.01 0.06 0.07 0.02 t h e b o t t o m ) 5 0,05 0.01 0,06 0,07 0.02 7 0.05 0,02 0.07 0.08 0.02 9 0.06 0.02 0.07 0.06 0.02 10 0.07 0.02 0.07 o;o6 0.02 B a r k 1 0.22 0.07 0.09 0.40 0.02 ( d i s c n o , f r o m 3 0.15 0.06 0.09 0.31 0.02 t h e b o t t o m ) 5 0.17 0.05 0.09 0.31 0.02 7 0.17 0.05 0.09 0.24 0.02 9 0.22 0.06 0.12 0.25 0.02 1 0 0.28 0.07 0.16 0.27 0.03 B i g b r a n c h e s 0.12 0.03 0.07 0.16 0.02 S m a l l b r a n c h e s 0.17 0.04 0.09 0.18 0.03 T w i g s 1 0.43 0.06 0.16 0.19 0.04 ( u p p e r , m i d d l e , 2 0.39 0.06 0.15 0.18 0.04 l o w e r c r o w n ) 3 0.34 0.06 0.17 0.17 0.05 F o l i a g e 1 0.80 0.11 0.28 0.31 0.08 ( u p p e r , m i d d l e , 2 0.91 0.12 0 . 3 1 0.28 0.08 l o w e r c r o w n ) 3 0.83 0.11 0.31 0.32 0.08 C o n e s 0.92 0.17 0.32 0.03 0.05 D a t a f o r w o o d a n d b a r k b a s e d o n 8 s a m p l e d t r e e s . D a t a f o r c r o w n c o m p o n e n t s b a s e d o n 5 s a m p l e d t r e e s . D a t a f o r c o n e s b a s e d o n 1 s a m p l e d t r e e . 6 0 T A B L E 2 1 : M e a n c o n c e n t r a t i o n ( p e r c e n t a g e o v e n d r y w e i g h t ) , o f N , P, K, C a a n d Mg i n c o m p o n e n t s a n d s u b - c o m p o n e n t s o f w e s t e r n h e m l o c k s a m p l e t r e e s . C o m p o n e n t N P K C a Mg Wood 1 0.04 0.02 0.07 0.05 0 . 0 1 ( d i s c n o . f r o m 3 0.04 0 . 0 1 0.07 0.06 0 . 0 1 t h e b o t t o m ) 5 0,04 0 . 0 1 0.06 0.05 0 . 0 1 7 0.05 0 . 0 1 0.06 0.05 0 . 0 1 9 0.06 0.01 0.07 0.04 0 . 0 1 1 0 0.06 0 . 0 1 0.07 0.04 0 . 0 1 B a r k 1 0.15 0.03 0.08 0.40 0.02 ( d i s c n o . f r o m 3 0.16 0.03 0.12 0.44 0.02 t h e b o t t o m ) 5 0.18 0.03 0.14 ,0.38 0.02 7 0.22 0.04 0.16 0.30 0.03 9 0.27 0.05 0.19 0.29 0.03 1 0 0.32 0.06 0.22 0.29 0.04 B i g b r a n c h e s 0.10 0,02 OgOS 0.17 0.02 S m a l l b r a n c h e s 0.14 0.02 0.09 . 0.19 0.02 T w i g s 1 0.53 0.08 0 . 2 1 0.29 0.06 ( u p p e r , m i d d l e , 2 0.45 0.07 0.19 0.29 0.05 l o w e r c r o w n ) 3 0.40 0.06 0.17 0.28 0.05 F o l i a g e 1 0.76 0 . 1 1 0.37 0.39 0.10 ( u p p e r , m i d d l e , 2 0.74 0.12 0.33 0.48 0.12 l o w e r c r o w n ) 3 0.74 0.14 0.34 0.49. 0.12 A l l d a t a b a s e d o n 8 s a m p l e d t r e e s . 61 T A B L E 2 2 : M e a n c o n c e n t r a t i o n ( p e r c e n t a g e o v e n d r y w e i g h t ) , o f N, P, K, C a a n d Mg i n c o m p o n e n t s a n d s u b - c o m p o n e n t s o f y e l l o w c e d a r s a m p l e t r e e s . C o m p o n e n t N P K C a Mg Wood 1 0,05 0 . 0 1 0.05 0.12 0 . 0 1 ( d i s c n o . f r o m 3 0,06 0 . 0 1 0,05 0.12 0 . 0 1 t h e b o t t o m ) . 5 0.05 0 . 0 1 0.05 .0.11 0 . 0 1 7 0.07 0 . 0 1 0.05 0.09 0 . 0 1 9 0.07 0 . 0 1 0.06 0 . 1 1 0 . 0 1 1 0 0,07 0 . 0 1 0.06 0.12 0 . 0 1 B a r k 1 0.27 0.05 0.20 0.99 0.04 ( d i s c n o . f r o m 3 0.27 0,05 0.18 0.40 0.04 t h e b o t t o m ) 5 0.27 0.06 0.25 0.49 0.05 7 0.33 0.07 0.27 0 . 1 1 0.05 9 0.37 0.08 0.30 0.87 0.06 1 0 0,44 0.08 0.30 0.62 0.06 B i g b r a n c h e s 0.15 0.02 0.09 0.64 0.02 S m a l l b r a n c h e s 0.23 0.03 0.14 0 . 8 1 0.03 T w i g s & F o l i a g e 1 0.75 0.09 0.37 0.24 0.07 ( u p p e r , m i d d l e , 2 0.70 0.08 0.32 0.33 0.07 l o w e r c r o w n ) 3 0.77 0.08 0.36 0 . 5 3 0.08 A l l d a t a b a s e d o n 4 s a m p l e d t r e e s . 62 c o n c e n t r a t i o n o f P i n b a r k a n d t w i g s i s a p p r o x i m a t e l y t h e same ( t h e m a i n c o m p o n e n t o f t w i g s i s b a r k ) . , T h e c o n c e n t r a t i o n o f C a i s h i g h e r i n b a r k t h a n i n t w i g s a n d b a r k c o n c e n t r a t i o n i s a p p r o x i m a t e l y e q u a l t o f o l i a g e c o n -c e n t r a t i o n . N u t r i e n t c o n c e n t r a t i o n s i n p a r t i c u l a r t r e e c o m p o n e n t s a p p e a r t o b e i n t h e s e q u e n c e : w o o d C a « s N « ? K > P * = : M g b a r k C a > N > K > P > M g b i g b r a n c h e s C a > N > K > P « M g s m a l l b r a n c h e s C a < » N > K > P > M g t w i g s N > C a > K > P > Mg f o l i a g e N > C a > K > P > M g c o n e s N > K > P > Mg > C a N u t r i e n t c o n c e n t r a t i o n s w i t h i n w e s t e r n h e m l o c k w e r e f o u n d t o b e i n t h e s e q u e n c e : s t e m w o o d < b i g b r a n c h e s <C s m a l l b r a n c h e s < s t e m b a r k < t w i g s < f o l i a g e f o r a l l n u t r i e n t s w i t h t h e e x c e p t i o n o f c a l c i u m . T h e c o n c e n t r a t i o n o f c a l c i u m i s h i g h e r i n t h e b a r k t h a n i n t h e t w i g s . N u t r i e n t c o n c e n t r a t i o n s i n p a r t i c u l a r t r e e c o m p o n e n t s w e r e f o u n d t o b e i n t h e s e q u e n c e : w o o d K > C a ~ N > P ? » M g b a r k C a > N > K > P > Mg b i g b r a n c h e s C a > N > K > P «2Mg s m a l l b r a n c h e s C a > N > K > P ^ M g t w i g s N > C a > K > P > Mg f o l i a g e N > C a > K > P > M g 63 N u t r i e n t c o n c e n t r a t i o n s i n y e l l o w c e d a r w e r e f o u n d t o b e i n t h e s e q u e n c e : s t e m w o o d < f b i g b r a n c h e s < f s m a l l branches<C b a r k < ^ t w i g s a n d f o l -i a g e f o r a l l n u t r i e n t s w i t h t h e e x c e p t i o n o f c a l c i u m . T h e c o n c e n t r a t i o n o f c a l c i u m i s h i g h e r i n b a r k t h a n i n t w i g s a n d f o l i a g e , N u t r i e n t c o n c e n t r a -t i o n s i n p a r t i c u l a r t r e e c o m p o n e n t s a p p e a r t o b e i n t h e o r d e r : w o o d C a > N > K > P <srMg b a r k C a > N > K > P > Mg b i g b r a n c h e s C a > N > K > P ^ M g s m a l l b r a n c h e s C a > N > K > P ^  Mg t w i g s a n d f o l i a g e C a ^> N > K > P 5 » M g Y e l l o w c e d a r h a s c o n s i d e r a b l y h i g h e r a m o u n t s o f c a l c i u m i n a l l t r e e c o m p o n e n t s t h a n t h e o t h e r s p e c i e s . C o n c e n t r a t i o n s o f n u t r i e n t s i n s t e m w o o d do n o t s h o w a n y s i g n i f i -c a n t t r e n d b e t w e e n d i f f e r e n t d i s c s f r o m t h e b o t t o m t o t h e t o p o f t h e t r e e . C o n c e n t r a t i o n s o f n u t r i e n t s i n s t e m b a r k , h o w e v e r , d o s h o w a n i n c r e a s i n g t r e n d f r o m b o t t o m t o t o p o f t h e t r e e f o r a l l n u t r i e n t s e x c e p t c a l c i u m . C o n c e n t r a -t i o n o f c a l c i u m i n b a r k o f P a c i f i c s i l v e r f i r i s c o n s t a n t , w h i l e f o r t h e o t h e r t r e e s p e c i e s i t d e c r e a s e s f r o m t h e b o t t o m t o t h e t o p o f a t r e e . T h e f o u r t r e e s p e c i e s c a n b e a r r a n g e d i n d i f f e r e n t o r d e r a c c o r d i n g t o t h e n u t r i e n t c o n c e n t r a t i o n s i n v a r i o u s t r e e c o m p o n e n t s . A l l s p e c i e s a r e a p p r o x i m a t e l y e q u a l i n c o n c e n t r a t i o n o f a l l n u t r i e n t s i n s t e m w o o d , w i t h t h e e x c e p t i o n o f y e l l o w c e d a r w h i c h e x h i b i t s c o n s i d e r a b l y h i g h e r c a l c i u m c o n c e n -t r a t i o n . T h e f o u r s p e c i e s a p p e a r i n t h e f o l l o w i n g o r d e r a c c o r d i n g t o c o n -c e n t r a t i o n s o f d i f f e r e n t e l e m e n t s i n v a r i o u s t r e e c o m p o n e n t s . 64 B a r k N Y . c . > P . s . f . > W.h. > M.h. P • Y , c , « P . s . f . ;= M,h. > W,h. K Y , c , > P . s . f . « = W.h, > M , h . C a a n d Mg Y . c . > P . s . f , > W.h, > M.h. B i g N Y . c . ^ P . s . f . > M . h . > W,h, B r a n c h e s C a Y . c , > P . s . f , > W.h. M.h. P, K, Mg Y , c . s = P . s . f . « W.h. « H , h ( S m a l l N Y . c , » P . s . f . > W . h , M , h , B r a n c h e s P, Mg Y , c . a s - P . s . f . ~ W.h. zz. M.h. K, C a Y . c , > P , s . f , > W.h, <=sM,h, T w i g s a n d f o l i a g e c a n n o t b e c o m p a r e d b e c a u s e o f t h e n a t u r e o f s a m p l i n g a n d a n a l y s e s . F o r b o t h h e m l o c k s p e c i e s t w i g s a n d f o l i a g e w e r e s a m p l e d a n d a n a l y s e d s e p a r a t e l y , w h i l e f o r P a c i f i c s i l v e r f i r a n d y e l l o w c e d a r t w i g s a n d f o l i a g e w e r e n o t s e p a r a t e d a n d w e r e a n a l y s e d t o g e t h e r . I n s u m m a r i z i n g t h e d a t a o n n u t r i e n t d i s t r i b u t i o n i n t h e a b o v e - g r o u n d t r e e b i o m a s s , t h e f o l l o w i n g p o i n t s c a n b e m a d e : a ) y e l l o w c e d a r e x h i b i t s c o n -s i d e r a b l y h i g h e r c a l c i u m c o n c e n t r a t i o n s i n a l l t r e e c o m p o n e n t s t h a n o t h e r s p e c i e s : b ) P a c i f i c s i l v e r f i r e x h i b i t s h i g h e r n u t r i e n t c o n c e n t r a t i o n s f o r a l -m o s t a l l e l e m e n t s i n a l l t r e e c o m p o n e n t s t h a n b o t h h e m l o c k s p e c i e s ; c ) n u t r i -e n t c o n c e n t r a t i o n s o f m o u n t a i n h e m l o c k a r e a p p r o x i m a t e l y e q u a l t o t h e c o n c e n -t r a t i o n s o f w e s t e r n h e m l o c k , One m u s t k e e p i n m i n d t h a t t h e s i t e c o n d i t i o n s o f t h e t w o s a m p l e p l o t s w e r e c o n s i d e r a b l y d i f f e r e n t . T r e e s p e c i e s m i g h t a p -p e a r i n a d i f f e r e n t o r d e r i f t h e y w e r e g r o w n u n d e r u n i f o r m s i t e c o n d i t i o n s . C o n c e n t r a t i o n o f m a c r o n u t r i e n t s i n f o l i a g e o r f o l i a g e a n d t w i g s w i t h i n t h e u p p e r t h i r d o f t r e e c r o w n s i s s u m m a r i z e d i n T a b l e 2 3 , T h e d a t a c a n n o t b e c o m p a r e d w i t h f o l i a g e c o n c e n t r a t i o n s f r o m o t h e r p u b l i c a t i o n s b e c a u s e , a ) , t h e s a m p l e s w e r e o b t a i n e d r a n d o m l y f r o m a n y l o c a t i o n w i t h i n t h e u p p e r t h i r d o f t h e t r e e c r o w n ; b ) t h e s a m p l e s i n c l u d e d f o l i a g e o f a l l 65 T A B L E 2 3 : C o n c e n t r a t i o n o f m a c r o n u t r i e n t s i n f o l i a g e w i t h i n u p p e r t h i r d o f t r e e c r o w n ( i n p e r c e n t ) , 1 T r e e T r e e S p e c i e s N o . N P C a Mg K M o u n t a i n h e m l o c k 68 0.80 0,10 0.31 0.10 0.23 74 0.65 0.11 0.39 0.08 0.28 80 0.86 0.12 0.22 0.06 0.31 105 0.83 0.10 0.32 0.07 0.27 119 0.84 0.10 0.32 0.08 0.30 R a n g e 0.66- 0.10- 0.22- 0.06- 0.24-0.87 0.12 0.39 0.10 0. M e a n 0.80 0.11 0.31 0.08 0.28 P a c i f i c s i l v e r f i r 2 22 0.76 0.11 0.41 0.06 0.31 41 0.80 0.10 0.30 0.07 0.32 110 0.69 0.09 0.34 0.05 0.37 159 0.52 0.07 0.51 0.06 0.44 175 0.73 0.10 0.44 0.08 0.33 213 0.76 0.10 0.40 0.06 0.38 243 0.82 0.11 0.37 0.10 0.35 R a n g e 0.52- 0.07- 0.30- 0.04- 0.31-0.82 0.11 0.51 0.10 o.. M e a n 0.73 0.10 0.40 0.07 0.36 W e s t e r n h e m l o c k 50 0.73 0.12 0.43 0.11 0.33 54 0.74 0.10 0.49 0.11 0.37 65 0.82 0.10 0.47 0.09 0.47 69 0.76 0.12 0.33 0.11 0.34 72 0.79 0.11 0.32 0.07 0.43 103 0.76 0.12 0.36 0.13 0.34 104 0.62 0.12 0.41 0.13 0.42 108 0.87 0.12 0.28 0.09 0.23 R a n g e 0.62- 0.10- 0.28- 0.07- 0.23-0.87 0.12 0.49 0.13 0,< M e a n 0.76 0.11 0.39 0.10 0.37 Y e l l o w c e d a r 8 0.70 0,08 1.14 0.06 0.31 63 0.81 0.09 1.22 0.09 0.48 76 0.73 0.09 1.18 0.06 0.30 102 0.77 0.08 1.43 0.08 0.40 R a n g e 0.70- 0.08- 1.14- 0.06- 0.30-0.81 0.09 1.43 0.09 o.. M e a n 0.75 0.09 1.24 0.07 0.37 1 - D a t a f r o m s a m p l e s i n c l u d i n g s e v e r a l f o l i a r a g e c l a s s e s t a k e n b e t w e e n l a t e M ay a n d t h e e n d o f J u n e f o r y e l l o w c e d a r a n d w e s t e r n h e m l o c k , a n d b e t w e e n e a r l y J u l y a n d l a t e A u g u s t f o r P a c i f i c s i l v e r f i r a n d m o u n t a i n h e m l o c k . 2 - D a t a f o r P a c i f i c s i l v e r f i r a n d y e l l o w c e d a r a r e f o r t w i g s a n d f o l i a g e t o g e t h e r a n d t h e r e f o r e t h e v a l u e s a r e l o w e r . 66 a g e s , i n some c a s e s u p t o 2 0 y e a r s o l d ; c ) . f o l i a g e a n d t w i g s w e r e n o t s e p -a r a t e d f o r P a c i f i c s i l v e r f i r a n d y e l l o w c e d a r ; d ) . f o l i a g e s a m p l e s f r o m d i f f e r e n t t r e e s w e r e o b t a i n e d t h r o u g h a t i m e - p e r i o d o f a b o u t t w o m o n t h s . T h e d i s t r i b u t i o n o f f i v e m a c r o n u t r i e n t s ( i n k g / h a ) i n t h e d i f f e r -e n t t r e e c o m p o n e n t s i s s u m m a r i z e d f o r t h e S q u a m i s h s a m p l e p l o t i n T a b l e 24 a n d f o r t h e H a n e y s a m p l e p l o t i n T a b l e 2 5 , A s c a n b e s e e n , t h e a b o v e -g r o u n d t r e e b i o m a s s o n t h e S q u a m i s h s a m p l e p l o t c o n t a i n s a b o u t s e v e n a n d o n e - h a l f t i m e s a s m u c h m a c r o n u t r i e n t s p e r h e c t a r e a s o n t h e H a n e y s a m p l e p l o t . T h e r a t i o o f b i o m a s s o n t h e t w o p l o t s i s s i m i l a r t o t h e n u t r i e n t r a t i o , w i t h t h e S q u a m i s h s a m p l e p l o t h a v i n g a p p r o x i m a t e l y e i g h t t i m e s a s m u c h b i o m a s s a s t h e H a n e y s a m p l e p l o t . T h i s r e f l e c t s t h e r e l a t i v e l y s m a l l v a r i a t i o n i n c o n c e n t r a t i o n o f m a c r o n u t r i e n t s r e l a t i v e t o t h e d i f f e r e n c e i n b i o m a s s . A v e r a g e r e l a t i v e a m o u n t s o f m a c r o n u t r i e n t s i n d i f f e r e n t a b o v e -g r o u n d t r e e c o m p o n e n t s a r e l i s t e d i n T a b l e 2 6 . T r e e b o l e s c o n t a i n f r o m 74 t o 85 p e r c e n t o f t h e a b o v e - g r o u n d t r e e b i o m a s s , b u t o n l y 4 1 t o 56 p e r c e n t o f t h e t r e e n i t r o g e n c o n t e n t , 47 t o 73 p e r c e n t o f t h e t r e e p h o s p h o r o u s c o n t e n t , 5 1 t o 76 p e r c e n t o f t h e t r e e c a l c i u m c o n t e n t , 4 9 t o 7 4 p e r c e n t o f t h e t r e e m a g n e s i u m c o n t e n t a n d 5 3 t o 74 p e r c e n t o f t h e t r e e p o t a s s i u m c o n t e n t . T h e s i t u a t i o n i n t h e t r e e c r o w n i s r e v e r s e d . T h e t r e e c r o w n s c o n t a i n f r o m 1 5 t o 26 p e r c e n t o f t h e a b o v e - g r o u n d t r e e b i o m a s s , b u t 44 t o 67 T A B L E 2 4 : D i s t r i b u t i o n o f m a c r o n u t r i e n t s b y p a r t i c u l a r t r e e c o m p o n e n t s o n t h e S q u a m i s h s a m p l e p l o t ( i n k g / h a ) . , n ; M o u n t a i n h e m l o c k = 7 4 ; P a c i f i c s i l v e r f i r = 1 8 2 , T r e e T r e e C o m p o n e n t S p . N P C a Mg K Wood M.h. 46 16 6 1 16 6 1 P . s . f , 1 0 7 1 3 1 4 2 26 1 3 3 Sum 1 5 4 29 2 0 3 4 2 1 9 4 B a r k M.h. 4 9 17 87 5 25 P . s . f . 1 0 9 20 2 1 9 1 3 6 0 Sum 1 5 8 37 3 0 6 18 8 5 B i g b r a n c h e s M.h. 13 3 16 2 7 P . s . f . 3 9 5 87 ' 7 25 Sum 5 2 8 1 0 2 9 3 2 S m a l l b r a n c h e s M.h. 3 1 3 1 2 P . s . f . 1 8 3 25 2 1 0 Sum 2 1 4 2 8 3 1 2 T w i g s M.h. 1 3 2 6 2 6 P . s . f . - - - - -Sum - - — — — F o l i a g e M.h. 2 8 4 10 3 1 0 P . s . f , - - - - -Sum — - - — — T w i g s & F o l i a g e M.h. 4 0 6 16 4 1 5 P . s . f . 1 3 2 1 8 86 1 2 .61 Sum 1 7 2 24 1 0 2 16 76 T r e e B o l e M.h. 95 3 3 1 4 8 2 1 86 ( t o 1 - i n c h o f P . s . f . 2 1 7 3 3 3 6 1 39 1 9 2 d i a m e t e r ) Sum 3 1 2 66 5 0 9 6 0 2 7 9 T r e e C r o w n M.h. 56 9 35 7 24 P . s . f . 1 8 9 2 6 1 9 7 2 0 96 Sum 2 4 6 35 2 3 2 27 1 2 0 T o t a l M.h. 1 5 1 42 1 8 3 27 1 1 1 P . s . f . 4 0 6 59 5 5 8 6 0 2 8 8 Sum 5 5 7 1 0 1 7 4 1 87 3 9 9 68 T A B L E 2 5 : D i s t r i b u t i o n o f m a c r o n u t r i e n t s b y p a r t i c u l a r t r e e c o m p o n e n t s o n t h e H a n e y s a m p l e p l o t ( i n k g / h a ) , n ; W e s t e r n h e m l o c k = 6 7 ; Y e l l o w c e d a r = 1 5 . T r e e T r e e C o m p o n e n t S p . N P C a Mg K Wood W.h. 1 2 4 14 3 2 1 Y . c , 4 1 8 1 3 Sum 16 5 2 2 4 2 4 B a r k W.h. 1 0 2 2 3 1 6 Y . c . 2 0.4 14 0.3 2 Sum 1 2 2 37 1 8 B i g b r a n c h e s W.h. 5 1 8 1 3 Y . c . 1 0.3 4 0.3 0.5 Sum 6 1 1 2 1 4 S m a l l b r a n c h e s W.h. 7 1 9 1 5 Y . c . 1 0.1 3 0.1 0.6 Sum 8 1 1 2 1 5 T w i g s W.h. 6 1 4 1 3 Y . c . - - - ' — _ Sum - - - - -F o l i a g e W.h. 15 2 9 2 7 Y . c . - - - — — Sum - - - - -T w i g s & F o l i a g e W.h. 2 1 3 1 3 3 9 Y . c . 6 1 1 1 0.6 3 Sum 27 4 24 3 12 T r e e B o l e W.h. 2 3 6 37 4 27 ( t o 1 - i n c h o f Y . c . 6 1 2 2 1 5 d i a m e t e r ) Sum 29 7 5 9 5 32 T r e e C r o w n W.h. 34 6 3 0 5 17 Y . c . 7 1 19 1 4 Sum. 4 1 7 49 6 2 1 T o t a l W.h. 57 1 1 66 9 44 Y . c . 14 2 4 1 2 9 Sum 70 13 1 0 7 1 1 5 3 69 T A B L E 2 6 : A v e r a g e p r o p o r t i o n s o f f i v e m a c r o n u t r i e n t s i n d i f f e r e n t t r e e c o m p o n e n t s o n t h e S q u a m i s h a n d H a n e y s a m p l e p l o t s ( i n p e r c e n t ) . n ; M o u n t a i n h e m l o c k = 5; P a c i f i c s i l v e r f i r = 7; w e s t e r n hem-l o c k = 8; Y e l l o w c e d a r = 4. T r e e C o m p o n e n t T r e e N P C a Mg K B i o m a s s S p . Wood M.h. 2 9 . 7 3 8 . 2 3 8 , 8 6 0 . 3 5 2 . 7 6 7 . 2 P . s . f . 2 5 . 0 2 0 . 9 2 4 . 9 4 3 . 8 4 5 . 9 6 6 . 3 W.h. 2 2 . 0 3 5 . 4 2 2 . 4 3 6 . 5 4 6 . 5 6 0 . 8 Y . c . 2 8 . 1 2 6 . 0 1 8 . 2 3 6 . 2 3 8 . 2 7 1 . 7 B a r k M.h. 2 6 . 0 3 5 . 1 3 7 . 3 1 3 . 8 2 1 . 1 1 7 . 9 P . s . f . 2 4 . 4 3 1 . 5 3 7 . 6 2 1 . 8 1 9 . 8 1 3 . 9 W.h. 1 8 . 8 1 5 . 7 3 4 . 6 1 2 . 8 1 5 . 3 1 3 . 3 Y . c . 1 3 . 2 2 1 . 3 3 2 . 7 1 3 . 8 1 5 . 2 6.4 B i g b r a n c h e s M.h. 8.2 6.7 9.2 6.9 6.6 6.9 P . s . f . 1 0 . 4 9.8 1 7 . 3 1 0 . 9 9.1 8.1 W.h. 6.9 6.1 1 0 . 0 6.9 6.4 6.6 Y . c . 7.8 9.1 1 2 . 2 9.3 7.1 5.9 S m a l l b r a n c h e s M.h. 2.3 1.8 2.2 1.9 1.7 1.9 P . s . f . 4.8 4.9 4.5 4.0 3.7 3.9 W.h. 1 1 . 3 9.6 1 2 . 5 1 0 . 5 9.0 1 1 . 3 Y . c . 7.6 7.6 8.1 6.9 6.9 6.2 T w i g s M.h. 8.3 4.8 3.6 4.9 5.0 2.5 P . s . f . - - - - - -W.h. 9.8 7.4 5.3 6.7 5.2 3.3 Y . c . — — — - — -F o l i a g e M.h. 2 5 . 5 1 3 . 4 8.9 1 2 . 2 1 2 . 9 3.6 P . s . f . - - - - - -W.h. 3 1 . 2 2 5 . 8 1 5 . 2 2 6 . 6 1 7 . 6 4.7 Y . c . — — — — — -T w i g s & F o l i a g e M.h. 3 3 . 8 1 8 . 2 1 2 . 5 1 7 . 1 1 7 . 9 6.1 P . s . f . 3 5 . 4 3 2 . 9 1 5 . 7 1 9 . 5 2 1 . 9 7.8 W.h, 4 1 . 0 3 3 . 2 2 0 . 5 3 3 . 3 2 2 . 8 8.0 Y . c . 4 3 . 3 3 6 . 0 2 8 . 8 3 3 . 8 3 2 . 6 9.8 T r e e B o l e M.h. 5 5 . 7 7 3 . 3 7 6 . 1 7 4 . 1 7 3 . 8 8 5 . 1 P . s . f . 49.. 4 5 2 . 4 6 2 . 5 6 5 . 6 6 4 . 8 8 0 . 2 W.h. 4 0 . 8 5 1 . 1 5 7 . 0 4 9 . 3 6 1 . 8 7 4 . 1 Y . c . 4 1 . 3 4 7 . 3 5 0 . 9 5 0 . 0 5 3 . 4 7 8 . 1 70 T A B L E 26 ( c o n t i n u e d ) . : T r e e T r e e C o m p o n e n t S p . N P C a Mg K B i o m a s s T r e e C r o w n M.h. 4 4 . 3 2 6 . 7 2 3 . 9 2 5 . 9 2 6 . 2 1 4 . 9 P . s . f . 5 0 . 6 4 7 . 6 3 7 . 5 3 4 . 4 3 5 . 2 1 9 . 3 W.h. 5 9 . 2 4 8 . 9 4 3 . 0 5 0 . 7 3 8 . 2 2 5 . 9 Y . c . 5 8 . 7 5 2 . 7 4 9 . 1 5 0 . 0 4 6 . 6 2 1 . 9 59 p e r c e n t o f t h e t r e e n i t r o g e n c o n t e n t , 27 t o 5 3 p e r c e n t o f t h e t r e e p h o s p h o r o u s c o n t e n t , 24 t o 4 9 p e r c e n t o f t h e t r e e c a l c i u m c o n t e n t , 26 t o 5 1 p e r c e n t o f t h e t r e e m a g n e s i u m c o n t e n t a n d 26 t o 47 p e r c e n t o f t h e t r e e p o t a s s i u m c o n t e n t . T o c o m p a r e w i t h d a t a f o r y o u n g s t a n d s , s e e T a b l e s 7 a n d 9. C o m p a r i n g d i f f e r e n t t r e e s p e c i e s , we f i n d t h a t m o u n t a i n h e m l o c k h a s t h e h i g h e s t p r o p o r t i o n o f c a t i o n s a n d p h o s p h o r o u s w i t h i n t h e b o l e , b u t t h e l o w e s t w i t h i n t h e c r o w n . C o n v e r s e l y , y e l l o w c e d a r h a s t h e l o w e s t p r o p o r -t i o n o f c a t i o n s a n d p h o s p h o r o u s w i t h i n t h e b o l e a n d t h e h i g h e s t w i t h i n t h e c r o w n . T h e d i s t r i b u t i o n o f n i t r o g e n i s m o r e p r o p o r t i o n a l t o t h e d i s t r i b u -t i o n o f b i o m a s s . T h u s , i n s u m m a r y , t h e f o u r s p e c i e s a p p e a r i n t h e f o l l o w i n g o r d e r a c c o r d i n g t o t h e p r o p o r t i o n a l n u t r i e n t a m o u n t i n t r e e t r u n k s : m o u n t a i n h e m l o c k ^ P a c i f i c s i l v e r f i r > w e s t e r n h e m l o c k ^> y e l l o w c e d a r , I f we c o m p a r e a m o u n t o f n u t r i e n t s w i t h i n t r e e c r o w n s , t h e s p e c i e s w o u l d a p p e a r i n t h e r e v e r s e o r d e r . 71 T h e n u t r i e n t c o n t e n t w i t h i n t h e a b o v e - g r o u n d t r e e b i o m a s s o n t h e S q u a m i s h p l o t i s 5 5 7 k g / h a o f n i t r o g e n , 1 0 1 k g / h a o f p h o s p h o r o u s , 7 4 1 k g / h a o f c a l c i u m , 8 7 k g / h a o f m a g n e s i u m a n d 3 9 9 k g / h a o f p o t a s s i u m . T h e c o r r e s -p o n d i n g v a l u e s f o r t h e H a n e y p l o t a r e s i g n i f i c a n t l y l o w e r : 7 0 k g / h a o f n i t r o g e n , 1 3 k g / h a o f p h o s p h o r o u s , 1 0 7 k g / h a o f c a l c i u m , 1 1 k g / h a o f m a g n e s -i u m a n d 5 3 k g / h a o f p o t a s s i u m ( T a b l e s 24 a n d 2 5 ) . T h e s e a m o u n t s a r e c o m -p a r e d w i t h d a t a f o r o t h e r t r e e s p e c i e s f r o m o t h e r g e o g r a p h i c a l a r e a s i n T a b l e 2 7 . B e c a u s e o f t h e v a r i a t i o n i n m a c r o n u t r i e n t c o n c e n t r a t i o n d i s c u s s e d a b o v e t h e d i s t r i b u t i o n o f m a c r o n u t r i e n t s d o e s n o t a c c u r a t e l y r e f l e c t b i o m a s s d i s t r i b u t i o n ( T a b l e 2 6 ) . Wood o f t r e e b o l e s r e p r e s e n t s f r o m 6 1 t o 72 p e r c e n t o f t h e w h o l e t r e e b i o m a s s b u t a c c o u n t s f o r o n l y 22 t o 3 0 p e r c e n t o f t h e n i t r o g e n , 2 1 t o 38 p e r c e n t o f t h e p h o s p h o r o u s , 1 8 t o 39 p e r c e n t o f t h e c a l c i u m , 3 6 t o 6 0 p e r c e n t o f t h e m a g n e s i u m a n d 38 t o 5 3 p e r c e n t o f t h e p o t a s s i u m . B a r k o f t r e e b o l e s r e p r e s e n t s f r o m 6 t o 1 8 p e r c e n t o f t h e w h o l e t r e e b i o m a s s b u t c o n t a i n s 1 3 t o 26 p e r c e n t o f t h e n i t r o g e n , 16 t o 35 p e r c e n t o f t h e p h o s p h o r o u s , 3 3 t o 38 p e r c e n t o f t h e c a l c i u m , 1 3 t o 22 p e r c e n t o f t h e m a g n e s i u m a n d 1 5 t o 2 1 p e r c e n t o f t h e p o t a s s i u m . B i g b r a n c h e s r e p r e s e n t f r o m 6 t o 8 p e r c e n t o f t h e t o t a l t r e e b i o m a s s a n d r e p -r e s e n t a p p r o x i m a t e l y t h e same p r o p o r t i o n o f m a c r o n u t r i e n t s : 7 t o 1 0 p e r c e n t o f t h e n i t r o g e n , 6 t o 1 0 p e r c e n t o f t h e p h o s p h o r o u s , 1 0 t o 17 p e r c e n t o f t h e c a l c i u m , 7 t o 1 1 p e r c e n t o f t h e m a g n e s i u m a n d 6 t o 9 p e r c e n t o f t h e p o t a s s i u m . S m a l l b r a n c h e s r e p r e s e n t f r o m 2 t o 1 1 p e r c e n t o f t h e w h o l e t r e e b i o m a s s a n d a c c o u n t f o r a p p r o x i m a t e l y t h e same a m o u n t o f t h e T A B L E 2 7 : T h e a m o u n t o f m a c r o n u t r i e n t s i m m o b i l i z e d i n a b o v e - g r o u n d t r e e b i o m a s s b y v a r i o u s s p e c i e s i n d i f f e r e n t c o u n t r i e s . i n X A u t h o r , y e a r D e s c r i p t i o n o f s t a n d A g e 1 0 0 0 k g / h a k g / h a B i o m a s s N. P K C a Mg R e n n i e , 1 9 5 5 P i n e s , a v e r a g e o f many s t a n d s E u r o p e . 1 0 0 - - 1 3 6 3 1 2 9 — R e n n i e , 1 9 5 5 O t h e r c o n i f e r s , a v e r a g e o f many s t a n d s , E u r o p e . 1 0 0 - - 3 1 1 7 0 3 0 7 — R e n n i e , 1 9 5 5 H a r d w o o d s , a v e r a g e o f many s t a n d s , E u r o p e , 1 0 0 - - 3 1 1 4 4 5 8 2 — W r i g h t & W i l l , 1 9 5 8 S c o t s p i n e , E n g l a n d , 64 1 1 9 2 0 5 2 3 1 0 0 2 0 2 3 5 W i l l , 1 9 6 4 R a d i a t a p i n e , New Z e a l a n d . 35 3 6 2 2 3 0 32 2 8 3 1 8 8 — C r a i g & M i l l e r , 1 9 6 6 C o r s i c a n p i n e , E n g l a n d . 35 2 0 0 5 8 6 75 2 0 7 2 3 0 9 3 S w i t z e r & N e l s o n , 1 9 7 2 L o b l o l l y p i n e , M i s s i s s i p p i , U.S.A. 20 9 0 1 7 4 1 9 9 8 9 0 2 3 S m i r n o v a , 1 9 5 1 S p r u c e f o r e s t , U.S.S.R. 9 3 - 4 6 8 85 2 7 8 5 5 2 9 0 C o l e , e t a l . , 1 9 6 7 D o u g l a s - f i r ( i n c l u d i n g r o o t s ) , 3 3 3 W a s h i n g t o n , U.S.A. 36 2 0 5 3 2 0 66 2 2 0 — M o r r i s o n , 1 9 7 2 12 j a c k p i n e s t a n d s , O n t a r i o . 65 1 0 6 1 6 1 1 3 8 1 1 1 6 1 8 O v i n g t o n , 1 9 6 8 D o u g l a s - f i r s t a n d , l i t t e r a n d 2 5 7 3 8 8 u n d e r s t o r e y v e g e t a t i o n . 47 2 6 4 6 3 6 6 2 — O v i n g t o n , 1 9 6 8 S c o t s p i n e s t a n d , l i t t e r a n d 2 2 9 u n d e r s t o r e y v e g e t a t i o n . 47 1 7 6 5 5 6 43 3 2 5 — O v i n g t o n , 1 9 6 8 N o r w a y s p r u c e s t a n d , l i t t e r & 5 6 1 9 1 2 7 7 u n d e r s t o r e y v e g e t a t i o n . 47 1 6 5 6 9 1 — O v i n g t o n , 1 9 6 8 Oak s t a n d , l i t t e r a n d u n d e r - 2 9 2 s t o r e y v e g e t a t i o n . 47 1 3 4 4 6 4 4 0 2 5 4 — M a t e r n a , 1 9 6 4 N o r w a y s p r u c e , C z e c h o s l o v a k i a . 1 0 0 — 8 3 0 1 0 9 3 1 7 4 5 5 3 0 T h e S q u a m i s h s a m p l e p l o t . 4 1 7 4 6 7 5 5 7 1 0 1 3 9 9 7 4 1 87 T h e H a n e y s a m p l e p l o t . 2 4 8 5 6 7 0 1 3 5 3 1 0 7 1 1 N3 73 m a c r o n u t r i e n t s : 2 t o 1 1 p e r c e n t o f t h e n i t r o g e n , 2 t p 1 0 p e r c e n t o f t h e m a g n e s i u m a n d 2 t o 9 p e r c e n t o f t h e p o t a s s i u m . T w i g s a n d f o l i a g e w e r e s t u d i e d s e p a r a t e l y f o r m o u n t a i n a n d w e s t e r n h e m l o c k o n l y . T w i g s r e p -r e s e n t 2.5 a n d 3.3 p e r c e n t o f t h e t o t a l t r e e b i o m a s s a n d c o n t a i n 8 a n d 1 0 p e r c e n t o f t h e n i t r o g e n , 5 a n d 7 p e r c e n t o f t h e p h o s p h o r u s , 4 a n d 5 p e r c e n t o f t h e c a l c i u m , 5 a n d 7 p e r c e n t o f t h e m a g n e s i u m , a n d 5 p e r c e n t o f t h e p o t a s s i u m f o r m o u n t a i n a n d w e s t e r n h e m l o c k , r e s p e c t i v e l y . F o l i a g e r e p r e -s e n t 3.7 a n d 4,8 p e r c e n t o f t h e w h o l e t r e e b i o m a s s b u t r e t a i n s a v e r y h i g h p r o p o r t i o n o f t r e e n u t r i e n t s : 25 a n d 3 1 p e r c e n t o f t h e n i t r o g e n , 1 3 a n d 26 p e r c e n t o f t h e p h o s p h o r u s , 9 a n d 15 p e r c e n t o f t h e c a l c i u m , 1 2 a n d 27 p e r c e n t o f t h e m a g n e s i u m , 1 3 a n d 18 p e r c e n t o f t h e p o t a s s i u m f o r m o u n t a i n a n d w e s t e r n h e m l o c k , r e s p e c t i v e l y . I f we c o n s i d e r t w i g s a n d f o l i a g e t o -g e t h e r f o r a l l f o u r s p e c i e s o n t w o s a m p l e p l o t s , i t a p p e a r s t h a t t w i g s a n d f o l i a g e r e p r e s e n t s 6 t o 10 p e r c e n t o f t h e w h o l e t r e e b i o m a s s a n d c o n t a i n 3 4 t o 4 3 p e r c e n t o f t h e n i t r o g e n , 1 8 t o 36 p e r c e n t o f t h e p h o s p h o r u s , 1 2 t o 29 p e r c e n t o f t h e c a l c i u m , 17 t o 39 p e r c e n t o f t h e m a g n e s i u m a n d 1 8 t o 3 3 p e r c e n t o f t h e p o t a s s i u m . R e m o v a l o f M a c r o n u t r i e n t s . f r o m t h e E c o s y s t e m Due t o D i f f e r e n t  L e v e l s o f T r e e U t i l i z a t i o n : T a b l e s 28 a n d 2 9 s u m m a r i z e t h e a m o u n t o f b i o m a s s a n d m a c r o n u t r i e n t s r e m o v e d f r o m t h e e c o s y s t e m d u e t o d i f f e r e n t l e v e l s o f t r e e u t i l i z a t i o n . L e a v i n g s m a l l b r a n c h e s , t w i g s a n d f o l i a g e a t s t u m p w o u l d r e s u l t i n a 7 p e r c e n t d e c r e a s e i n b i o m a s s y i e l d f r o m t h e S q u a m i s h s a m p l e p l o t . I t w o u l d T A B L E 2 8 : R e m o v a l o f b i o m a s s a n d m a c r o n u t r i e n t s a c c o m p a n y i n g d i f f e r e n t l e v e l s o f u t i l i z a t i o n i n t h e S q u a m i s h s a m p l e p l o t . B i o m a s s N P K C a Mg m e t r i c k g / k g / k g / k g / k g / t o n s / h a % h a % h a % h a % h a % h a % W h o l e t r e e l o g g i n g 4 6 7 1 0 0 5 5 7 1 0 0 1 0 1 1 0 0 3 9 9 1 0 0 7 4 1 1 0 0 87 1 0 0 U t i l i z a t i o n o f t r e e b o l e s u p t o 1 - i n c h o f d i a m e t e r a n d b r a n c h e s o v e r 1 - i n c h o f d i a m e t e r . 4 3 3 9 3 3 6 4 65 73 73 3 1 1 78 6 1 2 8 3 69 79 U t i l i z a t i o n o f t r e e b o l e s u p t o 1 - i n c h o f d i a m e t e r . 3 9 7 8 5 3 1 2 5 6 65 65 2 7 9 70 5 0 9 69 6 0 69 U t i l i z a t i o n o f s t e m w o o d o n l y u p t o 1 - i n c h o f d i a m e t e r . 3 1 9 68 1 5 4 28 29 29 1 9 4 49 2 0 3 27 42 48 T A B L E 2 9 : R e m o v a l o f b i o m a s s a n d m a c r o n u t r i e n t s a c c o m p a n y i n g d i f f e r e n t l e v e l s o f u t i l i z a t i o n i n t h e H a n e y s a m p l e p l o t . B i o m a s s N P K C a Mg m e t r i c k g / k g / k g / k g / k g / t o n s / h a % h a % h a % h a % h a % h a % W h o l e t r e e l o g g i n g 5 6 1 0 0 70 1 0 0 1 3 1 0 0 5 3 1 0 0 1 0 7 1 0 0 1 1 1 0 0 U t i l i z a t i o n o f t r e e b o l e s u p t o 1 - i n c h o f d i a m e t e r a n d b r a n c h e s o v e r 1 - i n c h d i a m e t e r . 47 84 3 5 4 9 8 57 36 68 71 66 6 5 5 U t i l i z a t i o n o f t r e e b o l e s u p t o 1 - i n c h o f d i a m e t e r . 4 1 7 3 29 4 1 7 5 0 32 6 0 59 5 5 5 4 5 U t i l i z a t i o n o f s t e m w o o d o n l y up t o 1 - i n c h o f d i a m e t e r . 27 4 8 16 23 5 36 24 45 22 2 1 4 36 76 d e c r e a s e t h e r e m o v a l o f n i t r o g e n b y 35 p e r c e n t a n d c a l c i u m b y 17 p e r c e n t . U t i l i z a t i o n o f t r e e b o l e o n l y u p t o 1 - i n c h o f d i a m e t e r w o u l d d e c r e a s e b i o -m a s s y i e l d b y 1 5 p e r c e n t b u t i t w o u l d d e c r e a s e t h e r e m o v a l o f m a c r o n u t r i -e n t s m o r e s u b s t a n t i a l l y . R e m o v a l o f n i t r o g e n f r o m t h e e c o s y s t e m w o u l d b e r e d u c e d b y 44 p e r c e n t , p h o s p h o r o u s b y 3 5 p e r c e n t a n d c a t i o n s b y 3 0 p e r c e n t . I n c o m p a r i s o n , h a r v e s t i n g o f s t e m w o o d o n l y u p t o 1 - i n c h o f d i a m e t e r w o u l d r e d u c e b i o m a s s y i e l d b y 3 2 p e r c e n t , r e m o v a l o f N, P, C a b y a s m u c h a s 72 p e r c e n t a n d r e m o v a l o f K a n d Mg b y 5 1 p e r c e n t . On t h e H a n e y s a m p l e p l o t , r e d u c t i o n o f b i o m a s s y i e l d w o u l d b e g r e a t e r a c c o m p a n i e d a l s o b y g r e a t e r r e d u c t i o n o f n u t r i e n t l o s s e s . 77 V I SUMMARY AND CONCLUSIONS T h e m e t h o d u s e d f o r t h e d e t e r m i n a t i o n o f t h e n u t r i e n t d e m a n d o n t h e s i t e o f t w o f o r e s t s t a n d s w a s b a s e d o n a m e a s u r e m e n t o f t h e t o t a l n u t r i -e n t i m m o b i l i z a t i o n b y t h e s t a n d ( e x c l u d i n g s t u m p a n d r o o t b i o m a s s ) : t h e s o - c a l l e d s u b s t i t u t i o n p r i n c i p l e (Tamm, 1 9 6 4 ) . A c c o r d i n g t o Tamm, t h i s i s o n l y a f i r s t a p p r o x i m a t i o n f o r d e t e r m i n i n g t r e e n u t r i e n t r e q u i r e m e n t s . N e v e r t h e l e s s , t h i s m e t h o d h a s b e e n u s e d b y a n u m b e r o f d i f f e r e n t a u t h o r s ( e . g . E b e r m a y e r 1 8 7 6 , 1 8 8 2 ; R e n n i e 1 9 5 5 ; O v i n g t o n 1 9 6 2 ; W e e t m a n a n d W e b b e r 1 9 7 2 , e t c . ) t o a p p r o x i m a t e t h e n e t n u t r i e n t u p t a k e o f f o r e s t s t a n d o v e r a r o t a t i o n a n d t h e l o s s o f n u t r i e n t s f r o m a f o r e s t s i t e i n h a r v e s t e d m a t e r i a l . T h e f o l l o w i n g c o n c l u s i o n s w e r e r e a c h e d o n t h e b a s i s o f t h i s s t u d y : 1. V a r i o u s a b o v e - g r o u n d t r e e c o m p o n e n t s o f f o u r s t u d i e d s p e c i e s a p p e a r t o b e i n t h e f o l l o w i n g o r d e r a c c o r d i n g t o t h e i r n u t r i e n t c o n c e n t r a -t i o n s : stemwood<C b i g b r a n c h e s < f s m a l l b r a n c h e s <^ s t e m b a r k <^ t w i g s < ^ f o l i a g e ( h e m l o c k spp„») o r t w i g s a n d f o l i a g e ( P a c i f i c s i l v e r f i r a n d y e l l o w c e d a r ) . , T h i s o r d e r i s v a l i d f o r a l l m a c r o n u t r i e n t s w i t h t h e e x c e p t i o n o f c a l c i u m . T h e c o n c e n t r a t i o n o f c a l c i u m i n s t e m b a r k i s h i g h e r t h a n i n t w i g s a n d a p p r o x -i m a t e l y e q u a l t o f o l i a g e c o n c e n t r a t i o n ( h e m l o c k s p p . ) o r t w i g s a n d f o l i a g e 78 c o n c e n t r a t i o n ( f i r a n d c e d a r ) . C o n e s , w h e r e s t u d i e d , e x h i b i t e d t h e h i g h e s t c o n c e n t r a t i o n o f N, P, K; t h e l o w e s t o f C a . T h e i r Mg c o n c e n t r a t i o n w a s a p p r o x i m a t e l y e q u a l t o t h a t o f t w i g s a n d f o l i a g e ( f i r ) o r t w i g s ( h e m l o c k ) . 2. T h e f o u r t r e e s p e c i e s i n v e s t i g a t e d a p p e a r t o b e i n t h e f o l -l o w i n g o r d e r a c c o r d i n g t o m a c r o n u t r i e n t c o n c e n t r a t i o n : W e s t e r n h e m l o c k ^ M o u n t a i n h e m l o c k <^ P a c i f i c s i l v e r f i r <^ Y e l l o w c e d a r . Y e l l o w c e d a r e x h i b i t s c o n s i d e r a b l y h i g h e r c a l c i u m c o n c e n t r a t i o n i n a l l t r e e c o m p o n e n t s t h a t d o t h e o t h e r s p e c i e s . P a c i f i c s i l v e r f i r e x h i b i t s h i g h e r n u t r i e n t c o n c e n t r a t i o n s f o r a l m o s t a l l e l e m e n t s a n d t r e e c o m p o n e n t s t h a n d o b o t h h e m l o c k s p e c i e s . N u t r i e n t c o n c e n t r a t i o n s i n m o u n t a i n h e m l o c k a r e a p -p r o x i m a t e l y e q u a l t o c o n c e n t r a t i o n s i n w e s t e r n h e m l o c k f o r a l l t r e e c o m p o n -e n t s . S i t e c o n d i t i o n s o f t h e t w o s a m p l e p l o t s w e r e c o n s i d e r a b l y d i f f e r e n t a n d t r e e s p e c i e s might a p p e a r i n a d i f f e r e n t o r d e r i f t h e y w e r e g r o w n u n d e r u n i f o r m s i t e c o n d i t i o n s . 3. T h e w e i g h t d i s t r i b u t i o n o f n u t r i e n t s w i t h i n t h e a b o v e - g r o u n d t r e e b i o m a s s i s n o t p r o p o r t i o n a l t o t h e b i o m a s s d i s t r i b u t i o n . T r e e c r o w n s ( b r a n c h e s , t w i g s a n d f o l i a g e ) c o n t a i n a s m u c h n u t r i e n t s a s t r e e b o l e s . W i t h i n t h e c r o w n , t h e l a r g e s t a m o u n t o f n u t r i e n t s i s b u i l t u p w i t h i n t h e f o l i a g e a n d t w i g s . T h e p r o p o r t i o n o f n u t r i e n t s i n t h e c r o w n c o m p o n e n t s v a r i e s f o r v a r i o u s e l e m e n t s a s w e l l a s f o r d i f f e r e n t t r e e s p e c i e s . 79 4. B a r k c o n t a i n s a p p r o x i m a t e l y h a l f o f t h e n u t r i e n t s o f t h e w h o l e t r e e b o l e . T h e p r o p o r t i o n s o f e l e m e n t s w i t h i n b a r k a n d w o o d v a r y f o r d i f f e r e n t e l e m e n t s a n d t r e e s p e c i e s , 5. T h e u p t a k e o f n u t r i e n t s i m m o b i l i z e d i n t h e a b o v e - g r o u n d t r e e b i o m a s s o v e r a t i m e p e r i o d o f 4 2 0 y e a r s i n t h e S q u a m i s h s a m p l e p l o t l o c a t e d i n t h e S u b a l p i n e M o u n t a i n H e m l o c k Z o n e - p l a n t a s s o c i a t i o n A b i e t o ( a m a b i l i s ) - T s u g e t u m m e r t e n s i a n a e v a r i a n t a b i e t o - t s u g o s u m m e r t e n s i a n a e ( B r o o k e , 1 9 7 0 ) w a s c a l c u l a t e d t o b e 5 5 7 k g o f n i t r o g e n , 1 0 1 k g o f p h o s p h o r u s , 3 9 9 k g o f p o t a s s i u m , 7 4 1 k g o f c a l c i u m a n d 87 k g o f m a g n e s i u m p e r h e c t a r e f o r o v e n d r y b i o m a s s o f 4 6 7 m e t r i c t o n s . 6. T h e u p t a k e o f n u t r i e n t s i m m o b i l i z e d i n t h e a b o v e - g r o u n d t r e e b i o m a s s o v e r a t i m e p e r i o d o f 2 5 0 y e a r s i n t h e H a n e y s a m p l e p l o t l o c a t e d i n t h e W e t S u b z o n e o f t h e C o a s t a l W e s t e r n H e m l o c k Z o n e - p l a n t a s s o c i a t i o n A b i e t o - T s u g e t u m h e t e r o p h y l l a e ( O r l o c i , 1 9 6 5 ) w a s c a l c u l a t e d t o b e 70 k g o f n i t r o g e n , 1 3 k g o f p h o s p h o r u s , 5 3 k g o f p o t a s s i u m , 1 0 7 k g o f c a l c i u m a n d 1 1 k g o f m a g n e s i u m p e r h e c t a r e f o r o v e n d r y b i o m a s s o f 56 m e t r i c t o n s . 7. I n c o m p a r i s o n w i t h w h o l e s - t r e e l o g g i n g , t h e a m o u n t o f n u t r i e n t s r e m o v e d f r o m t h e s t u d y s i t e s c a n b e r e d u c e d b y 5 0 - 7 0 p e r c e n t i f o n l y s t e m -w o o d w e r e h a r v e s t e d a n d o t h e r t r e e c o m p o n e n t s w e r e l e f t o n t h e s i t e . 8 0 R E F E R E N C E S A l l a n , J . E. 1 9 6 9 . T h e p r e p a r a t i o n o f a g r i c u l t u r a l s a m p l e s f o r a n a l y s i s b y a t o m i c a b s o r p t i o n s p e c t r o s c o p y , V a r i a n A e r o g r a p h , C a l i f o r n i a , U.S.A. B e a t o n , J . D., e t a l . 1 9 6 5 . O b s e r v a t i o n s o n f o l i a g e n u t r i e n t c o n t e n t o f s e v e r a l c o n i f e r o u s t r e e s p e c i e s i n B r i t i s h C o l u m b i a . F o r . C h r o n . 4 1 ( 2 ) : 2 2 2 - 3 6 . B r o o k e , R. C. 1 9 6 6 . V e g e t a t i o n - e n v i r o n m e n t r e l a t i o n s h i p s o f s u b a l p i n e m o u n t a i n h e m l o c k z o n e e c o s y s t e m s , P h . D . T h e s i s , D e p a r t m e n t o f B o t a n y , U n i v e r s i t y o f B r i t i s h C o l u m b i a , V a n c o u v e r , 2 2 5 p . , A p p . 1 1 0 p . B r o o k e , R. C , E . B. P e t e r s o n , V. J . K r a j i n a . 1 9 7 0 . T h e s u b a l p i n e m o u n t a i n h e m l o c k z o n e . E c o l . o f W e s t . N. A m e r . 2 ( 2 ) : 1 4 8 - 3 4 9 . C o l e , D. W., G e s s e l , S. P., D i c e , S. F. 1 9 6 7 . D i s t r i b u t i o n a n d c y c l i n g o f N, P, K, C a i n a s e c o n d - g r o x ^ t h D o u g l a s - f i r e c o s y s t e m . S y m p o s i u m o n p r i m a r y p r o d u c t i v i t y a n d m i n e r a l c y c l i n g i n n a t u r a l e c o s y s t e m s . A m e r i c a n A s s o c i a t i o n f o r t h e A d v a n c e s o f S c i e n c e 1 3 t h A n n u a l M e e t i n g , New Y o r k C i t y , D e c e m b e r 2 7 , 1 9 6 7 . P u b l i s h e d b y t h e U n i v e r s i t y o f M a i n e P r e s s . C u r r y , R. R. 1 9 7 3 . G e o l o g i c a n d h y d r o l o g i c e f f e c t s o f e v e n - a g e m a n a g e m e n t o n p r o d u c t i v i t y o f f o r e s t s o i l s , p a r t i c u l a r l y i n t h e D o u g l a s - f i r r e g i o n . E v e n - A g e M a n a g e m e n t S y m p o s i u m , A u g u s t 1 9 7 2 . E d . R. K. H e r m a n n , D. P. L a v e n d e r . O r e g o n S t a t e U n i v e r s i t y . D a v i d , J . D. 1 9 6 0 . T h e d e t e r m i n a t i o n o f e x c h a n g e a b l e N a , K, C a , Mg i n s o i l s b y a t o m i c - a b s o r p t i o n s p e c t r o p h o t o m e t r y . A n a l y s t 8 5 : 4 9 5 - 5 0 3 . D u k e U n i v e r s i t y S c h o o l o f F o r e s t r y B u l l e t i n 1 5 , 1 9 5 9 . M i n e r a l n u t r i t i o n o f t r e e s . A S y m p o s i u m . D u r h a m , N o r t h C a r o l i n a , M a y 1 9 5 9 . D u v i g n e a u d , P., D e n a e y e r - D e S m e t , S. 1 9 6 4 . L e c y c l e d e s e l e m e n t s b i o g e n e s d a n s l ' e c o s y s t e m e f o r e t . L ' U n i v e r s i t e L i b r e d e B r u x e l l e s , P u b l . 5 8 . 81 D u v i g n e a u d , P., D e n a e y e r - D e S m e t , S. 1 9 6 7 , B i o m a s s , P r o d u c t i v i t y a n d M i n e r a l C y c l i n g i n D e c i d u o u s F o r e s t s i n B e l g i u m . S y m p o s i u m o n P r i m a r y P r o d u c t i v i t y a n d M i n e r a l C y c l i n g i n N a t u r a l E c o s y s t e m s . A m e r i c a n A s s o c i a t i o n f o r t h e A d v a n c e m e n t o f S c i e n c e , 1 3 t h A n n u a l M e e t i n g , New Y o r k C i t y , D e c e m b e r 2 7 , 1 9 6 7 . P u b l i s h e d b y t h e U n i v e r s i t y o f M a i n e P r e s s . D y e r , R. F . 1 9 6 7 . 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I n g e s t a d , T. 1 9 7 0 . A d e f i n i t i o n o f o p t i m u m n u t r i e n t r e q u i r e m e n t s i n b i r c h s e e d l i n g s , P h y s i o l . P l a n t . 2 3 ( 6 ) : 1 1 2 7 - 3 8 , K a u l , 0. N., S r i v o s h a n a , P. B. L . 1 9 6 4 , M i n e r a l c o n t e n t o f f i r ( A b i e s  p i n d r o w ) . a n d K a i l ( P i n u s w a l l i c h i a n a ) . J o u r n a l o f t h e S o c i e t y o f I n d i a n F o r e s t e r s 4 ( 2 1 3 ) : 1 1 4 - 2 0 . K e a y , J . , A. G, T u r t o n . 1 9 6 9 . D i s t r i b u t i o n o f b i o m a s s a n d m a j o r n u t r i e n t s i n a m a r i t i m e p i n e p l a n t a t i o n , A u s t r a l i a n F o r e s t r y 3 5 : 3 9 - 4 8 , K e l l e y , J . D., S h i e r , R. W. 1 9 6 5 , S e a s o n a l c h a n g e s i n t h e m a c r o n u t r i e n t c o m p o s i t i o n o f l e a v e s a n d s t e m s o f T a x u s m e d i a . P r o c . A m e r . S o c . H o r t . S c i . 8 6 : 8 0 9 - 1 4 . K r a j i n a , V. 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M i n e r a l c o n t e n t o f p l a n t a t i o n s of- P i n u s s i l v e s t r i s L . A n n a l s o f B o t a n y , N , S. 2 3 ( 8 9 ) : 75-88 . O v i n g t o n , J , D. 1 9 5 9 , T h e c i r c u l a t i o n o f m i n e r a l s i n p l a n t a t i o n s o f P i n u s ' - ' s i l v e s t r i s L . A n n a l s o f B o t a n y , N, S. 2 3 ( 9 0 ) : 2 2 9 - 2 3 9 . O v i n g t o n , J . D. 1 9 6 0 . T h e n u t r i e n t c y c l e a n d i t s m o d i f i c a t i o n t h r o u g h s i l v i c u l t u r a l p r a c t i c e . 5 t h W o r l d F o r e s t r y C o n g r e s s , S e a t t l e , W a s h i n g t o n , O v i n g t o n , J . D, 1 9 6 2 , Q u a n t i t a t i v e e c o l o g y a n d t h e w o o d l a n d e c o s y s t e m c o n c e p t . A d v a n c e s i n E c o l o g i c a l R e s e a r c h , V o l . 1, A c a d e m i c P r e s s , L o n d o n , 1 9 6 2 , p p . 1 0 3 - 9 2 . O v i n g t o n , J . D., M a d g w i c k , A. A. I . 1 9 5 9 . 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T h e n u t r i e n t s t a t u s o f t w o h i g h e l e v a t i o n f o r e s t s o i l s a n d i m p l i c a t i o n s f o r some f o r e s t m a n a g e m e n t p r a c t i c e s . B . S . F . T h e s i s , F a c u l t y o f F o r e s t r y , U n i v e r s i t y o f B r i t i s h C o l u m b i a , V a n c o u v e r , B . C. R e n n i e , P. J . 1 9 5 5 . T h e u p t a k e o f n u t r i e n t s b y m a t u r e f o r e s t g r o w t h . P l a n t a n d S o i l 7 ( 1 ) : 4 9 - 9 1 . R e n n i e , P. J . 1 9 6 6 . A f o r e s t s a m p l i n g p r o c e d u r e f o r n u t r i e n t u p t a k e s t u d i e s . C o m m o n w e a l t h F r , R e v , 4 5 ( 2 ) : 1 1 9 - 2 8 . R o d i n , L . E . , B a z i l e v i c h , N. I . 1967. P r o d u c t i o n a n d m i n e r a l c y c l i n g i n t e r r e s t r i a l v e g e t a t i o n . O l i v e r a n d B o y d , E d i n b u r g , 1 9 6 7 . ( 2 8 8 p p . ) . 85 R o m e l l , L . G, 1 9 5 7 , Man-made " n a t u r e 1 ' o f n o r t h e r n l a n d s , I n " P r o c e e d i n g s a n d P a p e r s o f t h e S i x t h T e c h n i c a l M e e t i n g , I n t e r n a t i o n a l U n i o n o f C o n s e r v a t i o n o f N a t u r e a n d N a t u r a l R e s o u r c e s , E d i n b u r g , 1 9 5 6 " , L o n d o n , p p . 5 1 - 5 3 , R o w e , J , S, 1 9 5 9 . F o r e s t r e g i o n s o f C a n a d a . C a n a d a D e p t . N o r t h e r n A f f a i r s a n d N a t u r a l R e s o u r c e s , F o r e s t r y B r a n c h , B u l l e t i n 1 2 3 , Q u e e n ' s P r i n t e r , O t t a w a , 7 1 p p , S m i t h , W. H., L . E . N e l s o n , G. L , S w i t z e r , 1 9 6 3 . T h e c h a r a c t e r i z a t i o n o f d r y m a t t e r a n d n i t r o g e n a c c u m u l a t i o n b y l o b l o l l y p i n e ( P i n u s -- t a e d a L . ) o n p o o r s i t e s . S o i l S c , S o c . o f Am. P r o c , 2 7 : 4 6 5 - 4 6 8 . S m i t h , J . H. G. 1 9 7 4 . 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I n t e r n a t i o n a l R e v i e w o f F o r e s t r y R e s e a r c h A c a d e m i c P r e s s (New Y o r k ) 1 : 1 1 5 - 1 7 0 . Tamm, C. 0. 1 9 6 9 , S i t e d a m a g e s b y t h i n n i n g d u e t o r e m o v a l o f o r g a n i c m a t t e r a n d p l a n t n u t r i e n t s , P r o c , I U F R 0 M e e t . , S t o c k h o l m , S e p t e m b e r 1 9 6 9 . T e c h n i c o n A u t o - A n a l y s e r M e t h o d o l o g y , T e c h n i c o n C o r p o r a t i o n , T a r r y t o w n , New Y o r k , T s u t s u m i , T,, K a w a h a r a , T,, S h i d e i , T. 1 9 6 8 . T h e c i r c u l a t i o n o f n u t r i e n t s i n f o r e s t e c o s y s t e m I , On t h e a m o u n t o f n u t r i e n t s c o n t a i n e d i n t h e a b o v e - g r o u n d p a r t s o f s i n g l e t r e e a n d o f s t a n d . J . J a p . F o r . S o c . 5 0 ( 3 ) : 1 6 6 - 7 4 . 86 T u r t o n , A. G,, K e a y , J . 1 9 7 0 . 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V a r i a t i o n i n n u t r i e n t u p t a k e o f t r e e s . F o r e s t f e r t i l i z a -t i o n , t h e o r y a n d p r a c t i c e . P a p e r s p r e s e n t e d a t t h e s y m p o s i u m o n f o r e s t f e r t i l i z a t i o n , A p r i l 1 9 6 7 , a t G a i n e s v i l l e , F l o r i d a . T e n n e s s e e V a l l e y A u t h o r i t y N a t i o n a l F e r t i l i z e r D e v e l o p m e n t C e n t r e , M u s c l e S h o a l s , A l a . , 1 9 6 8 . 3 0 6 p p . W e e t m a n , G. F. 1 9 7 1 . E f f e c t o f t h i n n i n g a n d f e r t i l i z a t i o n o n t h e n u t r i e n t u p t a k e , g r o w t h a n d w o o d q u a l i t y o f u p l a n d b l a c k s p r u c e . W o o d l . P o p . P u l p P o p . R e s . I n s t . C o n . 2 8 : 1 8 . W e e t m a n , G, F . , W e b b e r , B. 1 9 7 2 . T h e i n f l u e n c e o f w o o d h a r v e s t i n g o n t h e n u t r i e n t s t a t u s o f t w o s p r u c e s t a n d s . C a n . J . 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M, 1 9 5 9 , N u t r i e n t r e t u r n i n l i t t e r a n d r a i n f a l l u n d e r some e x o t i c c o n i f e r s t a n d s i n New Z e a l a n d , New Z e a l a n d F o r e s t R e s . N o t e 1 8 , R e p t . New Z e a l a n d J o u r , o f A g , 2 ( 4 ) : 7 1 9 - 7 3 4 . 87 W i l l , G, M, 1 9 6 4 , D r y m a t t e r p r o d u c t i o n a n d n u t r i e n t u p t a k e b y P i n u s  r a d i a t a i n New Z e l a n d , Commonw, F o r , R e v , 4 3 (l).:57<-70. W i l l , G, M, 1 9 6 8 . T h e u p t a k e , c y c l i n g a n d r e m o v a l o f m i n e r a l n u t r i e n t s b y c r o p s o f P i n u s r a d i a t a , P r o c . N. A. E c o l , S o c . 1 5 : 2 0 - 2 4 , W i l l i a m s , C. H., J . R, T w i n e . 1 9 6 7 . D e t e r m i n a t i o n o f N, S, P, K, N a , C a a n d Mg i n p l a n t m a t e r i a l b y a u t o m a t i c a n a l y s i s . D i v i s i o n o f P l a n t I n d u s t r y T e c h n i c a l P a p e r N o . 2 4 . C o m m o n w e a l t h S c i e n t i f i c a n d I n d u s t r i a l R e s e a r c h O r g a n i z a t i o n , A u s t r a l i a , M e l b o u r n e , 1 9 6 7 , W r i g h t , T. W., G. M. W i l l , 1 9 5 8 * T h e n u t r i e n t c o n t e n t o f S c o t s a n d C o r s i c a n p i n e g r o w i n g o n s a n d d u n e s . F o r e s t r y 3 1 : 1 3 - 2 5 . Y o u n g , H. E . , L . S t r a n d , R. A l t e n b e r g e r . 1 9 6 4 . P r e l i m i n a r y f r e s h a n d d r y w e i g h t t a b l e s f o r s e v e n t r e e s p e c i e s i n M a i n e . M a i n e A g r i c u l t u r a l E x p e r i m e n t a l S t a t i o n T e c h . B u l l . 1 2 , U n i v e r s i t y o f M a i n e , N o v e m b e r 1 9 6 4 . Y o u n g , H. E . , P. N. C a r p e n t e r , R. A, A l t e n b e r g e r . 1 9 6 4 . P r e l i m i n a r y t a b l e s o f some c h e m i c a l e l e m e n t s i n s e v e n s p e c i e s i n M a i n e , M a i n e A g r i c u l t u r a l E x p e r i m e n t a l S t a t i o n T e c h . B u l l e t i n 2 0 , U n i v e r s i t y o f M a i n e , O c t o b e r 1 9 6 5 . Y o u n g , H, E . , G u i n n , V. P, 1 9 6 6 . C h e m i c a l e l e m e n t s i n c o m p l e t e m a t u r e t r e e s o f s e v e n s p e c i e s i n M a i n e . T a p p i 4 9 ( 5 ) . . Y a n g , H. E , , D y e r , R. F , , D u b e , G. F , 1 9 6 7 . N u t r i e n t d i s t r i b u t i o n i n t h e c r o w n o f p o l e s i z e r e d s p r u c e a n d w h i t e p i n e , M a i n e F a r m R e s e a r c h , O r o n o 1 9 6 7 , ( A p r i l ) , p . 5 . A P P E N D I X 1 S C I E N T I F I C AND COMMON NAMES OF T R E E S P E C I E S A b i e s - a m a b i l i s ( D o u g l . ) F o r b e s P a c i f i c s i l v e r f i r A b i e s l a s i o c a r p a ( H o o k . ) N u t t , A l p i n e f i r C h a m a e c y p a r i s ribotkatensis ( D . Don). S p a c h Y e l l o w c e d a r L a r i x l e p t o l e p i s ( S i e b . e t Z u c c ) . G o r d . J a p a n e s e l a r c h P i c e a a b i e s ( L . ) K a r s t N o r w a y s p r u c e P i c e a e n g e l m a n i i i i P a r r y E n g e l m a n s p r u c e P i c e a s i t c h e n s i s ( B o n g . ) . C a r r . S i t k a s p r u c e P i n u s b a n k s i a n a L a m b . J a c k p i n e P i n u s c o n t o r t a D o u g l * . L o d g e p o l e p i n e P i n u s r a d i a t a D. D o n R a d i a t a p i n e P i n u s s i l v e s t r i s L , S c o t s p i n e P i n u s t a e d a L . L o b l o l l y p i n e P s e u d o t s u g a m e n z i e s i i ( M i r b e l ) F r a n c o D o u g l a s - f i r T h u j a p l i c a t a D. D o n W e s t e r n r e d c e d a r T s u g a h e t e r o p h y l l a ( R a f . ) S a r g . W e s t e r n h e m l o c k T s u g a m e r t e n s i a n a ( B o n g . ) C a r r . M o u n t a i n h e m l o c k A P P E N D I X 2 B A S I C S T A T I S T I C S FOR THE INDEPENDENT V A R I A B L E S 91 P a c i f i c s i l v e r f i r = 7 ; w e s t e r n h e m l o c k =8. I n d e p e n d e n t V a r i a b l e T r e e S p e c . U n i t M e a n V a l u e S t a n d a r d D e v i a t i o n M i n i m u m V a l u e M a x i m u m V a l u e D 2 . H M.h. P . s . f . W.h. 2 nr-.rrt 12.59 13.83 2,55 6.64 11.81 2.11 4.79 1.71 0.44 19.86 33.67 6.65 BA.H M.h. P . s . f , W.h. 2 irr-.m 9.89 10.86 2.00 5.21 9.28 1.65 3.76 1.34 0.34 15.60 26.44 5.22 D.CL M.h. P . s . f . W.h. m.m 14.55 14.25 5.13 4.01 8.21 3.15 9.00 3.56 1.88 19.20 25.67 11.37 D . C L 2 M.h. P . s . f . W.h. m.m 2 353.51 361.81 89.57 116.62 262.08 80.37 185.44 41.00 19.41 483.90 729.13 263.74 D.CW 2 M.h. P . s . f . W.h. 2 m.m 30.98 33.09 9.76 24.54 20.70 8.51 9.65 9.38 1.54 67.33 63.79 26.11 D 2.H.CW M.h. P . s . f . W.h. 2 m .m.m 93.11 107.86 15.12 71.89 101.02 15.93 22.54 9.41 1.51 186.66 282.82 48.55 (BA.H)+CCW.CL) M.h. P . s . f . W.h. j m .rn+m.m 169.42 174.85 82.15 62.96 74.37 46.88 100.58 64.59 29.90 252.48 265.00 169.55 BA.H.CL M.h. P . s . f . W.h. 2 m^.m.m 242.09 285.52 35.93 135.21 273.24 38.97 77.55 15.45 4.20 393.03 751.00 121.18 BA M.h. P . s . f . W.h. ™2 m 0.296 0.304 0.087 0.127 0.210 0.058 0.150 0.075 0.020 0.456 0.642 0.189 cw2' M.h. P . s . f . W.h. m.m 46.68 52.10 27.10 28.56 18.44 15.98 22.59 30.25 8.41 88.36 72.25 53.29 CR.VOL M.h. P . s . f . W.h. m 3 299.58 321.99 123.47 194.54 172.49 102.84 119.13 91.07 22,68 582.94 546.64 323.67 CR.SURF M.h. P . s . f . W.h. m 2 253.39 260.98 130.67 93.46 104.28 74.59 153.07 102,15 47.38 378.51 390.01 269.30 A P P E N D I X 3 S I G N I F I C A N T INDEPENDENT V A R I A B L E S I N S I M P L E L I N E A R AND L O G A R I T H M I C R E G R E S S I O N EQUATIONS FOR BIOMASS P R E D I C T I O N OF P A R T I C U L A R T R E E COMPONENTS S i g n i f i c a n t i n d e p e n d e n t v a r i a b l e s i n s i m p l e l i n e a r r e g r e s s i o n e q u a t i o n s f o b i o m a s s p r e d i c t i o n o f p a r t i c u l a r c o m p o n e n t s ( v a r i a b l e s a p p e a r i n o r d e r o f d e c r e a s i n g r 2 v a l u e s ) , S q u a m i s h s a m p l e p l o t , n ; M o u n t a i n h e m l o c k = 5; P a c i f i c s i l v e r f i r = 7, T r e e S E E C o m p o n e n t T r e e I n d e p e n d e n t R e g r e s s , i n ( v a r i a b l e Y ) S p . V a r i a b l e X. I n t e r c e p t C o e f f , r 2 k g w h o l e t r e e M.h. D 2 . H 3 6 6 . 4 8 4 1 3 1 . 6 6 7 9 9 . 7 4 5 2 b i o m a s s D - 1 9 0 4 . 5 5 0 6 5 2 6 , 1 3 0 9 8 . 0 1 1 4 3 P . s . f . D Z . H 1 7 6 . 5 6 7 1 2 8 . 1 3 9 9 9 . 2 6 1 4 3 D - 2 0 2 9 . 0 5 0 6 7 7 5 . 6 4 0 9 7 . 5 3 2 6 1 w o o d M.h. D 2 . H 2 6 6 . 1 6 0 8 6 . 2 9 2 9 9 . 0 5 65 b i o m a s s D - 1 1 9 4 . 9 5 0 4 2 3 1 . 7 8 0 9 5 . 2 9 1 4 4 B A ' 4 2 . 5 3 8 4 4 3 0 . 2 2 0 9 5 . 1 2 1 4 7 P . s . f . D 2 . H 8 0 . 8 2 4 9 0 . 4 7 8 9 9 . 5 8 76 B A - 2 1 3 . 4 9 7 5 0 8 9 . 5 2 0 9 9 . 4 6 86 D - 1 4 6 7 . 7 2 0 4 7 6 9 . 2 1 0 9 7 . 2 3 1 9 5 b a r k M.h. B A - 1 5 . 1 9 3 1 2 8 2 . 7 4 0 9 8 . 7 0 2 2 b i o m a s s D - 3 6 9 . 5 1 9 1 2 1 8 . 6 7 0 9 7 . 8 0 28 D 2 . H 5 8 . 3 5 9 2 4 . 2 8 6 9 7 . 1 0 32 P . s . f . D 2 . H 1 7 . 7 5 3 1 8 . 7 6 3 9 6 . 1 2 4 9 B A - 4 2 . 3 2 4 1 0 5 2 . 2 8 0 9 5 . 4 3 5 3 b i o m a s s o f M.h. D 2.11.CW 2 2 . 0 4 8 1 . 3 7 8 9 9 . 2 3 1 0 b i g b r a n c h e s D.CW 2 2 5 . 6 3 2 4 . 0 2 7 9 8 . 7 0 13 B A - 7 9 . 4 0 2 7 7 6 . 9 9 3 9 8 . 0 0 16 CR.VOL - 0 . 5 8 6 0 . 5 0 4 9 7 . 1 4 19 D - 2 9 0 . 5 1 6 7 3 2 . 3 5 3 9 5 . 5 8 2 4 D 2 . H - 3 2 . 1 7 6 1 4 . 4 9 8 9 3 . 6 5 29 P . s . f . B A - 3 9 . 7 7 0 6 6 3 . 7 7 8 8 1 . 9 3 72 D - 2 0 2 . 4 1 3 6 2 0 . 4 1 1 7 9 . 6 9 76 D 2 . l i 1 . 0 4 1 1 1 . 6 2 5 7 9 . 6 1 76 B A . H . C L 2 2 . 0 2 5 0 . 4 9 0 7 5 . 6 3 83 b i o m a s s o f M.h. n o o n e i n d e p , v a r i a b l e i s s i g n i f i c a n t , s m a l l b r a n c h e s P . s . f . L - 2 8 . 9 4 4 2 . 6 6 0 8 8 . 0 5 9 D - 7 . 5 5 8 1 0 3 . 6 7 5 8 5 . 9 8 1 0 D.CL 1 3 . 6 3 2 2 . 7 8 4 3 5 . 3 3 1 0 ( B A . H ) . + (CL.CW) 0 . 9 5 0 0 . 2 9 9 8 0 . 9 3 1 2 C L - 2 4 . 6 0 9 3.472 8 0 . 4 9 12 CR.SURF - 1 . 8 3 3 0 . 2 1 1 7 9 . 2 4 12 94 T r e e S E E C o m p o n e n t T r e e I n d e p e n d e n t R e g r e s s , i n ( v a r i a b l e Y ) S p . V a r i a b l e X. I n t e r c e p t C o e f f . r 2 k g b i o m a s s o f M.h, D.CL - 2 0 . 6 3 7 9.679 9 7 . 0 3 8 t w i g s a n d D - 5 1 . 4 4 8 2 8 5 . 1 9 5 9 2 . 1 2 1 3 f o l i a g e B A.H.CL * 5 3 . 0 7 0 0 . 2 7 7 9 0 . 4 2 14 BA 3 3 . 8 9 7 2 9 1 . 9 8 8 8 7 . 9 5 16 D 2.H 5 0 . 1 8 5 5 . 5 6 4 8 7 . 1 6 16 (BA.H). + (CL.CW) 2 2 . 9 9 5 0 . 5 7 4 8 3 . 9 2 18 P . s . f . D.CL - 2 . 9 9 5 8 . 6 9 6 9 4 . 6 5 19 D - 6 4 . 8 4 9 3 1 6 . 4 1 0 9 1 . 0 9 24 3 A 2 1 . 9 4 7 3 2 5 . 8 5 9 8 6 . 7 8 29 (BA.H): + (CL.CW) - 3 9 . 6 6 1 0 . 9 1 8 8 6 . 5 8 29 C L - 1 1 8 . 1 0 7 1 0 . 6 5 1 8 6 . 0 8 3 0 BA.H.CL 5 0 . 4 5 0 0 . 247 8 4 . 9 3 32 b i o m a s s o f M.h. D.CL - 1 8 . 7 8 0 4 . 7 0 0 9 9 . 2 8 2 t w i g s D - 3 4 . 1 5 2 1 3 9 . 1 5 7 9 5 . 1 8 5 BA.H.CL 1 6 . 6 6 2 0 . 1 3 6 9 4 . 4 8 5 ( B A . H ) + (CL.CW) 0 . 1 4 3 0 . 2 9 2 9 4 . 2 9 5 CR.SURF 0.027 0 . 1 9 6 9 3 . 3 1 6 D 2.H * 1 5 . 2 8 6 2 . 7 2 7 9 1 . 3 9 6 b i o m a s s o f M.h. D.CL * - 1 . 8 5 7 4 . 9 8 0 8 8 . 1 6 8 f o l i a g e D * - 1 7 . 2 9 6 1 4 6 . 0 4 0 8 2 . 9 1 1 0 BA.H.CL * 3 6 . 4 0 8 0 . 1 4 1 8 0 . 5 2 1 1 D 2.H * 3 4 . 8 9 9 2 . 8 3 7 7 8 . 2 4 1 1 * s i g n i f i c a n t o n l y a t 5% l e v e l , o t h e r f i g u r e s s i g n i f i c a n t a t 1 % l e v e l . 95 S i g n i f i c a n t i n d e p e n d e n t v a r i a b l e s i n s i m p l e l o g a r i t h m i c r e g r e s s i o n e q u a t i o n s f o r b i o m a s s p r e d i c t i o n o f p a r t i c u l a r t r e e c o m p o n e n t s ( v a r i a b l e s a p p e a r i n o r d e r o f d e c r e a s i n g r 2 v a l u e s ) , S q u a m i s h s a m p l e p l o t , n ; M o u n t a i n hem-l o c k = 5; P a c i f i c s i l v e r f i r = 7, T r e e C o m p o n e n t T r e e I n d e p e n d e n t R e g r e s s , S E E ( v a r i a b l e &) S p . V a r i a b l e X. I n t e r c e p t C o e f f . r 2 ( l o g ) l o g w h o l e M.h. l o g D 2 . H 2 . 4 6 9 0.769 9 9 . 1 7 0 . 0 2 0 8 3 t r e e l o g D 3 . 7 3 3 2 . 0 1 6 9 8 . 6 6 0 . 0 2 6 4 1 b i o m a s s P . s . f . l o g D 2,H 2 . 2 7 9 0 . 8 9 9 9 9 . 6 5 0 . 0 2 8 2 8 l o g D 3 . 7 7 9 2 . 4 7 3 9 8 . 9 5 0 . 0 4 8 6 0 l o g w o o d M.h. l o g D 2.H + 2 . 3 1 9 0. 7 4 6 9 8 . 4 3 0 . 0 2 7 8 7 b i o m a s s l o g D 3 . 5 4 4 1.947 9 7 . 1 3 0 . 0 3 7 6 5 P . s . f . l o g D 2 . H + 2. 0 4 7 0 . 9 5 3 9 9 . 5 3 0 . 0 3 4 4 1 l o g D 3 . 6 3 6 2 . 6 1 8 9 8 . 6 1 0 . 0 5 9 4 3 b a r k M.h. l o g BA + 3.109 1.039 9 8 . 1 8 0 . 0 3 1 8 9 b i o m a s s l o g D 3. 0 0 0 2 . 0 7 8 9 8 . 1 7 0 . 0 3 1 9 0 l o g D S H 1 . 7 0 1 0 . 7 8 8 9 7 . 5 9 0 . 0 3 6 6 2 P . s . f . l o g D 2 . H 1.346 0 . 967 9 5 . 8 3 0 . 1 0 6 4 4 l o g BA + 3 . 0 9 6 1.327 9 4 . 6 5 0 . 1 2 0 5 9 l o g D 2.95 7 2 . 6 5 4 9 4 . 6 5 0 . 1 2 0 5 9 b i g M.h. l o g D 2.H.CW + 0.647 0 . 7 8 5 9 9 . 3 8 0 . 0 2 7 2 9 b r a n c h e s l o g D 2 . 8 0 0 3 . 0 7 4 9 9 . 1 0 0 . 0 3 2 9 1 b i o m a s s D.CL 1.017 0 . 0 7 4 9 7 . 8 3 0 . 0 5 1 2 4 D 2 . H 1.537 0 . 0 4 5 9 7 . 7 4 0 . 0 5 2 2 7 l o g D2.H 0.89 4 1 . 1 5 1 9 5 . 9 8 0 . 0 6 9 6 8 ( B A . H ) + (CL.CW) 1 . 2 9 1 0 . 0 0 5 9 4 . 7 1 0 . 0 7 9 9 8 CR.SURF 1.302 0 . 0 0 3 9 4 . 6 9 0 . 0 8 0 1 5 P . s . f . l o g D 2.6 6 5 2 . 4 9 3 9 1 . 9 0 0 . 1 4 1 5 3 D.CL + 1.269 0 . 0 5 3 9 0 . 3 0 0 . 1 5 4 8 5 l o g D 2 , H 1.169 0 . 8 9 0 8 9 . 3 3 0 . 1 6 2 4 3 D . C L 2 1 . 4 3 3 0 . 0 0 2 8 6 , 8 4 0 . 1 8 0 3 7 l o g D 2.H.CW 0 . 6 3 1 0 . 7 7 1 8 6 , 1 3 0 . 1 8 5 1 7 l o g s m a l l b r a n c h e s b i o m a s s M.h. n o o n e i n d e p e n d e n t v a r i a b l e i s s i g n i f i c a n t P . s . f , l o g D.CL + 0 . 8 6 1 0 . 7 6 0 9 5 . 0 6 0 . 0 5 8 7 5 l o g D . C L 2 0 . 4 2 3 0 . 5 2 1 9 4 . 5 2 0 . 0 6 1 8 7 l o g D 2,H 1.218 0 . 4 8 1 9 2 . 3 8 0 . 0 7 2 9 9 l o g D 2.H.CW 0.9 1 8 0 . 4 2 1 9 1 . 1 0 0 . 0 7 8 8 9 l o g D 2. 0 1 9 1.317 9 0 . 6 9 0 . 0 8 0 6 6 l o g CR.SURF - 1 . 0 2 2 1 . 1 3 3 9 0 . 0 0 0 . 0 8 3 6 2 96 T r e e C o m p o n e n t T r e e I n d e p e n d e n t R e g r e s s . S E E ( v a r i a b l e Y ) . S p . V a r i a b l e X, I n t e r c e p t C o e f f . r 2 ( l o g ) l o g t w i g a n d M.h. l o g D 2.H + D 2 H + 0 . 8 8 5 0 . 1 5 6 9 9 . 3 9 0 . 0 1 8 8 9 f o l i a g e l o g D.CL - 0 . 5 4 2 1.029 9 7 . 1 6 0 . 0 3 3 3 3 b i o m a s s l o g D.CL 0,567 1.297 9 5 . 9 3 0 . 0 3 9 9 2 l o g D * 2 . 4 3 4 1 . 6 5 1 8 8 . 3 1 0 . 0 6 7 6 5 l o g D 2.H.CW * 1.290 0 . 4 1 5 8 5 . 7 1 0 . 0 7 4 7 8 P . s . f . l o g D.CL + 0.879 1 . 0 3 8 9 7 . 4 1 0 . 0 5 7 3 9 l o g D . C L 2 0 . 2 7 6 0 . 7 1 3 9 7 . 2 4 0 . 0 5 9 2 8 l o g D 2 . H 1.374 0 . 6 4 8 9 2 . 0 2 0 . 1 0 0 8 0 l o g D 2.457 1.789 9 1 . 9 8 0 . 1 0 1 0 1 l o g D 2.H,CW 0 , 9 8 3 0 . 5 6 1 8 8 . 6 2 0 . 1 2 0 3 4 l o g t w i g M.h. l o g D . C L 2 + - 1 . 2 7 2 1 . 1 6 3 9 9 . 4 4 0 . 0 1 6 5 9 b i o m a s s l o g D.CL - 2 , 7 8 0 1.474 9 9 . 2 1 0 . 0 1 9 6 1 l o g D 2.1I.CW+D 2,H.CW 0.12 2 - 0 . 0 0 3 9 9 . 0 5 0 . 0 2 6 4 2 l o g D 2 . H 0.909 0.724 9 3 . 9 5 0 . 0 5 4 4 0 l o g D 2. 0 9 9 1 . 8 9 5 9 3 . 0 4 0 . 0 5 8 3 6 l o g f o l i a g e M.h. l o g D 2.H + 0 . 5 9 8 - 0 . 0 4 5 9 8 . 4 6 0 . 0 2 8 3 9 b i o m a s s l o g D . C L 2 - 0 . 5 3 7 0 . 9 3 6 9 0 . 4 5 0 . 0 5 7 6 8 l o g D.CL 0 . 4 7 8 1. 1 7 6 8 8 . 5 4 0 . 0 6 3 1 9 l o g D 2 . 1 6 8 1 . 4 8 6 8 0 . 2 7 0 . 0 3 2 9 1 + e q u a t i o n s u s e d f o r c a l c u l a t i n g b i o m a s s o n t h e w h o l e s a m p l e p l o t . * s i g n i f i c a n t o n l y a t 5% l e v e l , o t h e r v a r i a b l e s a r e s i g n i f i c a n t a t 1 % l e v e l . 97 S i g n i f i c a n t i n d p e n d e n t v a r i a b l e s i n s i m p l e l i n e a r r e g r e s s i o n e q u a t i o n s f o r b i o m a s s ^ p r e d i c t i o n o f p a r t i c u l a r t r e e c o m p o n e n t s ( v a r i a b l e s a p p e a r i n o r d e r o f d e c r e a s i n g r 2 v a l u e s ) . , H a n e y s a m p l e p l o t , n ; W e s t e r n h e m l o c k = 8, T r e e C o m p o n e n t ( v a r i a b l e Y ) . T r e e S p . I n d e p e n d e n t V a r i a b l e X. I n t e r c e p t R e g r e s s . C o e f f , r 2 S E E i n k g t r e e b i o m a s s W.h. D 2.H 7 1 . 6 7 8 1 5 5 . 5 2 4 9 6 . 2 2 70 w o o d b i o m a s s W.h. D 2 . H BA 4 8 . 8 5 6 - 9 . 2 3 6 9 0 , 9 5 9 3 3 4 2 . 3 8 0 9 6 . 3 3 9 8 . 1 1 4 0 29 b a r k b i o m a s s W.h. BA - 1 0 . 0 5 4 8 7 9 . 1 4 0 9 3 . 6 9 14 b i o m a s s o f W.h. D.CL - 2 7 . 1 4 7 1 3 . 1 6 1 9 1 . 5 5 14 b i g b r a n c h e s D 2.H.CW 1.759 2 . 5 5 4 8 7 . 9 7 16 BA.H.CL 2 . 9 1 9 1.042 8 7 . 6 4 16 CR.VOL - 8 . 7 1 7 0 . 3 9 8 8 8 . 7 8 16 CR.SURF - 3 0 . 6 4 6 0 . 5 4 4 8 7 . 2 9 17 CL - 9 9 . 3 1 0 9.100 8 0 . 7 8 21 b i o m a s s o f W.h. L - 7 2 . 4 9 3 5 . 6 4 7 8 7 . 3 9 1 0 s m a l l D.CL 8.504 7.119 7 5 . 9 7 14 b r a n c h e s D 2.H + CL - 1 8 . 7 8 0 3 . 5 6 5 7 2 . 6 9 15 CR.SURF 6.726 0 . 2 9 8 7 2 . 0 0 15 BA.H 1 8 . 8 0 7 1 3 . 1 0 1 7 0 . 7 2 15 b i o m a s s o f W.h. L - 8 2 . 7 5 9 5 . 6 9 5 9 3 . 0 0 7 t w i g s a n d D.CL - 2 . 6 0 1 7 . 4 8 0 8 7 . 7 4 1 0 f o l i a g e CW 2 - 3 . 7 5 8 1 . 459 8 5 . 5 8 1 0 BA.H 7.680 1 4 . 0 3 9 8 4 . 9 6 1 1 CR.VOL 8. 3 8 5 0 . 2 2 2 8 2 . 0 1 12 CR.SURF - 4 . 1 9 7 0 . 3 0 6 8 2 . 0 4 12 b i o m a s s o f W.h. CW 2 - 0 . 6 2 1 0 . 4 8 6 9 0 . 8 7 3 t w i g s ( B A . H ) + (CL.CW) - 0 . 6 1 8 0 . 1 6 0 8 5 . 1 2 3 CR.VOL 3 . 4 6 8 0.074 8 6 . 2 3 3 CR,SURF - 0 . 6 3 3 0 . 1 0 1 8 5 . 3 6 3 L - 2 3 . 7 5 1 1 . 7 4 4 8 3 . 4 3 4 D.CL 0 . 7 7 2 2 . 2 9 6 7 9 . 0 5 4 b i o m a s s o f W.h. L - 5 8 . 9 0 7 3 . 9 4 6 9 3 . 9 0 5 f o l i a g e D.CL - 3 . 3 4 0 5 . 1 7 6 8 8 . 3 6 6 BA.H 3 . 3 4 0 5 . 1 7 6 8 7 . 1 3 7 B A - 1 . 0 0 2 2 7 9 . 2 1 8 8 6 . 4 4 7 CR.SURF - 3 . 5 3 0 0 . 2 0 5 7 7 . 2 4 9 ( B A . H ) + (CL.CW) - 3 . 5 1 5 0 . 3 2 5 7 7 . 1 3 9 98 S i g n i f i c a n t i n d e p e n d e n t v a r i a b l e s i n s i m p l e l o g a r i t h m i c r e g r e s s i o n e q u a t i o n s f o r b i o m a s s p r e d i c t i o n o f p a r t i c u l a r t r e e c o m p o n e n t s ( v a r i a b l e s a p p e a r i n o r d e r o f d e c r e a s i n g r 2 v a l u e s ) . , H a n e y s a m p l e p l o t , n ; W e s t e r n h e m l o c k = 8, T r e e C o m p o n e n t T r e e I n d e p e n d e n t R e g r e s s , S E E ( v a r i a b l e Y ) S p . V a r i a b l e X I n t e r c e p t C o e f f . r 2 ( l o g ) l o g w h o l e W.h. l o g D 2.H 2.32 7 0 . 8 8 2 9 8 . 9 1 0 . 0 4 3 0 9 t r e e l o g D 3. 6 8 0 2 . 1 3 5 9 6 . 7 3 0 . 0 7 4 4 4 b i o m a s s l o g w o o d W.h. l o g D 2,H + 2 , 1 1 2 0 . 8 7 0 9 9 . 2 5 0 . 0 3 5 1 5 b i o m a s s l o g D 3. 4 5 5 2 . 1 2 0 9 8 . 3 2 0 . 0 5 2 5 7 l o g b a r k W.h. l o g D 2,H + 1 . 4 1 5 0 . 9 9 6 9 7 . 2 1 0 . 0 7 8 4 2 b i o m a s s l o g D 2. 9 3 4 2 . 3 9 4 9 3 . 7 3 0 . 1 1 7 4 8 l o g BA 3. 0 6 0 1.197 9 3 . 7 3 0 . 1 1 7 4 8 l o g b i g W.h. l o g D + 3 . 2 7 0 3 . 8 6 8 7 7 . 8 3 0 . 3 9 1 7 1 b r a n c h e s l o g D 2.H 0. 8 2 5 1 . 5 7 0 7 6 . 9 4 0 . 3 9 9 5 3 b i o m a s s l o g D 2.H.CW 0,07 2 1 . 2 3 1 7 5 . 4 0 0 . 4 1 2 6 6 l o g s m a l l W.h. l o g D 2.H.CW + 1.177 0 . 4 9 9 9 2 . 7 2 0 . 0 8 2 1 4 b r a n c h e s l o g D.CL 0. 9 3 6 1.007 9 1 . 1 9 0 . 0 9 1 3 2 b i o m a s s l o g D 2 . H 1.427 0 . 6 2 0 8 9 . 6 0 0 . 0 9 8 1 4 l o g D . C L 2 0.317 0 . 697 8 6 . 2 4 0 . 1 1 2 8 8 l o g D 2 . 3 6 6 1.477 8 4 . 8 3 0 . 1 1 8 5 3 l o g CR.VOL 0.406 0 . 6 0 2 6 9 . 4 2 0 . 1 6 8 3 0 l o g t w i g a n d W.h. l o g D.CL + 0 . 6 2 8 1 . 2 7 2 9 5 . 5 8 0 . 0 7 8 9 2 f o l i a g e l o g D 2.H.CW 0 . 8 6 3 0 . 6 2 6 9 5 . 5 1 0 . 0 7 9 6 1 b i o m a s s l o g D . C L 2 - 0 . 1 6 5 0 . 8 8 8 9 1 . 7 3 0 . 1 0 8 0 1 l o g D 2.H 0. 1 2 5 0 . 7 7 4 9 1 . 8 0 0 . 1 0 7 5 5 l o g D 2 . 4 2 0 1.837 8 6 . 1 3 0 . 1 3 9 9 1 l o g CR.VOL - 0 . 0 5 3 0 . 7 6 7 7 4 . 0 0 0 . 1 9 1 5 4 l o g t w i g W.h. ( B A . H ) . + (CL.CW). + 0.564 0. 0 0 6 8 2 . 8 1 0 . 1 2 7 3 0 b i o m a s s l o g C R.SURF 0 . 5 6 3 0 . 0 0 3 8 3 . 0 2 0 . 1 2 6 5 0 CR.VOL 0 . 7 0 8 0 . 0 0 3 . 8 2 . 1 3 0 . 1 2 9 7 9 l o g D . C L 2 - 0 . 2 1 3 0 . 6 7 9 8 0 . 3 9 0 . 1 3 5 9 7 l o g D.CL 0.42 2 0 . 9 3 1 7 6 . 6 0 0 . 1 4 8 5 1 l o g D2.H.CW 0 . 5 6 0 0 . 4 5 1 7 4 . 2 1 0 . 1 5 5 9 1 l o g f o l i a g e W.h. l o g D.CL+D.CL + - 0 . 1 5 0 - 0 . 1 4 4 9 7 . 4 5 0 . 0 8 6 3 3 b i o m a s s l o g D 2.H 0 . 9 5 1 1 . 0 2 2 9 2 . 4 3 0 . 1 3 5 2 8 l o g D 2,H.CW 0.455 0 . 807 9 1 . 9 0 0 . 1 4 0 4 9 l o g D 2. 5 0 8 2 . 4 5 4 8 8 . 9 8 0 . 1 6 3 9 4 l o g D . C L 2 - 0 . 8 1 6 1.114 8 3 . 6 5 0 . 1 9 9 6 7 + e q u a t i o n s u s e d f o r c a l c u l a t i n g b i o m a s s f o r t h e w h o l e s a m p l e p l o t . A l l v a r i a b l e s s i g n i f i c a n t a t 1 % l e v e l . A P P E N D I X 4 BIOMASS D I S T R I B U T I O N FOR 2 0 SAMPLED T R E E S ON THE SQUAMISH SAMPLE PLOT S P E C I E S 1 - MOUNTAIN HEMLOCK S P E C I E S 2 - P A C I F I C S I L V E R F I R • o o © © o o o o o J-o TREE SPECIE AGE LENGTH LENGTH LENGTH DBH CROWN CROWN BUTT BRANCH BRANCH CONES^  TWIGS FOLIAGE _TWIG*FOL NO TO 1 TO 6 OIAM LENG LENG OVER I IT01/4 (H) (M) (M) (M) (M) (M) (M) (KG) (KG) (KG) (KG) (KG) (KG) 22 440 37,3 37,0 33.6 0,780 7,5 25,9 11,4 411,4 66,4 0,6 0,0 0,0 TOP —1_TWIGS ANO FOLIAGE _ MID —2 ARE NOT SEPARATED LOW — 3 FOR ABIES AMAB1LIS 197,6 56,6 " 5 8 . 1 ' 83.0 SECTION N0| 1 2 J 4 5 6 7 a 9 10 WOOD 446,4 383,4 339,0 290,3 233,2 186,0 164,5 125,1 31,6 6.1 SUM WOODS 2205,8 KG BARK 75,4 47,1 42,2 41,0 35,1 49,8 53.1 41.5 18,2 » » * SUM BARK* 405,S KG TREE BIOMASS" 73 3;03f3 KG TREE SPECIE AGE LENGTH LENGTH LENGTH DBH CROWN CROWN BUTT BRANCH BRANCH CONES NO TO 1 TO 6 DIAM LENG LENG OVF.R 1 1T01/4 . ( M j TWIGS FOLIAGE TWIG*FOL (M) J i l l . ITO rsy 14,7 0,356 TMT 6.2 41 300 (M) t«».2 (M) 2.7 (KG) 33,2 (KG) 31.4 (KG) 0,2 (KG) 0,0 TKG) 0,0 (KGT 57.7 TOP--1 TWIGS AND FOLIAGE MID—2 ARE NOT SEPARATED LOW —3 FOR ABIES AMABILIS 9,6 29,3 18,8 SECTION" NO I 1 - 2 - — j j j - 5 - 6 r fl 9 _ WOOD 63,6 57,5 50,8 43.6 29,4 24,7 13.7 8,1 3,7 " 10" 1.1 SUM WOOOs 296,2 KG BARK 6,6 9,4 9,7 5,9 4,1 3,9 4,2 1,7 0,5 0,2 SUM BARKs 46,3 KG TREE BIOMASSs ' *t&S$Q KG TREE SPECIE ACE LENGTH LENGTH LENGTH OBH CROWN CROWN BUTT BRANCH BRANCH CONES _TWIGS_ FOLIAGE TWIG*FOL NO TO 1 (M) (H) TO 6 (H) (M) OIAM LENG LENG (M) CM) CM) OVER I" 1T01/4 (KG) (KG) (KG) (KG) (KG) (KG) 45 1 SOO 35,2 23,9 30,3 0.754 0,0 29,2 6,0 0,0 0,0 0,0 TOP--1 0,0 0,0 0.0 0,0 0,0 0,0 MID--2 LOW—.3 0,0 0,0 0,0 0,0 0,0 0.0 SECTION NOl 1 2 3 4 5 6 7 8 9 10 MOOD 544,1 443,7 362,5 288,8 221,0 169,3 120,3 63,0 41,4 8,6 SUM WOODS 2282,7 KG BARK 65,7 102,6 81.0 67.5 42.6 30.0 30,9 16,2 7,8 1,7 SUM BARKs 465,9 KG TREE BIOMASS8 2748,6 KG ' f t * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * TREE SPECIE AGE LENGTH LENGTH NO TO I LENGTH OBH TO 6 CROMN CROWN BUTT DIAM LENG LENG BRANCH BRANCH OVER 1 1T01/4 CONES TWIGS FOLIAGE TWIG»FOL (H) CM) CM) (M) CM) (M) CM) (KG) (KG) (KG) (KG) (KG) (KG) 55 1 460 34,A 33,6 2B,9 0',681 6,0 28,2 6,2 0,0 0,0 0,0 0,0 0,0 0,0 TOP—-1 MID--2 0,0 0,0 0,0 0,0 0,0 0,0 LOW--3 0,0 0,0 0,0 SECTION NOl 1 2 3 4 5 6 7 8 9 "to NOOO 415,4 357,5 269,6 227,4 198,3 135,9 88,3 60,1 22,3 5,2 SUM WOOD» 1779,9 KG BARK 94,0 68,1 74,6 62,3 37,7 34,2 23,9 15,4 4,7 1,1 SUM BARKa 436,0 KG TREE BIOMASSs 2216.0 KG •»«**«»«*»«***»*»****»*»»*»***»«**««»»»»**********«»****»**»*******«^ e TREE SPECIE AGE LENGTH LENGTH LENGTH OBH CROWN CROWN BUTT BRANCH BRANCH CONES TWIGS FOLIAGE TWIG+FOL 9 NO TO 1 CM) (H) TO 6 (M) (M) DIAM " LENG LENG CM) CM) (M) OVER 1 IT01/4 (KG) CKG) (KG) (KG) (KG) (KG) 69 1 325 21,3 20,7 14,7 0,320 0,0 12,4 9,0 0,0 0,0 0,0 0,0 0,0 0.0 © TOP--1 0,0 0,0 0,0 MI0--2 LOW—3 0,0 0,0 0,0 0,0 0,0 0,0 © SECTION NOl 1 2 3 4 5 6 7 8 9 10 MOOD 66,4 53,7 41,8 36,7 29,8 22,1 17,8 11,0 5,B 1,6 SUM WOODa 287,0 KG 0 BARK 9,2 13.7 10.0 8.3 6.1 5.0 3,2 1,6 0,6 0,3 SUM BARKa 58,0 KG TREE BIOMASSa 345,0 KG o o •»»»»»»»»«*»*«*«»•*»«•*»»»*»*«•»*»*****«**«*«***»******»**»* o o TREE SPECIE AGE LENGTH LENGTH NO TO 1 LENGTH DBH TO 6 CROWN CROWN BUTT DIAM LENG LENG BRANCH BRANCH OVER 1 1T01/4 CONES TWIGS FOLIAGE TWIG+FOL (M) (M) CM) (M) CM) CM) CM) (KG) (KG) (KG) (KG) (KG) (KG) o 66 1 530 38,0 36,9 33,4 0,711 7,9 24,5 13,5 217,5 31,3 0,0_ 64,2 84,0 148,2 0 TOP —1 MID — 2 14,3 11,8 16,6 15,5 30,8 27.3 o LOW — 3 38,1 51.9 90,1 SECTION NOl 1 2 3 4 5 6 7 8 9 10 •J MOOD 380,9 361,2 316,0 260,6 227,4 185,7 129,5 83,7 39,4 9,3 SUM WOODS 1993.7 KG BARK 64,2 76,8 86,3 92,8 60,5 36,7 30,4 22,9 15.4 5,3 SUM BARKs 491,4 KG TREE BIOMASSS KG •»*»»»»»*»*»**•*»««•«***»«»«»*»**«»***»»«******«****«*********•** o ] & <* TREE SPECIE AGE LENGTH LENGTH LENGTH OBH CROWN CROWN BUTT BRANCH BRANCH CONES TWIGS ; FOLIAGE TWIG*FOL NO TO 1 (M) (M) TO 6 DIAM (M) (M) (M) LENG (M) LENG OVER 1 "1T01/4 CM) CKG) CKG) (KG) CKG) (KG) • CKG) 7a I 410 34,2 33,4 28,6 0,762 9,4 25.2 9,0 289,6 53,8 0,0 70.1 90,5 160,6 TOP—1 32.9 30,9 63.8 MID—2 LOW — 3 17.9 19.3 23.7 35,9 41,6 55.2 SECTION NOI 1 2 3 4 5 6 7 8 9 10 WOOD 449,5 359,0 292,6 256,6 221.9 143,6 107,5 70,7 22 ,7 3,6 SUM WOOOn 1927,7 KG BARK 93,4 117,5 108,7 80,2 39,2 65,1 44,9 19,3 5 ,5 1,2 SUM BARKs 575,0 KG TREE BIOMASSs -SOOb,£ KG ft**********,.,*,*.****,.*,**!)******,*•**«***,•*•**«»*** A********.******************,*********************,,*,,•****•*•••*•**•<••*<* TREE SPECIE AGE LENGTH LENGTH NO TO 1 LENGTH OBH CROWN TO 6 DIAM CROWN LENG BUTT BRANCH BRANCH LENG OVER 1 1T01/4 CONES TWIGS FOLIAGE TWIG*FOL (M) CM) (M) (M) (H) CM) CM) CKG) CKG) CKG) CKG) CKG) (KG) 80 1 480 31,2 30,5 2 5 , 1 0 . 5 6 4 6.2 25,8 .5,5 1 0 7 , 1 3 5 , 5 0,0 51,6 70,1 121.7 TOP —1 MID—2 16,4 25,2 17,2 35,4 33.6 60,6 LOW — 3 10,1 17.S 27,5 SECTION NO 1 1 2 3 4 s ^ ? * 8 9 10 • » HOOD 260,5 216,2 174,5 135,4 103.2 75,4 55,8 26,3 10 ,5 2,1 SUM WOODs 1059,9 KG BARK 55,2 62,9 38,9 37,4 21,4 19,8 25,6 11,3 4 ,0 1,2 SUM BARKs 277,5 KG TREE BIOMASSs **»******»******»***«**»************************»***********^ . v : : _ : , to** U-: • j J o 9 • • • • M KI m o CO _» r- CO O! if. •< U-i X o CD bJ CO; UJ U •C * -• 0C •-t fr-_t d GO KI f\j Ai \- yj ^* * * w et rvj KI • l l eo . 1 t i Ui; «-» o OL • • z CJ • O •-• o CO o l »- a: _ i o »-» o ru —• X CJ 3 o a n z X, «»•» •> D ac o CD o ac CE O o 4 cc *^ m X X X z> => L> in CO CO z * « a: CD 9 or UJ it CO m > o m o o • <• i n »-«— u 3 z> z X * CD Ui CO o • * in z rvi X o ca «*>» * CE z X KI CO m o U Ui w •> • Kt 9 z r o X * r» i v CO or < X • • «j o o «o -* X CD Ki -o KI a X in •> • ^* • a-o . r*. X IA Ki o z -0 «~v • •> • Ul X rw _ J o w AI KI »-X • 9 in Kt • o 9 z •* • Kt K» UJ X _* o w KI K» m 9 X m CO ¥- •« m 13 •> z %^ •> Ui x • •* ru 9 —* KI • ' • o m Ui ru o O CO CO KI • * o in at 9 m ru u o Ui <•* z o OL o or t» z o •< o m *-4 i : X Z "V CE U l CD > O 1— IS Z3 Z CD Ui O O CE Z U Ui O X CE < X CO z « Ui _ i o X i — o CE O t - Z Kt ru nj ^ ( C O u. a. UJ O CO CO t-< UJ 13C 3t CE iO •< [U. f\J 'to o ,z « « * « « « « e*- * * * to * « * « « * n « CO * CO 4 * X « o * « CD « « Ui UJ « tr. « ¥— « « « « o o * ^c * * * CO in « « in X * 4> CO « o •o KI * « n u « o * o ac * o <t « X CO * « X X * * 00 to * * a « m Kt « ru * p» « « ru * ru « « ' 'i * ' o * • • * ru « ru KI « -w ; « : ru ; • • * KI in 1 CO KI « • • « 9 * ru « ru « KI CO * ! ru 9 « o O" i KI « | « • | i n o * 1 • * KI « eo Kl « KI v« « i « j O Kt « • • - * ru * Kl o « m « « o « • o- O m « r- « IT. « « tn o « • • « « 9 CO « « « « O « o or * o •< <* X CD - « « « « TREE SPECIE AGE LENGTH LENGTH LENGTH_ OBH CROWN CROWN BUTT BRANCH_ BRANCH CONES TWIGS FOLIAGE TWIG*FOL_ NO TO 1 TO 6 "' ' ~ D I A M LENG LENG OVER 1 1T01/4 (M) (M) (M) (M) (M) (M) (M) (KG) (KG) (KG) (KG) (KG) (KG) I 19 1 410 25.1 2«t» 21,5 0,437 4,7 20,6 4,5 53.1 23,8 0,0 TOP — 1 MID. LOW-23.6 _S.6_ 10,5 7,4 35.5 10.9 1 7 , 6 " 7.0 59,1 16,5 "28,2" 1«,5 SECTION NOI 1 10 WOOD 147,1 123,9 107,4 91,4 77,3 60,2 45,9 27,8 17,3 3,9 SUM WOOD" 702,2 KG BARK 32.2 31,5 29.8 26.5 18,1 iq,8 U . 7 7.2 5.1 1,2 SUM BARK" 178,1 KG TREE BIOMASS" fO/C',3 KG *************************************************** TREE SPECIE AGE LENGTH LENGTH LENGTH OBH CROWN CROWN BUTT BRANCH BRANCH CONES NO TO 1 TO 6 DIAM LENG LENG OVER 1 ITO 1/4 TWIGS FOLIAGE TWIG*FOL 159 018 (M) 29,9 (M) 29,0 (M) (M) 26,1 0,485 (M) 8,1 (M) (M) 18,1 11,8 (KG) 53,3 (KG) 41,9 (KG) TOP-MID" (KG) 0,0 (KG) 0,0 •1 TWIG3 AND FOLIAGE •2 ARE NOT SEPARATED LOW—-3 FOR ABIES AMABILIS (KG) 59.2 29.2 20.8 9.2 SECTION NO I 1 "2 J 4 5 T " " 8 ~ ~ " 9 " 1 0 " " WOOD 182,7 151,7 132,4 103.0 87.2 65.6 50.2 34.2 19.7 5,6 SUM WOOQ" 832.2 KG  BARK 37,9 38,3 34,6 28,7 30,8 23,4 16,2 11,0 5,7 2,6 SUM BARK" 229,0 KG TREE BIOMASS" 7-2.767° K<5 **««»««»««»««**««««»»«*«*«***.»**»****«»******««**********************«***«^ TREE SPECIE ACE LENGTH LENGTH LENGTH OBH CROWN_ CROWN BUTT BRANCH_ BRANCH CONES TWIGS NO TO 1 TO 6 DIAM LENG LENG OVER 1 1T01/4 (M) CM) (M) CM) CM) CM) CM) CKG) CKG) CKG) CKG) FOLIAGE TWIG*F0L_ CKG) CKG) 33,9 0,711 8,5 28,9 9,6 174,0 ci 175 530 38,4 38,0 73,6 0.3 0,0 0,0 TOP—1 TWIGS AND FOLIAGE MID — 2 ARE NOT "SEPARATEO"~ L0W--3 FOR ABIES AMABILI3 SECTION NOt 1 10 191,0 68,0 ~78.5~ 44,5 WOOD 409,3 336,3 286,9 243,5 192,4 148,9 106,4 76,7 37,9 10,2 SUM WOODa 1848,6 KG BARK 88.5 69,6 59.2 31.1 35.6 19.3 14.5 9.7 6,4 2.4 SUM BARK" 336.4 KG TREE BIOMASS" £6£ 3 , 9 KG ************************************************************************** TREE SPECIE AGE LENGTH LENGTH LENGTH OBH CROWN CROWN BUTT BRANCH BRANCH CONES NO TO 1 TO 6 DIAM LENG LENG OVER 1 1T01/4 CM) TWIGS FOLIAGE TWIGtPOL 176 335 CM) 26,3 25.5 CM) ( M l -lS,8 0,361 CM) 0,0 CM) 19,6 (M) 6,6 CKG) 0,0 (KG) 0,0 w 0,0 "(KG) 0,0 (KG) 0,0 TOP—1 TWIGS AND FOLIAGE MID—2 ARE NOT SEPARATED LOW — 5 FOR ABIES AMABILIS (KG 5 0,0 0,0 0,0 0,0 SECTION NO I 8 "10 WOOD 91,9 66.7 58.4 49,8 37.2 30,3 21.7 14.3 7.0 1.6 SUM WOODa 378.9 KG BARK 14,0 14,1 8,8 7,1 5,7 5,0 2,9 1,9 1,2 0,4 SUM BARKs 61,1 KG TREE BIOMASS" 4 - ; KG ********************************************************************************************************************************** TREE SPECIE _AGE _ LENGTH LENGTH_LENGTH DBH_ CROWN CROWN _BUTT_ BRANCH BRANCH CONES TWIGS FOLIAGE TWIG*FOL NO TO 1 TO 6 DIAM LENG LENG OVER 1 1T01/4 ~ "" "" (M) (M) (M) (M) (M) (M) (M) (KG) (KG) (KG) (KG) (KG) (KG) TTB 2 STo 38,9 38,5 32,3 o,88n b~7o 28,7 roTz o76 oTo b~7o : oTo oTo 'oTo , . T0P--1_TWIGS AND FOLIAGE 0,0 MID — 2" ARE" NOT "SEPARATED 0,0 LOW —3 FOR ABIES AMABILIS 0.0 SECTION NOl 1 2 3 0 5 6 7 8 9 10 S0OO~fj2,6 487,5~449,0 383,8 318,7 25T,TT6lu8HfTr,9 2~3.T if,l WTWoblyi 2931~6 KG : BARK 136.3 106.2 95.0 89.1 110.7 90.1 65.2 36.4 9,6 1.8 SUM BARK" 740.4 KG TREE BIOMASS" 3672,0 KG  »«**»»*•**»»«««*««**«••»***«»»«•*«**«*»»»**»**»******« * * * * * «k * *^<r* ****jr*****«^»»***ir******»r** * vr** * * *«*~* * *"* «r*~* *~sr«~*^rw * * * * »irir^ «^** v TREE SPECIE AGE LENGTH LENGTH LENGTH DBH CROWN CROWN BUTT BRANCH BRANCH CONES TWIGS FOLIAGE TWIG+FOL NO , TO 1 TO 6 DIAM LENG LENG OVER 1 IT01/4 (M) (M) (M) fMl CM) (M) (M) (KG) (KG) (KG) (KG) CKGl (KG) 213 2 365 30 «. 0 29,5 _2 4 , l \ , O.S64 5,6 25,1 4,9 92,1 51,3 0,5 0.0 0,0 116,3 TOP—1 TWIGS AND FOLIAGE 32.6 : : ; MID—2 ARE NOT SEPARATED 52,1 LOW —3 FOR ABIES AMABILIS 31,6 " SECTION NOi 1 2 " 3 4 5 6 7 " " 8 9 10 " " " """ WOOD 218,2 174,2 142,3 118,5 88,2 69,9 49.5 27,4 12,7 2,5 SUM WOOD" 903.4 KG ; BARK 53,7 38,3 25,8 22,7 23,3 15,4 11,7 3,8 3,9 1,4 SUM BARK" 200,0 KG TREE BIOMASS" I3&3t7. . KG ^ : : : : : : ) TREE SPECIE AGE _ LENGTH LENGTH LENGTH OBH CROWN__CROWN BUTT_ BRANCH BRANCH CONES NO TO i TO 6 DIAM LENG LENG OVER i 1T01/4 (M) (M) (M) CM) CM) CM) CM) CKG) (KG) CKG) TWIG»FOL CKG) 215 440 36,2 37.7 33.0 0,805 0,0 25,7 12.5 0,0 0.0 0,0 0,0 0,0 TOP-^1 TWIGS AND FOLIAGE "MI6 — 2 ARENOT SEPARATED" L0W--3 FOR ABIES AMABILIS 0,0 0.0 "0,0" 0,0 SECTION NOI 1 2 3 a 5 6 7 8 9 10 WOOD 480,2 299,9 306,5 266,3 216,5 171,0 112,6 87,0 33,9 5.2 SUM WOOD" 1979,0 KG BARK 125.2 81.0 80.8 77.1 89.6 72,« an.2 34,7 13.0 1.2 SUM BARKs 619,2 KG TREE BIOMASSs 2598,2 KG 0 •»*•••««**»«»»«««»•»*••»•««*««*****«»»•**«*****•»**»***•»*** © TREE SPECIE AGE LENGTH LENGTH LENGTH DBH CROWN CROWN BUTT BRANCH BRANCH CONES NO TO 1 TO 6 DIAM LENG LENG OVER 1 1T01/4 " CM)" TWIGS FOLIAGE TWIG*F0L 235 355 CM) 15.1 "W"" 14,6 CM) 12,5 0.343 CM) 0,0 CM) 8.4 CM) 6,6 (KG) 0,0 (KG) 0,0 "(KG) 0,0 (KG) 0,0 TKG") 0,0 T K G F " 0,0 TOP-M TWIGS AND FOLIAGE _MID«-2 ARENOT SEPARATED LOW—-3 t : o f ) ABIES AMABILIS 0,0 0,0 6,0 "SECTION NO! 1 ~r~ "3 W 5 6" 7 8 9" WOOD 48.2 41.4 36,6 29.8 25.7 22.7 16,2 10.9 8,1 1,5 SUM WOODS 241,0 KG BARK 8,1 8,1 7,8 7,4 5,2 5,2 3,9 3,8 1,4 0,5 SUM BARKs 51,3 KG TREiE BIOMASSs 292,4 KG **************** TREE SPECIE AGE LENGTH LENGTH_ LENGTH OBH CROWN_ CROWN BUTT_ BRANCH BRANCH CONES TWIGS FOLIAGE TWJG + FOL NO to i to 6 " O I A M " XENG "CENG" OVER i ¥Toi/a " (M) (M) (M) (M) (M) CM) (M) CKG) (KG) CKG) CKG) CKG) CKG) 243 380 J7.8 17.1 11,9 0,310 5,5 11,5 6,3 33,8 17,9 0,0 0,0 0,0 TOP —1_TWIGS AND FOLIAGE MID—2 ARE NOT SEPARATED LOW--3 FOR ABIES AMABILIS 30,0 6,B_ 17,8 5.4 SECTION NOI 1 8 10 WOOD 40,0 36,3 27,7 22,6 16,5 12,7 9,3 5,6 3,5 1,1 SUM WOOD* 175,4 KG BARK 5.5 7.0 6.8 5.8 4,8 3.1 2,1 1,4 0,7 0.3 SUM BARK" 37,5 KG TREE BIOMASS" KG ********************************************************************************************************************************** TREE SPECIE AGE LENGTH LENGTH LENGTH OBH CROWN CROWN BUTT BRANCH BRANCH CONES NO TO 1 TO 6 DIAM LENG LENG OVER I IT01/4 256 403 CM) CM) 27,9 27,5_ TM) CMT^ 19,5 0,475 CM) 0i°__ (M) 20,3 CM) 7.6 CKG) 0,0 (KG) 0,0 (KG) 0,0 TWIGS TKGT 0,0 FOLIAGE TWIG*FOL CKG) 0,0 TOP--1 TWIGS AND FOLIAGE MID—-2 ARE NOT SEPARATED CKG) 0.0 0.0 0,0 LOW--3 FOR ABIES AMABILIS 0,0 SECTION Nb i t '"""" 2 3 "a" s -6 — 7 - 8 9 1 0 WOOD 167,5 125,8 103,9 93.6 77.2 54,0 43,0 28,6 10,8 2,7 SUM WOOD* 707,3 KG BARK 27,6 29,2 29,5 20,2 17,4 10,4 10,1 8,4 1,6 0,7 SUM BARK" 155,1 KG TREE BIOMASS" 862,4 KG ****************************************************************************************************************** **************** A P P E N D I X 5 BIOMASS D I S T R I B U T I O N FOR 1 3 SAMPLED T R E E S ON THE HANEY SAMPLE PLOT S P E C I E S 1 - WESTERN HEMLOCK S P E C I E S 2 - WESTERN RED CEDAR S P E C I E S 3 - YELLOW CEDAR o o o o o o 0 o o o o TREE SPECIE ACE LENGTH LENGTH LENGTH DBH CROWN CROWN BUTT BRANCH BRANCH CONES _MO : 10-1 TO 6 DIAM LENG_ LENG„_O.V.ER_l IT017.4 (M) (M) (M) (M) CM) CM) CM) CKG) (KG) CKG) TWIGS _S0_ -17 Q 1J..L -15... 2_ 2,7 0.1611 *US 1A.3 2V9- _li*2_ _uuuu_ CKG) T...Q.-FOLIAGE TKT) 3 . 9 -TWIG*FOL CKG) TOP—1 M ID—2 LOW — 3 1.5 1.8 0,8 .2.1. 1.0 2,4 5.8 2.7 SECTION NOl 1 2 3 4 5 w. 0 Q D 15 , A 11 , 2_ 9 . 6 I*7_ BARK 2,5 1,9 1,7 2,2 1,3 1,2 6 7 8 9 10 _4,1 2,4 1,5 -0,5 3UM_-.W_Q.0DB .66,.3___.KG__ 0,7 0,6 0,3 0,3 SUM BARK* 12,7 KG TREE BIOMASSs jO%,±' KG •»«•«•»•«»»»»«*«««*•«,•»•««*»** »*»«***»**».»•*****»*«*«* * * * ** • * »*_*«_**** * *.****.**.•* * * «««»«««»«««««»«««»«*««««»»«« * ********* *_*_**.**._ TREE SPECIE AGE LENGTH LENGTH LENGTH DBH CROWN CROWN BUTT BRANCH BRANCH CONES TWIGS FOLIAGE TWIGtPOL NO TO 1 TO 6 DIAM LENG LENG OVER 1 1T01/4 (M) CM) CM) CM) (M) CM) (M) (KG) (KG) (KG) (KG) (KG) (KG) e 54 1 410 22,4 20,9 14,8 0,394 1.7 14,8 7,6 35,4 52,3 UUUU U . l 28,5 39,6 o TOP—1 M10—2 LOW — 3 2.1 5,6 3,5 6,2 15,2 7.1 8,2 20,8 10.6 . aECi icm NOJ. i I _1_ . 4 ? 6 _7. 8 9 10 WOOD 77,a 69,7 65,4 60,7 51,6 52,5 36,3 8,9 3,0 0,9 SUM WOOD* 427,0 KG BARK 16,7 14,6 14,3 11,8 12,4 10,6 7,1 2.1 2.5 0,4 SUM BARKS 92,5 KG TREE BIOMASSs •»•«•».«.»««•».**.««»»»*»»»««•««..«*****•»•«**«**•**»****»**«»*****«*•*«**«********»*««**«*««**•***«**«««****••«»•««•*.•***«»•***» 9 TREE SPECIE AGE LENGTH LENGTH LENGTH _H0 . 1.0-4. _tXL.6. _6JJ_ aoo (N> _2.a,u_ (M) CM) _44-*6-0l5. J 3 4 OBH CROHN CROWN BUTT BRANCH BRANCH CONES DIAM LENG LENG OVER-1—U.O.I/.« (M) (M) (M) (M) (KG) (KG) (KG) TWIGS FOLIAGE TWIG»FOL _ruu_ -.D.,.0_ _uuuu_ (KG) O..JL (KG) 0,0, T O P " ! TWIGS ANO FOLIAGE _MID»-2._.ARE.NUT...SEP.ARA TED-LOW—3 FOR CEDAR (KG) 0...0-0,0 .0,0 0,0 e> & SECTION NOI 1 2 3 4 5 6 7 8 9 10 _K0QD_44.T.*L_J).3.,.9....^ A.8 JUK-WOOD* 612,8 KG , BARK 12,9 14,5 11,1 7,0 9,5 0,0 0,0 0,0 0,0 0,0 SUM BARK" 55,0 KG TREE BIOMASS" 667,9 KG A * * * * * * * * * * * * * * * * » * * * « « * ***_»«_****_***_*_* **Jl*JL* ** i i i ************* ***** ****** *** *.* ****** *.* * +.***±**lL**L*±^**^^^±t-±^^J!^** **••*• ***** **~ TREE SPECIE AGE LENGTH LENGTH LENGTH DBH CROWN CROWN BUTT BRANCH BRANCH H 0 TO I TO 6 DIAM LENG LENG OVER 1 IT01/4 • (M) (M) (M) (M) (M) (M) JMJ OXGJ £J<GJ CONES (KG) TWIGS (KG) 65 1 340 23,5 22,3 15,7 0^343 6,1 18,5 4,9 80,7 73,4 UUUU TOP —1 MID—2 LOW—3 17,3 0.3 7.3 5.7 FOLIAGE (KG) 30,9 7.4 12.7 10.8 TWIG+POL (KG) 48,2 11.7 20,0 SECIX.QJLJiO.1 l_ MOOD 65,2 72,9 61,6 42,8 34.1 25,6 16.3 10,0 S.tt 1.8 SUM WOOD" 335,7 KG BARK 15,4 16,5 11.5 10.9 T7o 5^5 W 271 T72 074 SUM BARK" 74,6 KG TREE BIOMASS" KG TREE SPECIE AGE LENGTH LENGTH NO TO 1 LENGTH TO 6 DBH CROWN DIAM CROWN BUTT LENG LENG BRANCH BRANCH OVER t 1T01/0 CONES TWIGS FOLIAGE TWIG+POL (M) (M) (M) (M) (M) (M) (M) (KG) (KG) (KG) (KG) (KG) (KG) tyo | >\h 18,9 t 7, o 10,2 0,221 10,0 0,9 1,? 25,0 mum S 1 13,2 18,1 c TOP—-1 MIO — 2 2.1 ..2.4 6,2 .5.8... 8.3 8,2 c LOW--3 0,7 1.2 1.8 c SECTION NOl 1 2 5 0 5 6 • 7 8 9 10 WOOD ? 2 . « 19.0 16.9 10.6 12.5 4-l+0_ 2.« _J3,6 5,2 2,6 0,8 SUM WOODS 1.1-4,1 KG c BARK 5.9 5,5 0,6 3,1 3.3 1.9 1.1 0,7 0,3 SUM BARKS 28,8 KG TREE BIOMASSs o ******************************** c. c c TREE SPECIE AGE LENGTH LENGTH NO TO 1 (M) (M) LENGTH TO 6 (M) DBH (M) CROWN DIAM (M) CROWN BUTT LENG LENG (M) (M) BRANCH BRANCH OVER 1 1T01/4 (KG) (KG) CONES (KG) TWIGS (KG) FOLIAGE (KG) TWIG+FOL (KG) e 72 1 290 20,2 23,1 17,0 0,396 6,6 17,3 6,9 56,0 74,2 UUUU 24,0 44,4 68,0 e T0P--1 MID — 2 LOW —3 5,7 11.1 7,2 8,8 21.3 14.3 10,5 32,0 21,5 SECTION NOl 1 2 3 0 5 6 _J 8 9 10 WOOD 101,8 70,2 60,0 56,1 07,0 33,4 22,2 13,1 6,3 1,3 SUM WOODS 411,4 KG BARK 28,3 23,8 18,3 10,2 12,8 8,7 7,6 3,6 1,8 0,5 SUM BARKS 119,5 KG TREE BIOMASSs 7.2.% If • •*»*«***»»****•••»*••*»•**•*••<<**•****•** u -> TREE SPECIE AGE LENGTH LENGTH LENGTH DBH TO 6 CROHN OT AM CROWN BUTT BRANCH BRANCH .LENG. LENG OVER 1 1T01/4 CONES TWIGS FOLIAGE TWIG+FOL (M) (M) CH) CM) CM) CM) CM) CKG) CKG) CKG) CKG) CKG) CKG) 10* I I f n,o • 0,3 S,*. S,4 16,5 l l l l l l l l 6,4 3,1 9,5 TOP--1 MID — 2 1.5 2.1 0,7 1,0 2.2 3,2 L0W*-3 2.8 1.3 «.2 SECTION NOl I 2 3 0 5 6 7 8 9 10 . HOOD 13.3 12,3. 10,7 9,2_ 7,4 5,5. _4 ,6 .3 , <L 1., 7. 0 . 5 SUM_.WOODS .._ 66,6 ..KG. BARK 2,2 2,1 1,7 1,0 1.2 1,1 0,7 0,4 0,4 0,4 SUM BARK* 11,2 KG TREE BIOMASS* KG **«.»•*»*.*#•*«»«»»•.«..«**•**»•**»•««****»******»******•*«**»»**************«»*««*»««*****.•«•«»*•»**««*•«•»••«*««»«•»*«»»*««»«*» TREE SPECIE AGE LENGTH LENGTH NO TO 1 CM) fMl LENGTH DBH TO 6 CM) CM) CROWN DIAM , CM) CROWN BUTT BRANCH BRANCH LENG LENG OVER 1 1T01/4 CM) CM) CKG) CKG) CONES CKG) TWIGS CKG) FOLIAGE CKG) THIG»POL CKG) 105 1 360 17,2 16,5 7,3 0,325 4,0 10,4 . 6,9 17,7 29,8 UUUU 5.4 13,8 19.2 ' TOP--1 MID—2 LOW--3 0,8 1.8 2,6 1.7 5.6 6.5 2.5 7.4 9.3 SECTION NO 1 1 2 3 4 5 6 7 8 9 10 HOOD 52,8 47,6 48,0 43,4 11.2 6,8 5,3 3,9 1,3 0,5 SUM WOOD* 220,6 KG BARK 6,9 6,9 7,3 6,8 «,3 1.9 1.2 0,7 0,4 0,4 SUM BARK* 36,8 KG TREE BIOMASS* • • A * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * V ; : . : : : : : ; SECTION NOI t 2 3 4 5 6 7 8 9 10 WOOD 150,6 130.6__ _M.iL _7.5-.5_ 6-4,5 39,7 26,1 _ i _ . , . _ _ 5.8 1...2. - SUM-WOOD* 601,3. . KG BARK 25,a 33,9 29,5 22.3 15,8 12,4 8,5 3,5 1.6 0,4 SUM BARKs 153,2 KG TREE BIOMASSs TREE SPECIE AGE LENGTH LENGTH LENGTH OBH CROWN CROWN BUTT BRANCH BRANCH CONES MO T.Q_1 IO_6 DIAM LENG LENG.-_ftV.ER_l 1X01741 (M) (M) (M) CM) (M) CM) CM) CKG) CKG) (KG) TWIGS FOLIAGE TWIG*FOL 108 -4_ 365 27,7 26.(1 19.6 0,490 7-+_ 23,2 4.6 122.2- J _ U _ - uuuu (KG) 2".l (KG) «L,.9_ (KG) -X2,_-TOP--1 -MID--2.. L0W--3 4.5 -10,0-9.6 11,0 .20,8... 16,1 15,5 30,7 25,8 ********************************************************************************************************************************** 1 o © o o o o o o 0 TREE SPECIE AGE NO LENGTH LENGTH TO 1 LENGTH TO 6 OBH CROHN DIAM CROHN BUTT BRANCH LENG LENG OVER 1 BRANCH 1T01/4 CONES TWIGS FOLIAGE S TWIG*FOL CHJ CH) CM! - (H)— CM) (Ml CM) CKG) CKG) (KG) CKG) (KG) (KG) 8 J 115 0 17,3 9,9 0,231 5.8 13.5 5,5 4.4 13,7 UUUU 0,0 0,0 15.3 T0P--1 TWIGS AND FOLIAGE MID--2 ARE NOT SEPARATED 3,5 7,8 L0W--3 FOR CEDAR « . l SECTION NOI 1 i 3 4 5 6 7 6 9 10 WOOD 28,9 24,5 23, 1 18,0 15,5 14,0 9,2 5,0 2.2 0,4 SUM WOOD" 140,9 KG BARK 2,5 2.« 2. 5 2.1 1.6 1.3 1.3 0,9 0,6 0,3 SUM BARK" 15,6 KG ' TREE BIOMASS" —OBH" CM) ""BRANCH" 1T01/4 CKG) ' T K t t a r t e I t » e t LENGTH NO ( H ) LENGTH LENGTH TO 1 TO 6 CM) CM) CKUWfT CKUWN B U T T 8 K A N t n DIAM LENG LENG OVER 1 CM) CM) CM) CKG) CONES TWIGS FOLIAGE (KG) (KG) (KG) TwlG*FOL (KG) 63 3 22) 2T, 2 25,8 20,0 0,401 8.6 15,4 11,8 48,9 23.7 UUUU 0,0 0,0 42,9 - X0P*-1" "TWIGS AND-FOLIAGE MID—-2 ARE NOT SEPARATED L0W--3 FOR CEDAR 5,4 22,3 15.1 SECTION NOt 1 2 3 4 5 6 7 8 9 10 NOOO 114,0 95,1 TT, 4 66,9 59,8 55,9 37,7 16,4 9,3 2,6 SUM WOOD" 539,0 KG BARK 5,6 « . l •>. 5 6,3 4,8 1.5 4,5 3,9 1.0 0,8 SUM BARKS 42,4 KG TREE BIOMASS" o o o o o o o o TREE SPECIE AGE LENGTH LENGTH LENGTH DBH CROWN CROWN BUTT BRANCH BRANCH CONES NO TO 1 TO 6 DIAM LENG LENG OVER I 1T01/4 (H) (M J CM ) P fT (Ml" 76 195 14,8 - H)— 13.3 5,3 0,180 6,1 - (rO— 9.1 TMT~ 5.7 (KG) 1.7 (KG) 11.1 (KG) UUUU TWIGS "(KG) 0,0 FOLIAGE "(KG) 0,0 TWIG»FOL (KG) 17,5 C TOP--1 TWIGS AND FOLIAGE MID--2 ARE NOT SEPARATED -C0W--3-"-F0R~"CEDAR 2.4 5,2 "9,-9-SECTION NOl 1 WOOD 16,9 2 11,9 3 9,9 4 6.2 5 7.3 6 5.7 7 3,9 B 2.2 9 0.9 10 0,5 SUM WOODs 67,6 KG BARK 0,6 1.1 0.5 0,8 0,6 0,6 0,6 0,6 0.3 0,0 SUM BARKs 5.7 KG TREE BIOMASSs f'P^/S KG a************************************************** ************************** ******* "TREE—SPECIE AGE—LENGTH LENGTH—LENGTH—OBH—tROITN—CROTN BUTT—BR1TOCH—BR7SNCH—CONES TWIGS FOLIAGE TWTG+FOtT NO TO 1 TO 6 DIAM LENG LENG OVER 1 1T01/4 (M) (M) (M) (M) (M) (M) (M) (KG) (KG) (KG) (KG) (KG) (KG) 102 185 25,5 24,3 19,7 0,602 9,1 15,0 10,5 126,2 52,6 UUUU 0,0 0,0 109,6 T O P " - ! TWTGS-"AND-POtTAGE-MID—-2 ARE NOT SEPARATED LOW--3 FOR CEDAR -16-rr-49,2 43,8 SECTION NO I I 8 10 WOOD 245,7 167,1 140,5 111,5 99,5 75,5 52,5 31,2 11,5 2,7 SUM WOOD" BARK 12.7 I3T3 12,5 13,2 9y9 Ty4 4.8 2.0 2,5 0.7 SUM BARKs 937,8 KG 79,0 KB TREE BIOMASSs / 30 ********************************************************************************************************************************** A P P E N D I X 6 NUTRIENT CONCENTRATION AND N U T R I E N T D I S T R I B U T I O N W I T H I N 20 SAMPLED T R E E S ON THE SQUAMISH SAMPLE PLOT S P E C I E S 1 - MOUNTAIN HEMLOCK S P E C I E S 2 - P A C I F I C S I L V E R F I R TREE NO. 22 SPECIES NO. 2 LENGTH M l 37.3 OBH (Ml 0.780 CROWN DIA(M) 7.5 CROWN BUTT L.(M) L.(M) 25.9 11.* • ••PERCENT NUTRIENT CONTENT BY SAMPLE SPEC IF ICAT I 0 N * » * * * ********************************************************<•********# * * * » * < • * • » W000**SECTION— * * BARK**S ECT I ON *-I 3 5 7 J) 10 1 3 5 7 9 10 NITROGEN -j> HO S PHORU S_-_ "C'ATCIUM -MAGNESIUM -POTASSIUM -0.053 _0, OA 7_ 0.051 0.009 0. 03* 0.0*5 0.005_ 0. 07* 0.011 0.058 0.0*7 0.003 0. 056 0.005 0.063 0.008 0. 069 0. 010 NITROGEN -PHOSPHORUS-0.277 Oo 037 0. 063 0.012 0.05 9 0. 082 0. 017 0. 068 0.071 0.016 0.10* 0. 067 0. 016 0. 078 CALCIUM MAGNESIUM POTASSIUM 0.335 0. 019 Oc 090 0. 207 0.0 3J_ 0.126 0.028 0. 102 0.208 0.032_ ' O.*6"0 0.027 0.093 o.i e* "b."350 0.031 0.117 0.203 0, 03 *_ ' 0,'*26 0.030 0.096 0.297 0.0*7 "6". 6 88" 0.040 0.172 BRANCH BRANCH -TWIGS- -FOL I AGE- CONES >1 1-1/* (TOPI ( MIOI NITROGEN - 0. 169 0.227 0.000 0.000 (LOW) 0.000 (TOP) 0. OOP (MID) 0.000 (LOW) 0.000 *—-TWJGS + FOL I AGE ( TOPI " " ( MI D)~ flOW) 2.133 0.789 0 . 6 8 8 0.76* PHOSPHORUS-CALCIUM MAGNES IUM -0. 02* 0. 311 0.02* 0. 033 0. 356 0.031 0.000 0. 00 0 0.000 0.000 0.000 0. 000 POTASSIUM 0. 107 0.11* 0.000 0.000 0.000 0.000 0.000 0.000 0.000 0. 000 0. 00 0 0. 000 0. 000 0. 000 0.000 0.000 0„ 000 0. 000 0.000 0.000 0. 368 0. 067 J?„10JL 1.260 0.11* 0.*13 0.060 0.311 0. 102 0.)07 0.612 0.563 0.070 0.073_ 0.306 0. 261 ***NJTR!ENT AMOUNTS IN DECAGRAMS BY SAMPLE SPEC IFI CAT I ON*****************••*•***••«****•**•*»****»****•***•»***»*•**•***»***• »•*** -WO00«*SECTICN- -BARK**SECT!ON-1C2 3 £ * 5 £6 7£8 10 TOTAL l t2 3£* NITROGEN - * * .97 29.12 21.69 17.26 1.99 0.*2 11,5. *5 NITROGEN - 29.6* 17.26 PHOSPHORUS- 9.71 3.05 1.66 1.81 0.25 CALCIUM - 51.96 *3.15 30.3* 22.13 2.25 MAGNESIUM - 8. 17 7. 1* 6. 1 3 *. 83 0.50 0.06 16.55 0.*1 150.25 0.09 26.85 PHOSPHORUS-CALC1UM MAGNESIUM -*.17 2.62 *0i*6 33.51 2. 83 2.26 5t6 16.61 '2.7* 35.20 2.*6 7C8 18. 29 " '3V'1'3 36.71 2.91 9 1 0 3, 69 0. 56 "0", 61 " " OYOS 7,76 1.31 0. 55 0.08 TOTAL 8 6 . 06_ "" 13."3 6 1 5 * . 0 6 11 . 09 POTASSIUM - 59.03 36.89 26.5* 2*.80 3.27 _BRANCH_ BRANCH " >i" ~ "i-i>* (TOP) ---TWIGS (MID) "(LOW)' 0.*8 151.00 * * TOTAL (TOPI POTASSIUM --FOLIAGE-(MlUI ' (lOW) 1 1. 77 8.11 8.90 10.06 1,7* 0.33 * CONES * TWIGS»FOLIAGE — "TOTAL " "(TOP) (MIDI ~ (LOW) TOTAL NITROGEN - 69. 5* 19. 57 0. 00 0.00 0. 00 0.00 0. 00 0. 00 0. 00 0.00 0. 00 PHOSPHORUS- 9. 77 2. 89 0. 00 0.00 . 0. 00 0.00 0. 00 0. 00 0. 00 0.00 0.00 CALCIUM 128. 1* 3 0. 7* 0. 00 0.00 0. 00 0.00 0. 00 0. 00 0. 00 0. 00 0. 00 MAGNESIUM - 9. 96 2. 66 0. 00 0. 00 0. 00 0.00 0. 00 0. 00 0. 00 0. 00 0.00 POTASSIUM - * 3 . 96 9. 85" "0. bo ~ d^ dd " " b." 00 0.00 " o. "00" d. 00 0. 00 " "r b.oo" " "0.00 **.65 39.95 63. 39 1*7.99 *0. 92 TP. EE TOTAL" 428,61 6.*3 23.37 3.38 17.61 5.95" 35„5* * t 05 15. 16 8. H<5~ 2TT27" *6,77 105.68 6.06 1 3 . 4 9 25.36 " 58.13 6i.7T~ 569.7B 6 4 , OS 303. 87" r TREE SPECIES LENGTH OBH CROWN CROWN BUTT NO. NO. (Hi (Hi DIAIMI L.(M) L.(M) 41 2 21.9 0.356 6.2 19.2 2 .7 NITROGEN - 0. 051 0. 350 0.05 3 0.057 0.067 0. 090 NITROGEN - 0.276 0.273 0. 307 0. 325 Oc 431 0. 587 PHCSPHOBUS- 0.005 o.ooa 0. 01 0 0. 003 0.021 0.023 PHOSPHORUS- 0. 040 0.056 0.058 0.069 Oo 095 0.134 CAICIUM 0.06 9 0.056 0.051 0. 048 0.052 0. 037 CALCIUM 0.4 54 0.519 0.472 " 0.3"3 6 0"o"318" 0.327 MAGNESIUM - 0.015 0. 010 0.01 0 0.011 0.011 0.013 MAGNESIUM - 0. 018 0.042 0. 043 0. 042 0. 044 . 0.060 POTASSIUM - 0.06 7 0.060 0. 052 0. 060 0. 061 0. 078 POTASSIUM - 0.123 0.167 0.198 0.227 0. 312 0.395 • ••PERCENT NUTRIENT CONTENT BY SAMPLE SPECIF 1 * . W00D^*SECTION-10 -BARK**SECT ION-5 7 10 BRANCH BRANCH • TWIGS * _ _* — " " f PL I AGE — — — r _ * CONES * —TWI GS*FOL I AGE — - * >1 1-1/4 (TOP) (MID) (LOW) (TOP) (MID) (LOW) (TOP) (MID) (LOW) NITROGEN - 0.143 0.214 0.000 0.000 0.000 0.000 0.000 0.000 0.990 0.836 0.735 0. 712 PHOSPHORUS- 0.021. 0.031 0.000 0.000 0.000 0.000 0.000 0.000 0.181 0.098 0.087 0.078 CALCIUM - 0.254 0.239 0.000 0.000 0.000 0. 000 0. 000 0.000 0. 025 0.301 0.346 0.380 MAGNESIUM - 0. 02 5 0. 030 0. 000 0. OOP 0.000 0.000 0.000 0.000 O^Obl 0.073 0. 069 0. 053 POTASSIUM - 0.073 0.114 0.000~~ 0.000 0.000 0. 000 0, 000 0.000 0.486 0.324 0.297 6.267 •••NUTRIENT AMOUNTS IN DECAGRAMS 8Y SAMPLE SPECIF] -WOOO^SECTION- -BARK**SECTION-TOTAL _NI TROGEN -PHOSPHORUS-CALCIUM HAGNESIUH -1C2 6.05 3C4 4. 32 5 66 2.96 7£8 1.35 0.25 10 0.10 TOTAL 15. 55 NITROGEN 1C2 4.39 3C4 4.53 5G6 2.53 7£8 10 0.73 0.32 7.54 5.05 1.51 0.93 0 .48 2.63 0.57 0.32 0.08 1.09 0.19 0.25 0.04 0.03 0.04 0.01 2.50 16. 60 3. 32 PHOSPHORUS-CALCIUM MAGNESIUM -0.76 0.88 7.78 7.73 0.48 0.66 0.51 3.23 0.34 2.2 3 "0.4 8" 0.22 '0."0'5~ 1.93 0. 16 0. 25 0, 02 1. 59 0. 16 0.12 14.01 "0703 2771 ' 0.07 20. 89 0.01 1.77 POTASSIUM - 7.63 5.25 3.00 1.31 0.22 0.09 17.55 POTASSIUM 2. 32 2.85 1.70 0. 08 BRANCH BRANCH •- -TWIGS->1 1-1/4 (TOP) (MID) (LOW) TOTAL (TOP) ^ F 0 LI AG E - - * CON ES_ (Mib) (LOW] TOTAL TWIGS*FOLI AGE--( TOP) "(MID) "(LOW)" NITROGEN - 4.93 6.71 0.00 0.00 0. 00 0.00 0.00 . 0.00 0. 00 0.00 0.00 8.03 21.53 13.39 2.56 n w 10.14 7.15 2.01 1.00 "8.70 ~ 5 . 0 3 "TOTAL" 42.95 8. 69 TP EE "TOTAL" e4.14 PHOSPHORUS-CALCIUM fAGNES HIM -0.70 0.98 8. 42 7. 49 0.82 0.94 0. 00 0. 00 0. 00 0.00 0.00 0.00 0.00 0.00 0.00 POTASSIUM - 2.59 3.59 0.00 0.00 0. 00 0.00 0.00 0.00 0.00 0.00 0.00 "0.00 oVoo 0. 00 0. 00 0.00 0.00 0.00 0^0 0_ 0. 00 0. 00 ^JTOT5orrnr 0.00 0.00 0.00 0. 00_ "d.'O'O 0.00 0.94 2.89 0.70 "3.11" 4.96 20. 18 3.71 16.84" I I.s; 73. 59 10.57 -49726"" TREE SPECIES LENGTH 08H CROWN NO. NO. (HI (Ml OIAIMI 45 1 35.2 0.754 0.0 CROWN BUTT L.(M) L.(H) 29.2 6.0 •••PERCENT NUTRIENT CONTENT BY SAMPLE SPEC IF ICATION************************************************************** -WOOD**SECTION-- -BARK**SECTION-1 3 5 7 9 10 1 3 5 7 <J 10 NITROGEN - 0. 057 0. 052 0.053 0. 051 0. 059 0. 067 NITROGEN - 0.259 0.171 0.208 0. 198 0. 247 0. 280 PHOSPHORUS- 0. 017 0. 007 0.009 0.012 0. 020 0.021 PHOSPHORUS- 0. 061 0. 054 0. 052 0. 049 0. 055 0.062 CALCIUM 0. 053 0. 057 0. 070 0. 078 0. 064 0. 081 CALCIUM 0.693 0.615 0.854 0. 374 0. 435 0. 3 59 MAGNESIUM - 3. 012 0. 012 0.016 0. 014 0. 015 0. 016 MAGNESIUM - 0. 016 0.016 0.017 0. 017 0. 020 C.024 . POTASSIUM - 0, 094 0. 057 0.050 0. 054 0. 080 0. 062 POTASSIUM - 0. 084 0.094 0.094 0. 101 0. 109 0.161 BRANCH BRANCH -TWIGS-- -FOL IAGE- CONES * TWIGS+ FOLIAGE---* >1 1- 1/4 (TOP) (M10) (LOW 1 (TOP) (MID) (LOW) ( TOP) (MID) ( LOW) MTROGEN - 0. 000 0. 000 0.000 0. 000 0. 000 0. 000 0. 000 0.000 0.000 0.000 0.000 0. 000 PHOSPHORUS- 0. 000 0. 000 0.000 0.000 0. 000 0. 00 0 0. 000 0.000 0. 000 0. 000 0. 000 0. 000 CALCIUM - 0. 000 0. 000 0. 000 0.000 0. 000 0. 00 0 0. 000 0.000 0.000 0.000 0.000 0. 000 MAGNESIUM - 0. 000 0. 000 0.000 0. 000 0. 000 0. 000 0.000 0.000 0.000 0.000 0.000 0. 000 POTASSIUM - 0. 00 J 0. 000 0.000 0.000 0. 000 0.000 0. 000 0. 000 0.000 0. 000 0. 000 0. 000 •••NUTRIENT AMOUNTS IN DECAGRAMS BY SAMPLE SPECIF1 » . WOOD*»SECTION -BARK**SECTION-1C2 3C4 5C6 7E8 10 TOTAL NITROGEN PHOSPHORUS-CALCIUM MAGNESIUM -54.03 34. 20 20.2 5 11 ^ 17 11.86 5.33 4.16 3.30 54.42 41.30 23.83 14.43 11.35 9. 11 5. 39 3.02 2.44 0.58 122.72 NITROGEN 1&2 40. 50 0.83 2.65 0.63 0.18 0.70 0.13 25.66 142.34 30.62 PHOSPHORUS-CALCIUM MAGNESIUM -3 £4 28^16 ,8 6" 10.84 7. 123.16 109.06 3.00 2.43 5 £6 J 4 . 7 6 "3.65 4 4 . 5 6 1.23 7C8 10.48 2";v4 19.04 0.87 9 10 1.92 0.48 "0.43 " o;n 3.39 0.61 0.16 0.04 TOTAL 96. 31 " "2'5. 32"" 299.83 7. 72 POTASSIUM - 74.56 34.86 20.38 13.65 3.33 0.54 147. 32 POTASSIUM- 16.82 13.98 7.09 4 .96 "0785 0.27 _BRANCH_8RANCH_*-- - -"~>l" ""1-1/4 (TOP I ---TWIGS (MID)" (LOW) • *. TOTAL ( TOP) — FOL I AGE-(MIDI (LOW) TOTAL CONES TWIGS«-FOLIAGE--(TOP)" (MID) " (LOW) TOTAL 43. 98 TREE TOTAL" NITROGEN - 0. 00 0.00 0. 00 0. 00 0. 00 0. 00 0. 00 0. 00 0. 00 0.00 0. 00 0.00 0. 00 0. 00 0. 00 219.03 PHOSPHORUS- 0. OJ 0.00 0. 00 0. 00 0. 00 0. 00 0. 00 0. 00 0. 00 0.00 0. 00 0.00 0. 00 0. 00 0.00 50.98 CALCIUM 0. 00 0.00 0. 00 0. 00 0. 00 0. 00 0. 00 0. 00 0. 00 0. 00 0. 00 0.00 Oo 00 0. 00 0.00 442.16 MAGNESIUM - 0. 00 0. 00 0. 00 0. 00 0. 00 0. 00 0. 00 0. 00 0. 00 0.00 0. 00 0.00 0. 00 0. 00 0.00 38.35 POTASSIUM - 0. do o.oo" 6. 00 0. 00" """"0. ob" 0. 00 "0. 00"" 0. 00 ' 0. do"""" ~ "b". oo'" "b. 00" " 0.00 0. 00 0. 00 " 0.00 ' " 191.29 TREE NO. 55 SPECIES NO. 1 LENGTH ( H I 34. 4 OBH (Ml 0.631 CROWN OIA( Ml 0.0 CROWN BUTT L.(M) L.(M) 28. 2 6.2 •••PERCENT NUTRIENT CONTENT BY SAMPLE S P E C I F I C A T I 0 N » * * * * ^ * * * * ^ * * * » * * * * * * * * * * * * * * * * * * * * • * • * * * • • • * • • • * • * • « • • • • * • • • * • • • • * • * • * • * • « » * » * • WCOD+*SECTION * * BARK**SEC Tl ON * 1 . 3 5 7 9 10 1 3 5 7 9 10 NITROGEN -PHOSPHORUS-CALCIUM MAGNESIUM -POTASSIUM -0. 055 .0 -.021_ 6. 057 0. 016 0.073 0. 051 __.020_ 0.1)76 0. 022 0.083 0. 04 6 0. 01 2__ 6.079 0. 022 0.048 0. 049 _0. 015_ 0. 079 0. 021 0. 060 0.054 _0.014_ 0.0 51 0.013 0.066 0. 06 0 _0._013_ 0. 05 0 0. O i l 0. 057 NITROGEN -PHOSPHORUS-CALCIUM MAGNESIUM -POTASSIUM -0.285 _0i08_3_ 0. 2 26 0.017 0.099 0. 162 0.063 0.286 0.016 0.073 0. 177 0.070_ 0. 386 0.018 0.106 0. 188 0.064 "0. 1 92" 0.020 0.101 0. 236 0. 308 0. 075 0.081 0.292 "0.313" 0.026 0. 037 0.120 0. 166 NITROGEN BRANCH >i 0.000 JBRAil_CH_ 1-1/4 0. 000 ( TOP I -TWI_ S-( MIDI ( L O W ) (TOP I -FOLI AGE-(MI D I ( L O W I 0.000 0.000 0.000 0.000 0.000 0.000 CONES 0.000 * TWIGS* FOL I AGE » (TOPI (MID) " (LOW) 0.000 0.000 0.000 PHOSPHORUS-CALCIUM MAGNESIUM -0. 000 0.000 0. 000 0.000 0.000 0.000 POTASSIUM - 0.000 0.000 0.000 0.000 0. 000 o."doo 0.000 0.000 0.000 0.000 0.000 0.000 0. 000 0. 000 0. 000 0. 000 0. 000 0.000 0. 000 0.000 0.000 0.000 0.000 0. 000 0. 000 0.000 0.000 0.000 0.000_ "0". 000 0.000 0.000 0.000 0.000 0.000 0.000 0.000 0.000 0.000 0.000 0^000 o". bob" • ••NUTRIENT AMOUNTS IN DECAGRAMS BY SAMPLE SPEC I F I C A T I O N * * * * * * * * * * ^ * * * * * * * * * ^ * * * * * * * * * • * • • * • * • * * * « * * * * • * • * * » * * * « • • • « * * * * • * * * * » * » * • -WOOO^«SECTION- -BARK**SECTION-1C2 3£4 5£6 7£8 9 10 TOTAL U 2 3C4 5£6 7C8 9 10 TOTAL NITROGEN - 41.02 24. 23 15.95 7.70 1.21 0.31 90.42 NITROGEN - 40. 74 23.22 13.13 8.34 1.11 0. 34 86. 88 PHOSPHORUS- 15.74 7.97 4.58 2.17 0.32 0.07 30. 85 PHOSPHORUS- 13.32 9.11 4.84 — 2 7 7 3 "07'3"S~ "0.09" 30.44 CALCIUM 51.21 38. 38 26.43 9.65 1.13 0.26 127.06 CALCIUM 46.60 46.01 20.77 9.51 1.37 0. 34 124.60 MAGNESIUM - 14.54 10.92 7.11 2.52 0.30 0.06 35.44 MAGNESIUM - 3. 00 2.30 1.34 0.89 0.12 0.04 7.69 POTASSIUM - 62.16 32.59 18.14 9.3 5 1.47 0.30 124.00 POTASSIUM - 15.67 12.28 7.43 4.33 0. 56 0. 18 40.45 BRANCH BRANCH •- -TWIGS :* »->1 1-1/4 (TOPI ( M I D I (LOWI TOTAL NITROGEN - 0. 00 0. 00 0.00 0. 00 0. 00 0. 00 0. 00 PHOSPHORUS- 0. 00 0. 00 0.00 0. 00 0. 00 0. 00 0. 00 CALCIUM - 0. 00 0. 00 0. 00 0. 00 0. 00 0. 00 0. 00 MAGNESIUM - 0. 00 0. 00 0.00 0. 00 0. 00 0. 00 0. 00 POTASSIUM - 0. 00 0. 00 0. 00 "o"; 00 0. 00 0. 00 0. bo FQ L IAGE----T T C J P V ( M I D I " (LOW! 0. 00 TOTAL 0.00 CONES * TW I GS + FOL IAGE--< T O P ) ( M I D I " " ( L O W ' l ' 0.00 0.00 0.00 0.00 "TOTAL" 0.00 TREE "TOTAL" 177.30 0. 00 0.00 0. 00 0.00 0.00 0.00 0. 00 0.00 0.00 0.00 0.00 b.oo" 0. 00 0.00 0. 00 0.00 0.00 0.00 "0.00" 0.013 5701) 0.00 0.00 0.00 _ 0.00 "0. 00 "0. 00 "DT30" 0. 00 0.00 "0.00 61.29 251.66 43. 13 164.45"' TREE NO. 64 SPECIES NO. 1 LENGTH 1 M I 21.3 OBH (M) 0.320 CROWN DIA(Ml 0.0 CROWN L.(M) 12.4 BUTT L. (Ml 9.0 »*»OERCENT NUTRIENT CONTENT BY SAMPLE SPECIF ICATION******************************************************** * WOOO**SECTION * * BARK**S ECT ION « 1 3_ 5 7 9 10 1 3 5 7 9 10 NITROGEN -PHOSPHCRUS-CALCIUM MAGNESIUM POTASS!UM 0.062 0.017 0.053 0.027 0.061 0.095 0.016 0.025 0.066 0.081 0.051 0.01 7 0. 06 9 0.018 0.072 0. 055 0±0_ZQ_ 6. 053 0.017 0. 067 0.066 0.014 0. 075 0. 012 0.047 0. 042 0.013 0.013 0.055 0.065 NITROGEN -PHOSPHORUS-CALCIUM MAGNESIUM -POTASSIUM -0.251 0. 059 0.178 0.075 0.704 0.345 0.018 0.016 0.072 0.075 0. 198 _0._04J _ 0.308 0.018 0. 084 0.208 0^08 0__ 6.216 0.018 0.093 0.248 0.095 0.339 0.103 0. 194 0.226 0.030 0.045 0. 106 0.159 BRANCH BRANCH >1 1-1/4 -TWIGS- -FOLIAGE-(TOP) (MID) ( LOW) (TOP) (MID) (LOWl CONES * TWIGS + FOlI AGE * ( TOP) (M ID ) ( LOW ) NITROGEN 0.030 0.000 0.000 0.000 0.000 0. 000 0.000 0.000 0.000 0.000 0.000 0.000 PHOSPHORUS-CALCl 'JM MAGNESIUM -POTASSIUM -0. 000 0.003 0.03J 0.003 0. 000 0.000 0.003 3.000 0.03 0 0.300 0.00 0 0. 000 0.000 0. 000 0.000 0.000 0.000 0. 000 0. 000 0. 000 0. 000 0. 000 0. 000 0.000 0.000 0. 000 0.000 0.000 0.000 0.000 0.000 0.000 0.000 0.000 0. 000 0. 000 0. 000 0.000 0.000 0.000 0.000 0.000 0. 000 0.000 0.000 0.000 0.000 0.000 •••NUTRIENT AMOUNTS IN DECAGRAMS BY SAMPLE SPEC IF ICAT ION**** * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * -WOOD**SECTI0N- -BARK**SECTION-NITROGEN 162 6.92 3 64 4. 09 PHOSPHORUS-CALCIUM MAGNESIUM -2.67 3.76 2.46 1.73 6.04 1.69 566 2.76 0.95 3.13 0. 92 7£8 1.75 0.39 10 0.13 TOTAL 16.03 NITROGEN -162 4.92 364 3.45 0.49 1.45 0.43 0.08 0.28 0.07 0.02 0.08 0.02 5. 95 19.77 5.60 PHOSPHORUS-CALCIUM MAGNESIUM -1.53 12. 01 0.40 1.06 5.97 0.32 5 £6 2. 25 "6.67 2.91 0.20 7£8 U 09 0.42" 0.98 0.11 10 TOTAL 0. 15_ "0. 06 0. 12 0. 02 11. 96 ,77" 0. 10 "OTOT 3. 0.07 22.05 0.01 1.06 POTASSIUM 8.82 6.02 3.62 1.76 0.32 0.12 20.65 POTASSIUM 1.68 1.46 0.98 0.48 0.06 0.05 3RANCH BRANCH _*----------TWIGS : >1 1-1/4 1 TOP) (MID I (LOW) * * - - - -FOL I AGEj^----f b TA L " ( TUP) (MIDI ( L 0 W) » "TOTAL CONES * TWIGS+FOLIAGE--" " " " " ( T O P ) , .' (MID) " (LOW) T3TAL 4.71 TREE TOTAL" NITROGEN 3. 30 0.00 0.00 0.00 ' 0. 00 0.00 0.00 0.00 0.00 0. 00 0.00 0.00 0. 00 0. 30 0. 00 28.00 PHOSPHORUS" 0.33 6.00 6.00 0.00 3.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 9.73 CALCIUM - 0.03 0.30 0.30 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.60 0.00 0.00 41.63 _MAGNF SI UM _- 3. 33 0.00 0. 00 0.00 0. 00 _ 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 6.65 POTASSIUM -~ 3.66 0.00" 0.00 "6 . 00 Of. 00' "6 .00 "6.00 0 . 0 0 0 . 0 0 6.00 ' 0 . 0 0 0 . 0 0 " 0.00 0.00 0.00 '25.35 r TREE NO. 63 SPECIES NO. 1 LENGTH ( M) 33.0 DBH ( .11 0. 711 CROWN DlAIM) 7.9 CROWN BUTT L.IMI L.(M) 24.5 13.5 • •••PERCENT NUTRIENT CONTENT BY SAMPLE SPECIFICATI ON********************************* ********•***•*•***•••*•••••»••***••»*•**•**** * • WOOD**SECT ION * * BARK**SECTI ON * 1 3. 5 7 9 10 1 3 5 7 9_ 10 NITROGEN -JLHO.SiHOjyj.Sj; CALCIUM MAGNESIUM POTASSIUM 3. 053 _0._Q32 0.073 0. 026 0.070 0.054 _0,_P13_ 6.067 0. 320 0.063 0.04 8 0. 012 0.08 8 0.02 9 0. 056 0.055 _P„P16_ 0.078 0.024 0. 071 0.068 J>.026_ 6.090 0.029 0.076 0.075 __0^029_ 0. 076 0. 025 0. 08 5 NITROGEN -PHOSPHORUS-CALCIUM MAGNESIUM -POTASSIUM -0. 190 0.070 0.307 0.019 0. 075 0.137 _0.061 6.229"" 0.017 0.084 0. 148 0.053_ " 6. i"96 0.016 0.087 0.155 3.047 '~6".2 45"~ 0.01 7 0.092 0.2 26 0.057 0. 185 0, 047 "0.177 0.282 0.019 0.022 0. 100 0.110 _B?ANCH_ >1 BRANCH 1-1/4 • TWIGS-ITOPT (MID) ( LOW) (TOP) -FOLIAGE-(MID) (LOW) CONES * TWIGS*FOLI AGE * "(TOP) (MID) (LOW I NITROGEN 0. 146 0. 202 0.475 0.449 0.243 0. 840 1.404 0.805 0.000 TQ>^ 57 0.927 0.1>^* PHDSPHORUS-CALCIUM MAGNESIUM -POTASSIUM 0. 02 7 0.110 0. 022 0. 046 0.125 0.032 0.072 0.178 0.045 0. 067 0. 170 0.050 0.0 34 0.125 0.045 0. 108 0. 313 0. 100 0. 118 0.282 0.094 0. 103 0.279 0.089 0.000 0. 000 0.000 0.074 0.091 0.155 0.156 0.169 0.235 0.250 0.273 0.000 • ••NUTRIENT AMOUNTS IN DECAGRAMS BY SAMPLE s PEC IF ICAT I ON**************************** *************************************"*** ***** * • WOOD** SECT ION * * BARK** SECT I ON 1C2 3C4 5 £6 7£8 10 TOTAL 1(12 3£4 5£6 7&B TO" TOTAL NITROGEN - 41.52 29.24 21.16 13.15 2.70 0.70 108.47 NlTROGEN PHOSPHORUS-CALCIUM MAGNESIUM -16. 86 53.59 16.92 7. 30 44.66 13. 13 5.34 34.45 11.00 4.54 17.94 5.66 1.03 3.54 1.13 0.27 0.71 0.23 35. 84 154.90 48. 88 PHOSPHORUS-CALCIUM MAGNESIUM -23.02 25.44 14.69 9.26 37.79 2.53 5.11 T7T3 1.20 76.25 o;"30 27rer 1.49 112.75 0.12 8.48 TJ7F8 42. 57 POTASSIUM 49.06 34.09 26.23 1 5.67 2.99 0.79 128. 83 POTASSIUM - 11.27 15.36 BT71 NITROGEN BRANCH 9RANCH *->1 1-1/4 31.79 6.32 (TOPI 6. 79 •—TWIGS (MIDI (LOW"! 5.30 9. 24 # *. "TO TAT -FOLIAGE- CONES (TOP) (MID) (COW) TOTAL 21.34 13.94 21.77 41.78 1.80 5.20 1. 66 "3^9"0" 77.49 8. 97 24.05 7.72 21.92 TWIGS* FOLIAGE--"'( TOP )""""< MI Or" (LOWI 0.00 0.00 0.00 0.00 o. Ob" 20.73 2.83 7.74 2.31 6.12 27.07 2.61 6.37 2.05 '"""5.71" "TOTAL"" 98.82 51.03 6.63 19.23 33.34 6.33 ' 10.69 20.61 " 32.44 TREE "TOTAL" 321.64 85.03 328.77 73.79 "2 22." 69" PHOSPHORUS- 5.84 CALCIUM - 23.86 MAGNFSIUM - 4.73 POTASSIUM - 16. 02 1. 43 3. 92 W 01 2.8 3" 1. 03 2. 54 0.65 2.22 0.79 2. 01 0^59 1". 85 " 1.28 4. 75 1. 73 6.46 3.10 9.29 2.97 16152' 1.82 4.37 1.45 3.87 5.35 14.48 4. 60 14.15 TREE NO. 74 SPECIES NO. 1 LENGTH (Ml 34.2 CBH (Ml 0.762 CROWN DIA(M) 9.4 CROWN BUTT L.IM) L.(M) 25.2 9.0 • ••PERCENT NUTRIENT CONTENT BY SAMPLE SPEC IF I CAT I ON*** * • * * * * * * • * * • * • • • • * • • • • * * • • * • • • » * • • • • • « » • • • * • • « • • * » " » *M~j+^*&Vi"rr*«^i~M#**** • WOOD^SECTION * * BARK**S ECT I ON » 1 3 5 7 9 10 1 3 5 7 9 10 NITROGEN -PHOSPHnRUS-CALCIUM MAGNESIUM -POTASSIUM -0.056 J? • 0.064 0.015 0. 084 0.045 J?.Q13_ 0. 072 0.018 0.052 0.051 0.014 0. 044 0.021 0.056 0.014 0. 080 0.014 NITROGEN -PHOSPHORUS-0. 076 0.018 0.052 0. 081 0.020 0. 060 0.063 0.012 0.054 0. 052 0. 010 0.062 CALCIUM MAGNESIUM -POTASSIUM -0.218 _0._083_ 0.308" 0.017 0. 092 0.136 0.060 0.344 0.013 0.087 0. 144 0. 054_ " 0.243 0.014 0.115 0.147 0.046_ "0.226 0.014 0. 085 0.229 0.307 0. 06b 3.0 79 0.261 " 0~. 2 73" 0.019 0.028 0. 143 0.181 NITROGEN BRANCH BRANCH >1 0. 114 1-1/4 0.156 (TOP I 0.477 -TWIGS- -FOL I AGE * (MID) 0. 390 (LOWI 0. 3 72 (TOP) 0. 683 (MIDI 0. 866 (LOWI 0.819 CONES 0.000 ? r ~ I _ I G J . i F O l - _ *_E - -"(TOPI (MIDI" ( LOWI 0.62S O.^&^y PHOSPHORUS-CALCIUM MAGNESIUM -POTASSIUM -0. 025 0. 162 0.013 0. 069 0.029 0.184 0.023 0.091 0.067 0.191 0.042 0.156 0. 059 0. 156 0. 038 0.061 0.163 0.043 0. 112 0. 394 0. 077 0. 121 0.335 0. 074 0. 114 0.335 0. 067 0.176 0.192 0.281 0. 332 0.311 0.000 0.37 0.000 0.293 0.000 0 0. 000 440 0.249 055 • ••NUTRIENT AMOUNTS IN DECAGRAMS BY SAMPLE SPECIF I CAT I ON******** * * * * * * * * * * * * * * * • » * * * # * * « • * * • « * * • • * • • * • • • • * • * * ••••••^•••* »•••"•"* •»"*"•" -W00D**SECT10N- -BARK**SECTION-1£2 3£4 5(16 NITROGEN 40.90 26.50 17.44 7£8 8.90 PHOSPHORUS-CALCIUM MAGNESIUM -14.60 54.87 12.97 7.53 40.66 9. 30 6.37 28.63 6. 96 3.12 12.82 2. 85 1.26 0.33 1.43 0.28 10 0.29 0.05 0.19 0.04 TOTAL 95.29 32. 00 138.61 32.89 NITROGEN PHOSPHORUS-CALCIUM MAGNESIUM -1E2 15.01 68. 69 3.13 3 £4 37^31 26.42_ 1077 3 55.47 2.56 5 £6 15. 18 5.22 24.50 1.47 7£8 J2 .06 "3.59 15.64 1.07 1. 2 6 "0'. 36 1,44 0. 11 10 TOTAL 0. 37 92. 60 O.'l'O 35700" 0.33 166. 07 0.03 8.33 POTASSIUM 55.25 28.77 20.60 10.18 1.23 0.22 116.25 POTASSIUM 18. 79 19.00 10.40 7770 0"77~B~ 0771 56.49 BRANCH BRANCH •- -TWIGS->1 1-1/4 (TOPI (MIDI (LOWI NITROGEN - 32. 99 8.41 15. 70 6. 97 7. 18 PHOSPHORUS- 7. 25 1.56 2.19 1. 06 1. 19 CALCIUM 46. 83 9.91 6.27 2. 80 3. 15 MAGNESIUM - 5. 13 1. 26 1. 37 0. 69 0. 82 POTASSIUM - 19.99 4.39" "5. 13 3. 16 "3. 70" • *. TOTAL (TOP) — F O L I A G E — - ; (MIDI" "(LOW) CONES 29.85 21.10 20.53 29.39 TOTAL 71. 02 TWIGS+.FOLIAGE--"(TOPI" (M10 I (LOW I TOTAL" 0.00 36.80 27. 50 36. 57 100.88 TREE TOTAL" 330. 17 4.43 12.22 2.88 11.98" 3.45 12. 19 2.38 "8.67 2.87 7. 94 1.74 "7.86" 4.10 12.04 2.42 11.18 10.42 32.16 6. 54 '27.71" 0. 00 0. 00 0.00 "0. 00" 5.64 18.46 3. 75 13.80 " 3.92 10.74 2.43 11.01 5.29 147T5" 15.18 44.38 3.24 9.42 14.88 " 39.69 90.65 405.85 57.05 237. 32 TREE NO. 80 SPECIES NO. 1 LENGTH ( M) 31.2 DBH CM) 0. 564 CROWN DIA(M) 6.2 CROWN BUTT L.tM) L.tMl 25.8 5.5 1 3 5 7 9 10 1 3 5 7 9 1 0 NITROGEN - 0. 05 3 0.036 0.052 0.049 0.064 0.071 NITROGEN - 0.199 0.158 0. 168 0. 186 0. 264 0.319 PHOSPHORUS- 0. 020 0. 006 0.010 0. 023 0.0 24 0. 013 PHOSPHORUS- 0.073 0.055 0.047 0.051 0. 058 0.072 CAICIU" 0.056 0.058 0.053 0. 078 0.065 0. 043 CALCIUM 0.3 71 0.208 0. 147 0.235 "" 0; 155 0.185 MAGNESIUM - 0. 01 7 0. 015 0.014 0.024 0.019 0. 014 MAGNESIUM - 0.016 0.020 0. 017 0.021 0. 023 0.034 POTASSIUM - 0.091 0.051 0.052 0. 069 0.093 0. 063 POTASSIUM - 0.072 0.115 0.093 0.104 0. 129 0. 187 BRANCH BRANCH —TW1GS- * -FOL IAGE- * CONES * TWIG S+FOLI AGE * >1 1-1/4 (TOPI ( MID) ( LOW) ' (TOP ) ( M ID) " (LOW) (TOP ) (MID) 1 LOWI NITROGEN - 0. 135 0. 162 0.471 0.353 0.386 0. 902 0. 854 0. 845 0. 000 T_xijS6 0.6 04 O.bJJf PHOSPHORUS- 0. 033 0. 037 0. 351 0. 061 0.066 0. 121 0.133 0.132 0.000 0.475--. --0T456 CALCIUM 0. 146 0.172 0.204 0. 194 0.183 0. 222 0. 208 0.265 0.000 0.213 0.224 MAGNESIUM - 3. 022 0. 027 0.040 0. 042 0.048 0.060 0. 057 0.072 0.000 0.05JJ---"0. 050^ -»J3_t060 POTASSIUM - 0. 37 3 0.083 0.164 0.139 0. 173 0. 307 0.357 0.278 0.000 j8<?35 0.243 o7z>6. •••PERCENT NUTRIENT CONTENT BY SAMPLE SPECIF! -WOOD^SECT ION- « BAP.K**SECTION-•••NUTRIENT AMOUNTS IN DECAGRAMS BY SAMPLE SPECIFICATION************************************************************************** -WOOD**SECTION- -BARK**SECTI ON- T O 0.38 ""07"0"9~ 0. 22 0.04 TOTAL 50.59 ~T5784~ 63. 64 5.31 2 8. 04 1C2 3C4 NITROGEN - 21.0313.62 5C6 9.05 7t8 4.65 0.68 10 TOTAL 0.15 49.22 1C2 3S4 NITROGEN - 21.11 12.45 5C6 7.29 PHOSPHORUS-CALCIUM MAGNESIUM -6.39 2.51 2.90 27.06 17. 15 11.66 7.64 4.51 3.44 7C8 8.30 1.92 0.25 0.03 14.00 PHOSPHORUS- 7.57 3.92 2.03 2.0l~ 5.87 0.68 0.09 62.50 CALCIUM - 34. 19 13.52 7.37 7.21 1.78 0.20 0.03 17.60 MAGNESIUM - 2.16 1.43 0.78 0.80 1.06 "0723" 0. 62 0.09 POTASSIUM 31.45 15.97 10.88 6.67 0.98 0.13 66.09 POTASSIUM - 11.03 7.93 4.05 4.2 9 0. 51 0. 22 BRANCH BRANCH •- -TWIGS- -FOLIAGE- CONES >1 1-1/4 (TOP) (MID I (LOW) TOTAL (TOP I (MID) NITROGEN - 14.47 0. 79 7.72 8.91 3. 90 20. 53 15.51 30. 23 PHOSPHORUS- 3. 51 0. 18 0.84 1.53 0.67 3.04 2.08 4.71 CALCIUM 15.62 0. 34 3. 35 4. 90 1. 85 10.10 3.83 7.37 MAGNESIUM - 2.37 0. 13 0.66 1.07 0. 49 2.22 1.02 2. 01 POTASSIUM - 7.47 0. 43 2.68 3.50 1. 75 7.93 5.29" 12.62 "(LOW) TOTAL TWIGS•FOLI AGE (TOP) (MID) (LOW) 14.78 60.53 0.00 23.23 39.14 18.69 1 96.13 9. 09 0. 00 15.83 0.00 4.29 0.00 "22777 7j;-ffO" 2.91 7.18 1.69 "7.97 ~6"7Z4" T T W 12.27 6.48 3.08 1.75 16. 12 " "6761 "T27TT" 25.93 6. 51 30.70" 45.67 168. 53 31.92 T32. 73' 2.31 4.64 1.26 "578 7"" TREE NO. 105 SPECIES NO. 1 LENGTH <M> 31.9 OBH (H ) 0.5 36 CROWN DIA(MI 4.7 CROWN BUTT L.(H) L.(MI 23.5 8.4 •••PERCENT NUTRIENT CONTENT * I 3__ BY SAMPLE SPECIFI CAT I ON** ********* ****••» *»»**»«**«****«*•* *»****•»* «*###**** — WOOD**SECTION -* * 6ARK**SECT ION * 5 1 2 10_ 1 3 5 7 g 10 NITROGEN -PHOSPHORUS- 0.030 _0i.031 CALCIUM MAGNESIUM -POTASSIUM -0.052 0.015 0.052 0.057 0.015 0.015 0.065 0.054 0.055 0.02 0 0. 061 0.030 0.062 0.024 0. 075 0.019 0. 062 0.014 0.056 0. 074 0. 023 0.071 NITROGEN -PHOSPHORUS- 0.208 0. 083 0.057 0. 019 0.073 0. 04 8 0. 015 0. 064 CALCIUM MAGNESIUM POTASSIUM 0.280 0.01 8 0. 083 0.174 0. 072 0.275 0.018 0.090 0. 161 0. 071 0. 184 0.017 0. 095 0.179 0. 066 " 0". 2 36" 0.021 0.098 0.222 0.074 0.152 0 0.025 0.034 0. 128 0.169 0.275 0.081 1 59" NITROGEN PHOSPHORUS-CALCIUM MAGNESIUM -BRANCH BRANCH 1-1/4 >1 0. 127 0. 03 3 0. 177 0.02 2 POTASSIUM - 0.075 0. 194 0.043 0. 161 0.092 " (TOP) 0.462 -TWIGS-(MID) 0.483 ( LOW) 0. 396 0.063 0. 173 0.038 0.146 0.079 0. 187 0. 043 0.069 0.163 0.053 (TOP) 0. 102 0. 322 0. 068 -FOLIAGE-(MIDI 0.870 0.845 0. 148 0.176 0.273 0. 278 (LOW I 0.854 CONES _* TW_LGS*FOl I*.G_E (TOP) (MID)"" " (LOW) • ••NUTRIENT AMOUNTS IN DECAGRAMS BY SAMPLE SPECIFICATION**"*************************"***»• ******"****-*TS***^»T(.^^t?»^«^S1«T«W*^* -WOOD** SECT I ON * 162 3E4 5 66 7C8 10 TOTAL -BARK**SECTION-1£2 3C4 NITROGEN - _29.71 16.27 11.08 6.38 PHOSPHORUS-CALCIUM MAGNESIUM -1 0.49 24.65 6. 86 1.34 5.39 18.09 4.41 4. 84 12.96 3. 55 0.42 65.20 NITROGEN 2.83 6.81 2.20 0.52 1.23 0.41 0.11 0.26 0.08 24.17 63. 99 17. 52 POTASSIUM- 26.72 16.71 12.17 7.54 1.59 . 0.35 .65. 07 PHOSPHORUS-CALCIUM MAGNESIUM -21.42 _6.05 6.8 5 21.96 1 .69 8.68 31. 09 2.02 5C6 9.70 "3.91 12.01 1.08 768 5_.3 7_ "1.86 5.20 0.62 POTASSIUM 9 10 1.27 0.27 "0.42 0.08" 0.86 0.16 0,14 0.03 TOTAL 54. 08 '21."8 0" 71.28 5. 59 9.68 8.82 5.51 3.03 0.73 0. 17 _BRANCH_BRANCH_*: " " 1 - 1 / 4 >1 r r — r — T W I G S ^ * *—-----FOLIAGE-— * CONES * TWIGS*FOLIAGE * (TOP) (MID) (LOW) TOTAL (TOP)"" (MIDI (LOW I TOTAL (TOPI (MID) " (LOW) "TQTAL" NITROGEN 10. 72 7. 36 6. 43 FHOSPHORUS- 2. 83 1.64 0. 95 CALCIUM - 14.92 6. 12 2.40 MAGNESIUM - 1. .33 1.11 0.52 POTASS IUM - 6.33 3.49 2.03 6.09 .00 4.79 17.30 15.14 21.72 25.55 62.40 1 . 2.36 0.54 1.87 0. 83 2. 04 0.64 2. 13 2.73 6.79 1.70 6.03 1.78 5. 60 1.19 4.76 1.92 21.57 27. 81 30. 34 2. 95 8.64 1.81 7.14 " 3.30 11.61 2.29 7.69" 79. 71 8.03 25. 85 5.30 19.59 0.35 0.05 0.10 0.64 2.73 8.01 1.71 6.79 27.94 TREE TOTAL" 218.99 3.95 10.99 2.35 9. 01 4. 13 10.81 13. 65 32.65 2.93 6.99 9.62 25.62 61 . 6l 189.01 33. 13 "129.09 TREE NO. 110 SPECIES NO. 2 LENGTH (HI 41.2 DBH ( M) 0. 904 CROWN DIA(M) 8.4 CROWN BUTT L.(M) L.(M) 28.4 12.8 1 3 5 • 7 9 10 1 3 5 7 9 10 — i NITROGEN - 0, 045 0. 045 0.036 0. 046 0.048 0. 059 NITROGEN - 0.212 0. 178 0. 177 0.175 0. 235 0. 251 £ PHOSPHORUS- 0. 00. 0. 004 0. 004 0.007 0.006 0. 009 PHOSPHORUS- 0.051 0. 042 0.035 0.045 0. 052 0. 046 1 CALCIUM 0. 054 0. 058 0.05 5 0.070 .0.06 5 0. 062 CALCIUM 0.298 0. 487 0. 507 ' 0.433 " 0. 426 "" " "0. 566 t' MAGNESIUM - 0. 010 0. Oi l 0.009 0.014 0.013 0. 012 MAGNESIUM - 0. 022 0. 029 0. 030 0. 033 0. 041 0. 042 POTASSIUM - 0. 060 0. 053 0.051 0. 057 0.068 0. 068 POTASSIUM - 0.108 0. 116 0. 130 0.1 21 0. 143 0. 1 26 BRANCH BRANCH -TWIGS — « *- -FOLIAGE- _CQNES__ • ---TWIGS+FOLIAGE * i j i >1 1-1/4 (TOPI (MID) ( LOW) (TOP) (MID) (LOW 1 (TOP) (MID) (LOW) • NITROGEN - 0. 157 0. 231 0.000 o.odo 0.000 0. 000 0. 000 0.000 1.041 0. 715 0. 814 0.794 | PHOSPHORUS- 0. 020 0. 036 0. 000 0. 000 0.000 0. 000 0. 000 0.000 0.226 0. 094 0. 104 0.110 't CALCIUM 0. 440 0. 331 0.000 0.000 0.000 0. 000 0. 000 0.000 0. 049 0. 345 0.410 0.457 MAGNESIUM - 0. 031 0. 02 7 0.000 0.000 0.000 0. 000 0.000 0.000 0. 070 0. 045 0. 053 0.067 I I POTASSIUM - 0. 10J 0. 137 0.000 0.000 0.000 0. 000 0. 000 0.000 0.694 0. 369 0.311 0.372 • ••PERCENT NUTRIENT CONTENT BY SAMPLE SPEC IF ICAT I ON****************************************************************************** * -WOOD**SECTION- -BARK**SECTION-•••NUTRIENT AMOUNTS IN DECAGRAMS BY SAMPLE SPECIFICATION******************************************************* •wnOD*«SECTION- -BARK**SECTION-U2 364 5£6 7C8 10 TOTAL 1£2 3£4 5C6 NITROGEN 55.00 37.37 21.69 13.78 3.83 1.52 133. 18 NITROGEN 30. 42 41.77 29.80 PHOSPHORUS-CALCIUM MAGNESIUM -4.54 3.65 67.82 52. 22 13.10 9.63 3. 35 33.36 6.26 1.92 0.45 19.77 5.14 4.00 1.06 0.24 13.84 1.61 179.92 0.32 34.40 PHOSPHORUS-CALCIUM MAGNESIUM -7.23 9.00 61.24 116.92 4.01 6.94 6.73 79.43 5.28 7£8 18.49 4. 38.71 3.30 9 10 5. 19 3.42_ 1". 14 "0.62 9,41 7.69 0.90 0.57 TOTAL 129.10 " 29.08 313.41 21 .00 POTASSIUM 63.21 48.03 23.70 18.35 5.43 1.76 170.49 POTASSIUM - 17.53 28.94 21.22 11.92 3. 16 1.71 BRANCH 8RANCH *- -TWIGS- -FOLI AGE- CONES *• >1 1-1/4 (TOPI (MID) (LOWI TOTAL (TOP I (MIDI "(LOW) TOTAL JWIGS+FOLIAGE— (TOP I (MID)(LOW) TOTAL' 84.50 TREE "TOTAL" NITROGEN - 52.66 16.32 0.00 0.00 0. 00 0.00 0. 00 0.00 0.00 0.00 1.25 54.59 63.66 31.90 150.15 482.66 0.00 0.2 7 0.00 0.06 0.00 0.08 "0.00 0.83" 4.41 T577T" 18.37 76.82 2.70 10.25 14.97 67.45 7TT2T" 741. 04 77.91 "3 66."53" PHOSPHORUS-CALCIUM MAGNESIUM -6. 77 147. 45 10. 26 2. 53 2 3. 38 1.91 POTASSIUM - 33. 57 9. 70 0. 00 0. 00 0. 00 0.00" 0.00 0.00 0.00 0.00 0. 00 0. 00 "0.00 0.00" 0.00 0.00 0.00 0.00 q.oo o.oo 0.00 0.00 0.00 0.00 0.00 0.00 0. 00 0.00 0.00 0.00 7.21 26.35 3.42 "28;i7"' a. io 32.10 4.13 24. 31 W.*E9 1E*ST . O r » . . : » 0 ' l . M * V 00*0 Ei. * t T S0*£ 9»7*S ZZ*£ 8S*E _50 * I 85*1 St'0 _ 6 i M 69**>Z 6T*^ 90*1 96 *T 8 f l 00*0 80*E ZA*0 L^'l 06*6 11 ° I "\l *6"" 6<7 '0"~" 6 Z *0"' £9 °0 15*021 TZ'ST. 8 i c E 03*9 £9**/ 00 *0 l^'OT LZ'Z 09** <?**£ 6£**> T V T 6t "Z 61*1 »<9 *S 59*5Z _.8'S 8£*T 86*Z OS'T 00*0 »*>"7 £6*0 S£*Z 9TT, £*>*! S*/*0 £9 '0 SE*0 99'0 *S '£ - KPISSVlOd £ T T. - '(.IHSaNOVh OZMl - kfllDlWD e i ' I -SOMOHdSOHd 18*911 *>8*3E 98*6 *0Z'AT 8A*II 00*0 9!'0£ £6*9 8S*EI 99*6 j w i o i _ i v i c i 3381 * 89*8 £6 °Z Z9*E £ T Z 8/ *E SE "9 N3'JfJblIN I MOT )....•( 01 W)...(d01). jy.l01_._.(MDl)_...(OIW)_ (dOl) 1V101_ (MOD ( 01W ) J dOl) *t I 1-1 --39VI103+S9IM1 * S3N03 * 39VI103 * * S9 I Ml - — . ; _ » HONVfcB '"HONVoB 9*r'<7 U*S - wnissviod TO t-8*0 1Z*T 6 T *S £8*6 68 * ET 56*iT - Hfl l SSVlOd 86*6 i6*0 V9 *5T _09*Z 9T*£I - rtfllSSNOVW wnmvD -SflTOHdSOHd - N390H1IN 80*91 60*0S 9E*££ 60*0 61*0 60*0 8Z*0 8E*0 £1*1 8E*0 56*1 8i*S SS'T i9*E T*7*TT LX'Z LZ'<7 61 *£T 8Z*Z ZA'S 1V8T. £?** •;6*0 86*£ 65 *9 6«7*8 6 C E I - Wfll S=M)VK -snautidSOHd - N390»1IN 1?101 01 83A 93 5 *3£ Z3I "1V101 01 83A 93 5 <;3£ -NOU03S*#X'dV8- -N0li33S*»Q0OM- Z31 _jt*aA»*JL»»-»»JL«JJL»«^ d I 03d5 3TdHV5 A8 SHVMOV330 NI S1NDCHV lN3 IK inN»»» 1 •") o OATO SAO'C A90*0 - wnissviod 65 I *0 - HOI S3N'9Vh -SnsOHdSOHd - N320biIN (MOT ) t a i w ) IdOl) -39»nO3+S0IMl » S3N03 (MOD IOIW) (dOl) (MOD (01W) ( d C U ) Wl-T T< -39VI103- —S9IM1- HDNV H3NS'o8 901*0 ZZO'O 91 £ *0_ SSO'C <7AZ*0 160*0 610*0 _80_£_!0_ ^6 *b 6ZZ*0 080*0 <7l0*0 08T *0_ 5 £0*0 £8T*0 580*0 M0*0 T i l *0 AEO *0 8AI *0 EAO'O ZIZ'O 8E0' 9AI' 901*0 910*0 6ZZ*0 WO *0 A£Z*0 - wnissviod - wniS3N9VW wnmvo -SfldOHdSOHd - N390dilN AIZ*0 zzo *0 BW *0 £Z0*0 ZAO *0 *<A0*0 ZZ0°O 590*0 ZZO'O */S0*0 190*0 TEO *0 ?60 *0 OZO °0 *50 *0 Z90*0 EZO*0 VAO *0 UC'O Z<70*0 8A0 *0 OZC'O 850*0 Z7.6~*0 £<70*0 £60 *0 I zo • c AAO *C ZZO *0 *>50*0 - wnissviod - KflIS3N9VH -sn-aoHdSOHd - N390M1IN 01 6 A 5 £ 1 01 6 Z : I £ I „ NOI103S**iiaV8 * » N01i03S**OOOM o 31dWVS AB 1N31N0D !N'3HiinN lN33«3d*»» S**/ 9*0Z i ' V lt<?'0 T S Z T 611 (W)*1 (H)*T (W)VIO (H> (H) *0N *0N 1108 NMOBD NMOdO H8J H19N31 S3 ID3dS 33U1 o V r TREE NO. 159 SPECIES NO. 2 LENGTH ( Ml 29.9 D8H ( M ) 0. 485 CROWN DIA(M) 8.1 CROWN BUTT L.IMI L.(M) 18.1 11.8 • ••PERCENT NUTRIENT CONTENT BY SAMPLE SPEC IFICATION****************•*•*•••***••*»**•****•«•**•*••»•**»••••••••*«**•*••••»»»*****•* • WOOD**SECT ION * * BARK**SECTI ON * 1 3 5 7 9 10 1 3 5 7 9 10 NITROGEN -JIHOSPHORUS; CALCIUM MAGNESIUM -POTASSIUM -0. 048 jj.00_6_ 0.056 0. C03 0.075 0.047 _0.CJ7_ 6.064 0.011 0.054 0.053 . JJ.006 0.06 7 0.014 0.031 0.042 _0_.„005 6. 067 0. 012 0.076 0.050 _0.009 0.051' 0.010 0.061 0. 068 J ) . 010_ 0. 061 " 0.011 0. 07 0 NITROGEN -PHOSPHORUS-0.2 54 0.037 CALCIUM MAGNESIUM -POTASSIUM -0.631 0.018 0.112 0.203 _C_PJ33 0.596" 0.028 0.100 0.232 0. 048_ 0.632" 0.038 0. 127 0.254 0.044 6.540 0. 046 0.132 0.369 0.440 0. 056 0.066 0.795" 0.616" 0, 051 0.053 0.199 0.263 BRANCH BRANCH i - i / 4 >1 _ • - - - — ' (TOP) -JWIGS-( MID) -----*_ "(LOW) (TOP I -FOLIAGE-\ (MIDI "" (LOW) CONES _* TWIGS* FOL I AGE * ""(TOP) """ (MID) "" (LOW) NITROGEN - 0. 177 0.249 0.000 0.000 0.000 0. 000 0. 000 0.000 1.211 0.543 0. 881 0. 835 PHOSPHORUS-CALCIUM MAGNESIUM -POTASSIUM 0. 024 0. 282 0.024 0. 035 0.274 0. 030 0.099 0.123 0. 000 0.000 0.00 0 0. 000 0.000 0. 000 0.000 0.000 0.000 0. 000 0. 000 0.000 0.000 0. 000 0. 000 0.000 0. 000 0.000 0.216 0.030 0.075 0.000 0.000 0.000 0.000 0.000 0.000 0.468 0.076 0.509 0.063 0.442 0.117 0.123 0.536 0.534 0. 064 0.063 0.397 0.383 • ••NUTRIENT AMOUNTS IN DECAGRAMS BY SAMPLE SPECIFICAT] »•••*•«•••••• -W000+* SECTION— -BARK**SECTION-TOTAL 162 364 566 768 10 TOTAL 162 364 566 768 T O -NITROGEN - 15.97 11.32 7.26 3.8 8 0.99 0. 3e 40. 30 NITROGEN -PHOSPHORUS-CALCIUM MAGNESIUM -2.15 20.15 3,24 1.50 15.47 2. 89 0.85 10.23 1.96 0.59 4.97 0.91 0.17 1.00 0. 19 0.05 0.34 0.06 5. 32 52. 15 9.25 PHOSPHORUS-CALCIUM MAGNESIUM -17.42 13.78 13.18 2. 68 46.77 1. 78 2.55 38.88 2.11 2.48 31.76 2.30 8.48 1. 36 18.14 1.32 POTASSIUM - 21.56 15.91 11.96 5. 75 1. 0.39 56.76 POTASSIUM - 8.06 7.18 7.03 4.50 2.11 "0."3'2 4. 53 0. 29 1. 13 1.14 56.11 **07i7 '9757"" 1.59 141.67 0.14 7.93 0. 68 28. 59 BRANCH BRANCH •- -TWIGS * * >1 NITROGEN - 9.46 10.44 0. 00 0.00 0. 00 FOLI AGE -1-1/4 (TOP) (MID) (LOW) TOTAL (TOP) (MID) (LOW) TOTAL 0.00 0.00 0.00 0. 00 CONES • TW I GS • FOL I AGE — (TOP I (MID) (LOWI 0.00 0.00 15.87 18.33 7.68 TOTAL" 41.88 TREE "TOTAL 158.19 PHOSPHORUS- 1. 30 1.47 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0. 00 2.23 2. 43 *"", 1 3" 5*779 2"2.44 CALCIUM - 15.05 11.46 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 14.84 11.15 4.91 30.91 251.24 MAGNESIUM - 1.29 O._00'_ 0.00_ 0.00 0.00 0.00 0.00 0.00 0.00 0. 00 _ 1.85 1. 33 0.58 3. 76 23. 51 POTASSIUM - 5. 26 5. 36 0.00* 0.00 0. OO" "6.66 0.66" 0.00"* 0.00 0.60 "070"0" "12.90" 7.97 3.65 " 24. 53 " 1 20.50 TREE N O . 1 7 5 S P E C I E S N O . 2 L E N G T H (Ml 3 8 . 4 OBH (Ml 0 . 7 1 1 CR OWN 01 A I M ) 8 . 5 CROWN BUTT L.(M) L.(M) 2 8 . 9 9 . 6 • • • P E R C E N T N U T R I E N T CONTENT BY SAMPLE S P E C I F I CAT I 0 N » * * * * * * * * * * * * * * * * * * # * * * * * * * * » * * * * * « » « ^ ^ * WOOD*^SECT ION * * B A R K * * S E C T ION » 1 2 5 7 9 10 1 3 5 7 9 1 0 N I T R O G E N -P H O S P H O R U S -0 . 0 4 4 _0_. 0 0 3 C A L C I U M MAGNESIUM POTASS IUM 0 . 0 6 2 0 . 0 1 2 0 . 0 5 3 0 . 0 4 5 <L.°OJ_ 0 . 112 0 . 0 2 5 0 . 0 8 1 0 . 0 5 3 0 . 0 0 9 0 . 0 6 7 0 . 0 1 3 0 . 0 5 1 0 . 038 _0_.J3 03_ 0 . 076 0 . 0 1 3 0 . 0 5 4 0 . 0 6 1 0 . 0 1 0 0 . 0 4 9 0 . 0 1 1 0 . 0 6 7 0 . 066 0 . 0 1 1 0 . 061 0 . 015 0 . 0 6 4 N ITROGEN -PHOSPHORUS-0 . 2 3 8 0 . 0 3 8 0 . 2 3 4 0 . 0 5 0 0 . 2 7 2 0 . 0 5 7 C A L C I U M MAGNESIUM -P O T A S S I U M -0 . 4 6 5 0 . 0 1 9 0 . 1 3 4 0 . 4 3 1 0 . 0 3 3 0 . 1 4 3 0 . 4 1 1 0 . 0 4 3 0 . 126 0 . 2 6 4 0 . 0 5 9 0 . 4 1 8 0 . 0 4 4 0 . 165 0 . 4 0 0 0 . 075_ ' 0 . 4 1 5 0 . 0 4 5 0 . 199 0 . 4 1 5 _ 0 . 0 83_ 6 . 321 0 . 0 4 5 0 . 2 5 2 BRANCH BRANCH >1 1 - 1 / 4 ( T O P ) - T W I G S -( MID) (LOW) - F O L I A G E -IT OP) (MID) I LOW) CONES * TWIGS *FOLI AGE (TOP I "(MID)' ( LOW) N I T R O G E N - 0 . 1 6 0 0 . 2 4 6 P H O S P H O R U S -C A L C I U M MAGNESIUM -P O T A S S I U M 0. 025 0 . 252 _ 3 . 3 2 7_ 0 . 101 0 . 0 3 6 0 . 2 3 9 0 . 0 2 9 0 . 1 3 7 0 . 0 0 0 0 . 0 0 0 0 . 0 0 0 0 . 0 0 0 0 . 0 0 0 o. oop_ o. odo o. obo 0 . 0 0 0 0 . 0 0 0 0. 0 0 0 0 . 0 0 0 0 . 0 0 0 1 . 5 6 1 0 . 7 5 9 , 0 . 501 0 . 0 0 0 0 . 0 0 0 0 . 0 0 0 0 . 0 0 0 o. ooo 0. 00 0 0 . 8 3 3 0 . 0 0 0 0 . 0 0 0 0 . 0 0 0 0 . 0 0 0 0 . 0 0 0 0 . 0 0 0 0 . 0 0 0 0 . 0 0 0 0 . 0 0 0 0 . 0 0 0 0 . 0 0 0 0 . 2 7 7 0 . 0 3 0 0 . 088 0 . 734 0 . 1 0 6 0 . 4 4 3 0 . 0 7 6 0 . 3 2 8 0 . 0 9 2 0 . 1 11 0 . 4 6 2 0 . 4 9 4 0 . 0 6 0 0 . 0 5 8 0 . 2 76 0 . 3 0 6 • ••NUTRIENT AMOUNTS IN DECAGRAMS BY SAMPLE S P E C IF ICAT I 0N + * * * * * * » •••********•***•*>*• * * » * * » * * * * » * « * " " ^ ^ - W P 0 0 ^ * S E C T I 0 N -_NI T R 3 G E N P H O S P H O R U S -C A L C I U M MAGNESIUM -1£2 2 . 2 6 6 4 . 3 1 1 4 . 0 3 3 £4 3 . 1 0 4 7 . 3 6 1 0 . 2 7 5 £6 3 3 . 0 3 2 6 . 0 0 1 5 . 6 6 7E8 9 . 0 8 1 . 9 7 2 4 . 3 1 4 . 5 0 1 . 1 9 1 1 . 4 4 2 . 2 3 2 . 3 1 0 . 3 9 1 . 8 7 0 . 4 3 1 0 J?i 6JL 0 . 1 1 0 . 6 2 0 . 1 5 TOTAL 8 6 . 7 6 9 . 02 1 5 0 . 4 1 3 1 . 6 0 P O T A S S I U M - 5 0 . 1 0 3 5 . 0 5 1 7 . 9 2 1 1 . 0 4 2 . 5 3 0 . 6 5 1 1 7 . 30 - B A R K * » S E C T I O N -1£2 P H O S P H O R U S -C A L C I U M MAGNESIUM -3 £4 N ITROGEN - 3 7 . 25 2 2 . 8 2 6 . 9 8 4 . 8 3 7 0 . 80 3 8 . 0 0 4 . 0 8 3 . 4 4 5 £6 J 4 . 7 0_ " 3 . 1 8 2 2 . 7 5 2 . 3 9 7 £ 8 _ 8 ' 0 3 _ " " 1 . 6 2 1 0 . 0 7 1 . 0 7 9 1 0 2 . 5 6 1 . 0 0 0 . 4 8 0 . 2 0 2 . 6 5 0 . 7 7 0 . 2 9 0 . 1 1 TOTAL 8 6 . 3 5 1 7 . 2 8 " 1 4 5 . 0 5 1 1 . 3 8 P O T A S S I U M 2 1 . 9 0 1 2 . 1 3 7 . 9 8 4 . 4 0 1 7 2 7 0 . 6 0 4 8 . 2 9 BRANCH J 3 R A N C H _ ">1 1 - 1 / 4 ( T O P ) - - T W I G S • (MID) "(LOW) * *. TOTAL ( TOP") - F O L I A G E - - - - -(MID) " (LOWI « "TOTAL" _CONES_ • - TWIGS* FOL I AGE — " ( T O P ) " (MID) " (LOW) T O T A L " T R E E TOTAL" N I T R O G E N - 2 7 . 76 1 8 . 11 0 . 0 0 0 . 0 0 0 . 0 0 0 . 0 0 0 . 0 0 0 . 0 0 0 . 0 0 0 . 0 0 0 . 0 0 5 1 . 5 9 3 9 . 32 3 7 . 0 8 1 2 7 . 9 8 P H O S P H O R U S - 4 . 37 2 . 6 6 0 . 0 0 0 . 0 0 0 . 0 0 0 . 0 0 0 . 0 0 0 . 0 0 0 . 0 0 0 . 0 0 0 . 0 0 7 . T 8 7 , 22 C A L C I U M - 4 3 . 32 1 7 . 6 1 0 . 0 0 0 . 0 0 0 . 0 0 0 . 0 0 0 . 0 0 0 . 00 0 . 0 0 0 . 0 0 0 . 0 0 3 0 . 1 2 3 6 . 2 8 _M.AGNESXU_M_J: 4._7.1_ 2 . 1 4 0 . 0 0 _ 0 . 0 0 _ 0 . 0 0 _ 0 . 0 0 ^ 0 . 0 0 0 . 0 0 0 . 0 0 0 . 0 0 0 . 0 0 5 . 1 4 4 . 74 P O T A S S I U M - 1 7 . 5 7 1 0 . 1 1 0 . 0 0 0 . 0 0 0 . 0 0 0 . 0 0 ' " " " 6 . 0 0 ' 0 . 0 0 " " 6 . 0 0 " 0 . 0 0 "0". 0 0 " ' 2 2 . 3 3 ' " 21". 64 ' 4 . 9 3 1 9 . 3 3 2 1 . 9 9 8 8 . 3 9 2 . 5 9 1 2 . 4 8 1 3 . 6 1 5 7 . 5 8 3 4 6 . 9 6 5 2 . 6 7 4 4 5 . 2 8 6 2 . 3 1 "2 5 0 . 8 5 TREE SPECIES LENGTH DBH CROWN CROWN BUTT NO. NO. (M) (M) DIA(M) L.(M) L.(M) 176 2 26. 3 0. 361 0.0 19.6 6.6 *** PER CENT NUTR I ENT CONTENT BY SAMPLE SPECIF I CANTON ********************************** ***'"* * WOOD** SECT I ON * * BARK** SECT I ON * 1 3 5 7 9 10 1 3 5 7 9 10 NITROGEN - 0. 063 0.041 0.045 0.050 0. 059 0. 063 NITROGEN - 0.242 0.226 0. 246 0. 265 0. 359 0.409 PHOSPHORUS- 0. Oil 0. 004 0. 007 0.008 0. 009 0. 012 PHOSPHORUS- 0.041 0.044 0.051 0. 053 0. 074 0.095 CALCIUM 0. 043 0.061 0.060 0. 059 0. 060 0. 051 CALCIUM 0.675 0.480 0.482 "0. "591" b. 413 0.550 MAGNESIUM - 0. 007 0. OIJ 0.007 0. 009 0. 009 0. 010 MAGNESIUM - 0.010 0.022 0.022 0. 023 0. 02 5 0.021 POTASSIUM - 0. 054 0.054 0. 059 0. 065 0. 073 0. 073 POTASSIUM - 0.097 0.158 0.134 0. 146 0. 143 0.237 BRANCH BRANCH -TWIGS-- --* -FOLIAGE- CONES * TWIGS+ FOLIAGE---* >1 1-1/4 (TOP) (MID) ( LOW) (TOP) (M ID) (LOW) (TOP) (MID) ( LOW) NITROGEN - 0. 00) 0.000 0. 000 0.000 0. 000 0. 000 0.000 0.000 0. 000 0.000 0. 000 0. 000 PHOSOHORUS- 0. 000 0. 000 0. 00 0 0. 000 0. 000 0. 000 0.000 0.000 0.000 O.COO 0.000 0. 000 CALCI'JM 0. 000 ' 0.000 0.000 0. 000 0. 000 0. 000 0. 000 0. 000 0. 000 0.000 0.000 0. 000 MAGNF SIUM - 0. 000 0.000 0. 000 0.000 0. 000 0. 000 0. 000 0.000 0.000 0.000 0.000 0. 000 POTASSIUM - 0. 000 0.000 0. 000 0. 000 0. 000 0. 000 0. 000 0.000 0.000 0.000 0.000 0. 000 •••NUTRIENT AMOUNTS IN DECAGRAMS BY SAMPLE SPECIFICATION************************************************************************** a WCOD**SECT ION * * BARK**SECTION * 162 3£4 5C6 768 9 10 TOTAL 112 3C4 5C6 7C8 9 10 TOTAL NITROGEN - 9.25 4. 64 3.19 1.95 0.41 0.10 18. 55 NITROGEN - 6. 58 3.76 2.74 1.50 0.43 0.16 15. 17 PHOSPHORUS-CALCIUM "AGNESIUM -1.17 9.25 1.27 0. 56 6. 55 0.90 0.49 4.01 0.55 0.2 9 2.14 0.3 3 0.06 0.42 0.07 0.02 0.08 0.02 2.60 21.46 3. 13 PHOSPHORUS-CALCIUM MAGNESIUM -1.20 16.23 0. 44 0.76 7.65 0.35 0.55 5.74 0.24 0.30 2.41 0.11 "OTO'9" 0. 5 0 0.03 0.0"4 0.22 0.01 2.94 32.74 1.19 POTASSIUM - 3. 53 6.09 4.19 2.49 0.51 0. 12 21.97 POTASSIUM . 58 2.32 1.49 0. 69 0. 17 0.09 BRANCH BRANCH *- -TWIGS-NITROGEN >1 0.00 1-1/4 (TOP) (MID) (LOW) TOTAL - - _- FOL I AG E "(TOP) (MID) (LOW) _«. CONES * TWI GS«-FOL I AGE * TOTAL ' "(TOP)'"""( MID)' (LOW) TOTAL 0.00 0.00 0. 00 0. 00 0.00 0.00 0. 00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 e.35 TREE "TOTAL"" 33. 72 PHOSPHORUS- 0. 00 0. 00 0. 00 0.00 0. 00 0. 00 0. 00 0.00 0.00 0.00 0. 00 0.00 0.00 0. 00 0.00 5.53 CALCIUM 0. 00 0. 00 0.00 0. 00 0. 00 0. 00 0. 00 0. 00 0.00 0.00 0.00 0.00 0.00 0. 00 0.00 54.20 MAGNESIUM - 0. 00 0. 00 0.00 0.00 0. 00 0. 00 0. 00 0. 00 0.00 0.00 0. 00 0.00 0.00 0. 00 0.00 4.32 POTASSIUM - 0. 00 0. 00 0.00 0.00 0. bo" 0. 00 b. bo " 0.00 0.00 0. 00 ' 0. 00 " "0.00 " "0. 00 0. 00" " " 0.00 30.32" TREE NO. 178 SPECIES NO. 2 LENGTH (Ml 33.9 DBH (Ml 0.884 CROWN DIA(Ml 0.0 CROWN L.IMI 28.7 BUTT L. (M) 10.2 •••PERCENT NUTRIENT CONTENT BY SAMPLE SPEC IF I CAT 10N»**«»*»«»»»»» ••*»*»•*••»«••••»»•••••••••»»»••••••»»»> • -NITROGEN -J?HO_S_PHOR_US:L_ CALCIUM MAGNESIUM -POTASSIUM -0. 067 _9i00_9_ 6. 04d O.OOd 0. 043 0.037 _9 .Q_05_ 6.076 0.013 0.048 -WOO0**SECTI0N * _> 7 9 10 0.036 0.005 0. 052 0. 006 0.095 0.013 0. 071 0.01 5 0.05 6 0. 058 0. 013 0. 093 0.080 0.025 0.1 39 0. 064 _0. 041_ 0."654 0. 012 0.064 • — — NITROGEN -PHOSPHORUS-CALCIUM MAGNESIUM -POTASSIUM -0.303 0. 045 0.496 0.023 0. 130 • • * • « • « • * * • • • • • » « • • • • • • A A A A 8ARK^*SECTI0N * 5 7 9 10 0.172 0.035 673 53 ' 0.030 0.109 0.168 0. 038_ "67313 0.025 0. 100 0.180 0.035_ ' 0.314 0.031 0.115 0.214 0.376 _0. 034_ _;0.O83_ 0.319 " 0.427 0.045 0.040 0. 172 0.267 BRANCH BRANCH >1 NITROGEN - 0.000 PHOSPHORUS-CALCIUM .MAGNESIUM -POTASSIUM -0. 00 J 0.000 0.000 1-1/4 0.000 0.000 0.000 • TWIGS * (TOP ) (MID) FOL I AGE • (LOW) (TOP ) 0.000 0. 000 0.000 0. 000 0. 000 0. 000 0.000 0.000 0.000 o.ooo ' o.ooo o.ooo 6.o66~ o.ooo 0. 000 0. 000 0. 000 (MID) 0.000 0.000 0.000 0.000 0. 000 0. 000 0.000 0. 000 0. 000 ( LOW) 0.000 0.000 0.000 0.000 0.000 0.000 0.000 CONES 0.000 0.000 0.000 _* TWJ.GS+FOLIAGE--(TOP I (MID) 0.000 0.000 0.000 o.ooo o.ooo" I LOW) 0.000 0.000 0.000 0.000 o.ooo o.ooo o.ooo o.ooo 0.000 0.000 0. 000 "o.ooo" •••NUTRIENT AMOUNTS IN DECAGRAMS BY SAMPLE SPECIFICATION*****""""""""***^  • WOOO**SECTI ON • • 162 3 64 • 5 66 768 10 TOTAL 162 BARK**SEC TION 3 64 566 768 NITROGEN PHOSPHORUS-CALCIUM MAGNESIUM -_6_*b_18 30.09 25.22 2 0.57 2.27 3.13 75.36 12.33 4.10 61.11 11.31 3.35 36.69 8.05 2.68 19.19 5.27 0.31 1.92 0.60 _0.26_141.60_ 0.T7" ' 1 8. 74 0,22 194.99 0.05 38.59 NITROGEN - 57.64 31.28 34.87 19.98 2.05 PHOSPHORUS- 9.71" 6".76" '7.36 3751 0."33" CALCIUM - 102. 88 61.28 62.91 32.13 3. 06 MAGNESIUM - 6.40 5.16 5.69 3.86 0. 43 POTASSIUM - 53.32 43.30 42.70 32.31 3.31 . 0.26 180.20 POTASSIUM - 29.00 19.22 21.57 14. 59 JTT! 10 0.68 0.15 0. 77 0.07 TOTAL 146.49 27. 32"" 263.03 21.62 J ^ A N 1 " ! 3RANCH_*R-:---------TWIGS • • >1 *"' 1-1/4 (TOP) (MID) (LOW) TOTAL (TOP) ^ F O L I A G E - — - * CONES * T Wl GS* FOL I AGE--(MID) (LOW) TOTAL "'" ( TOP) (MID) (LOWI NITROGEN - 0.00 PHOSPHORUS-CALCIUM _MAGNES IUM ^ POTASSTUM -0.00 0.00 0. 00 0. 00 0. 00 0. 00 0. 00 "o'. 00* 0. 00 0.00 0. 00 0. 00* 0. 00 0.00 0. 00 0. 00 6. oo • 0. 00 0.00 0.00 0.00 0. 00 0. 00 _ 0. 00 6. o o 6 . oo 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0. 00 6.00* " 0.00 0.00 0.00 0.00 0.00 0.00 0.00 "0.00*"" 0.00" 0.00 0.00 0.00 0.00 0. 00 0.00 0.00 "0. 00 0. 00 0. 00 0.00 "0. 00" 0. 00 0. 48 TOTAL 0.00 0.00 0.00 0.00 0.00 86. 52 TF EE TOTAL" 288. 09 0, 00 0.00 0.00 0.00 0.00 0.00 0. 00 * 0. 00 0.00 0.00 0.00 0.00 46.55 458.02 60.21 266. 7? T R E E N O . 2 1 3 S P E C I E S NO. 2 L E N G T H (Ml 30.0 0 3 H ( M ) 0 . 5 6 4 C R O W N 01 A( Ml 5 . 6 C R O W N B U T T L . ( M ) L . I M I 2 5 . 1 4 . 9 «««p£RCgNT NUTRIENT CONTENT BY SAMPLE SPECIFICATION*"****************************^ • WOOD**SECT ION * * BARK**SECT ION * 1 - 3 5 7 ? 10 1 3 5 7 9 10 0.046 0.052 0.063 0. 076 NITROGEN - 0.259 0.230 0.230 0.306 0.365 0.419 _0_.006 Oj.008 0..014 0_.01_2 PHOSPHORUS- 0.036 0.040 _ 0. 041 0.052 _ 0.062 0.072 0. 072 6.057" 0.057 6.054 CALCIUM - 6.666 0.621" "0.515 0.690 0.579 0.463 0.011 0.009 0.009 0.009 MAGNESIUM - 0.016 0.026 0.031 0.032 0.034 0.031 0.063 0.048 0.061 0.065 POTASSIUM- 0. 132 0.117 0. 148 0.131 0.172 0.226 NITROGEN -_P_HQ_$ PJ±ORUi^_ CALCIUM MAGNESIUM -POTASSIUM -0.054 0.0 07_ 051 0 0. 011 0. 061 0.047 0.006_ 6.05_4 0.010 0.055 BR ANCH BRANCH NITROGEN > 1 0. 172 PHOSPHORUS-CALCIUM MAGNESIUM -0. 022 0. 330 0.020 1 - 1 / 4 0.209 0.031 0. 313 0.022 ( T O P ) 0.000 -TWIGS-( MID) ( LOW) 0.000 0.000 (TOP I F^OL I A G E -(M I D I 0. 000 0. 000 (LOW) 0.000 CONES * TWIGS + FOL I AGE • "" '"""(TOP) "• (MID) (LOW) 1.493 0.795 0.829 0.829 0.000 0. 00 0 0 . 6b o 0. 000 0.000 0. 000 0.000 0.000 0.000 0.000 0. 000 0. 000 0. 000 0. 000 0. 000 0.000 0.000 0.000 0. 261 0.025 0. 076 0.106 0.398 0.062 0. 099 0.099 0.510 0.510 0.058 0.058 P O T A S S I U M - 0.090 0.103 0.000 0. 000 0.000 0.000 0. 000 0.000 0.564 0.384 0. 341 0.341 ~**VNUTRIENT A M O U N T S I N O E C A G R A M S B Y S A M P L E S P E C I F I C A T I O N * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * R * * * * * * * * * * * * * ^ * * * * - W O O O * * S E C T I O N - B A R K » * S E C T I O N -TC5 T O T A L 1 C 2 3 C 4 5 £ 6 768 N I T R O G E N 19.31 12. 05 7.70 4.43 0.80 10 0 . 1 9 TOTAL 4 4 . 9 9 NITROGEN -1£2 22.53 PHOSPHORUS-CALCIUM MAGNESIUM -2.53 1.65 1.09 20.35 16.39 10.20 3.95 2.66 1.56 0.33 0.18 4.36 0.72 0.6 7 0.11 0.03 0.14 0.02 6.30 52.66 8.97 PHOSPHORUS-CALCIUM MAGNESIUM -3.51 59.22 1.92 3 £4 11.16 — n s T " 27.57 1.38 5 £6 10.41 T O " TT&TT 23.33 1.22 5^22 1. 42 0.8 9 "0.24 9.84 2.26 0.52 0. 13 2.35 0.67 0.59 51.32 "OTTO -"e."5i"" 0.65 122.86 0.04 5.22 POTASSIUM - 22.74 15.39 8.35 4.22 0.78 0.16 52. 14 POTASSIUM - 11.46 6.42 5.39 0.32 BRANCH BRANCH *- -TW1GS->1 1 - 1 / 4 (TOP) (MIDI (LOW) TOTAL ( T O P ) ^FOLIAGE--— (MID) (LOW) - * _ C O N E S _ _ " T O T A L TWIGS+FOL1AGE--"(TOPT"""(MID)' ' (LOW)" TOTAL" NITROGEN 15.81 10.74 0, 00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 2 5.92 12.98 2.03 12.51 43. 17 —57 26. 18 26.58 16.1 2 3.00 1.82 "17.75 -"10.77 9 5 . 2 7 T T T 7 T " 26.60 TREE "TOTAL" 218.13 PHOSPHORUS-CALCIUM _MAGNE SIUM -POTASSIUM -2. 05 1. 57 0. 00 35.04 16.04 0.00 1.35 1. 1 4 0.00 8.25 5.55 0.00" • 0 . 0 0 0 . 0 0 0 . 0 0 0 . 0 0 _o1oo 0 . qo_ "0.06 o.'oo" 0 . 0 0 0 . 00 0 . 0 0 0 . 0 0 o.oq 0 . 0 0 "oio'o 6 . 0 0 ' 0 . 0 0 0 . 0 0 0 . 0 0 0 . 0 0 0 . 0 0 __o_oo_ 0 . 0 0 0 . 0 0 0.66 0.06 0 . 0 0 0 . 0 0 0 . 0 0 0 . 00 "0V00 o.oo 55.68 282.29 6. 65 24. 03 "41.04" 133. 58" TREE SPECIES LENGTH OBH CROWN CROWN BUTT NO. NO. (Ml (Ml DIAIMI L.(M) L.(M) 215 2 38.2 0. 805 0.0 25. 7 12.5 •••PERCENT NUTRIENT CONTENT BY SAMPLE SP EC IF I CAT I ON*** ******************* ****^^ * -WOOO**SECTION * * -8AP.K**SECTI0N * 1 2 : § 7 9 10 1 3 5 7 9 1 0 NITROGEN - 0. 061 0.044 0.046 0.045 0.086 0. 085 NITROGEN - 0.267 0.194 J 0. 208 1 0.216 1 0. 307 I u 0.391 PHOSPHORUS- 0.003 0. 006 0. 009 0.009 0.015 0. 021 PHOSPHORUS- 0.039 0.038 0.037 0.043 0. 041 0. 075 CALCIUM 0.055 0.055 0.055 0.071 0.090 0. 060 CALCIUM 0.265 0.350 0.296 0.311 0. 743 0.356 MAGNESIUM - 0.C09 0.003 0. 01 0 0. 012 0.017 0. Oi l MAGNESIUM - 0. 016 0.020 0. 022 0.031 0. 030 0. 029 POTASSIUM - 0. 051 0.042 0.042 0.054 0.060 0. 058 POTASSIUM - 0.121 0.099 0.083 0.090 0.093 0.196 BRA MCH BRANCH —TWIGS- -FOLIAGE- CONES * Tul r.^+Fni T AGP * >1 1-1/4 ( TOP 1 ( MIDI ( LOW) (TOP) (MIDI (LOW 1 ( TOP 1 (MIDI ( LOW) NITROGEN - 0. 000 0. 000 0.000 0. 000 0.000 0. 000 0.000 0.000 0.000 0.000 0. 000 0. 000 PHOSPHORUS- 0. 00) 0. 000 0. 00 0 0. 000 0.000 0. 000 0.000 0.000 0.000 0.000 0.000 0.000 CALCIUM 0. 000 0. 000 0.000 0. 000 0.000 0. 000 0. 000 0. 000 0. 000 0.000 0.000 0.000 MAGNESIUM - 0. 000 0. 000 0.00 0 0. 000 0.000 0.000 0. 000 0.000 0. 000 0. 000 0. 000 0. 000 POTASSIUM - 0. 000 0. 000 0.000 0.000 0.000 0. 000 0. 000 0.000 0. 000 0.000 0.000 0.000 • ••NUTRIENT AMOUNTS IN DECAGRAMS BY SAMPLE SPEC IF ICAT'l0N*"*>*************** *******"*T*~*T^ -WOOO»«SECTION-1£2 3£4 5(16 7£8 10 TOTAL -BARK**SECT ION-1£2 3C4 5 C 6 NITROGEN PHOSPHORUS-CALCIUM MAGNESIUM -41.12 25.73 17.46 13.07 2.93 5.43 42.61 6.85 4. 20 31.42 5.39 0.44 100.75 3.53 24.48 4.38 2.45 16.12 2.96 0.51 3.06 0.59 POTASSIUM - 36.20 24.01 18.58 11.36 2.04 0.11 0.31 0.06 NITROGEN - 47.51 31.73 34.34 16.28 118.00 20.23 PHOSPHORUS-CALCIUM MAGNESIUM -7.93 5.97 63.40 50.98 3.69 3.28 0.30 92.49 6.52 49.12 4.30 SRANCH_ BRANCH J*j; " > 1 1 - 1 / 4 ~z—r—TWIGS — * * (TOPI" TMlb") (LOW I " TOTAL ' (TOP) POTASSIUM ---FOLIAGE—— (MID)" (LOW I 7£8 9 _20.63 3.99 3.32 0.53" 41.59 9.66 2.43 0.40 10 0.47 0.0'9 0.43 0.03 22.68 14.36 14.01 7.26 TOTAL 1^ 38. 66 ~ 24."37 215.17 14.12 1.21 * _CONES T O T A L * " • TW I GS*FOL I AGE — "(TOP) (MID) " (LOW) NITROGEN - 0.00 0.00 PHOSPHORUS-CALCIUM _M_AGMES IUM_^ _ POTASS I UM -0. 00 0.00 0.00 0. 00 0. 30 0. 00 o". 66 o. oo o. oo o. oo o. oo" 0.00 0. 00 0. 00 0.00 6. oo" 0.00 0.00 0,00 0.00 0. 00 _ 0. 00 b.oo". "6".o6r 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 "" 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 "0.00 "0.00 0.00 0.00 0.24 • TOTAL ' 0.00 0.00 0. 00 0.00 0.00 0.00 0.00 "0.00 "' "0.00" 0.00 0.00 0.00 0.00 * 0.00 OT075 0.00 0.00 0.00 0.00 0.00 0.00 59.76 TREE TOTAL" 239.42 0.00 0.00 0.00 0.00 40.65 333. 18 34.35 "152.25 TREE NO. 235 SPEC!ES NO. 2 LENGTH < M ) 15.1 09H (M) 0.3*3 CROWN DIA(Ml 0.0 CROWN L.(M) 8.4 BUTT L. (M) 6. 8 • ••PERCENT NUTRIENT CONTENT BY SAMPLE SPEC I F I CAT I ON********************************************** • WOOD** SEC T I ON * * B ARK**SECT I ON * 1 3 5 7 9 10 1 3 5 7 9 10 NITROGEN -PHOSPHORUS-CALCIUM MAGNESIUM -POTASSIUM -0.053 JJ.Ol 0_ 0. 052 0.009 0. 053 0.051 _0.0 07_ 0. 06 0 0.011 0. 050 0.049. _0_.0i)6_ 0. 06 7 0.012 0.054 0. 061 J).008_ 0. 062 0. 013 0. 055 0.054 _0_.009 0.054* 0.010 0.062 0. 066 0. 012_ "0.~"053 0. O i l 0.069 NITROGEN -PHOSPHORUS-CALCIUM MAGNESIUM -POTASSIUM -0.229 0. 037_ 0.627 0. 02 0 0. 122 0.224 _0_039_ 0.583 " 0.029 0.112 0. 251 0. 046 ' 0.492" 0.042 0. 157 0.254 0.043 "O". 431 0.037 0. 182 0.313 0. 056 0.418"' 0.037 0. 180 0.377 0.058 " 0. 6 34" 0.038 0.217 _BR_ANCH_ >1 _3RANCH_ 1-1/4 (TOP) -J WIGS-(MID) * (LOW) (TOP) -FOLIAGE-(MID) (LOW I CONES * TWIGS* FOLI AGE * "(TOP)" (MIDI " (LOW) NITROGEN - 0. OOP 0.000 0.00 0 0. 000 0.000 0. 000 0. 000 0.000 0.000 0.000 0.000 0.000 PHOSPHORUS-CALCIUM MAGNESIUM -POTASSIUM P. 000 0. 000 0.000 0.000 0.000 0.000 0.000 0.000 0.000 0. OJO 0.000 0. 000 0. 000 0. 000 0.000 0.000 0.000 0. 000 0. 000 0. 000 0. 000 0.000 0. 000 0.000 0.000 0. 000 0. 000 0. 000 0. 000 0.000 0.000 0.000 0.000 0.000 0.000 0.000 0. 000 0. 000 0. 000 0.000 0.000 0.000 0.000 0.000 0.000 0.000 o_oo_o_ 0.000 • ••NUTRIENT AMOUNTS IN DECAGRAMS BY SAMPLE SPECIFICAT ION*********************************************************************»**»* -WOOD««SECTIQN- -BARK**SECTION-162 364 566 768 9 10 TOTAL 162 364 5 66 768 9 10 TOTAL NITROGEN - 4.65 3.33 2.68 1.57 0.44 0.10 12.77 NITROGEN - 3.67 3.61 2.63 2.18 0. 44 0. 19 12.71 PHOSPHORUS-CALCIUM MAGNESIUM -0.77 5.02 0. 94 0.43 4.22 0.79 0.34 3.14 0. 62 0.24 1.58 0.32 0.08 0.44 0.08 0.02 0.08 0.02 1.87 14.48 2.76 PHOSPHORUS-CALCIUM MAGNESIUM -0.61 9. 80 0.40 0.65 8.16 0.54 0.47 4.80 0.41 0.38 3.27 0.29 0.08 0, 59 0. 05 0".03 0. 32 0. 02 " 2 . 21" 26.93 1. 72 POTASSIUM - 4.36 3.48 2.66 1.59 0.50 0.10 13.19 POTASSIUM - 1.90 2.04 1.76 1.39 0. 25 0.11 7.46 BRANCH BRANCH * TWIGS->1 1-1/4 (TOP) (MID) (LOW) TOTAL FO LIAGE — (TOP) (MIO) ' (LOW) TOTAL CONES *- TWIGS* FOL1 AGE — (TOP )" (MID) (LOW) TOTAL" TREE TOTAL" NITROGEN - 0. 00 0. 00 0. 00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0, 00 0. 00 0.00 25.48 PHOSPHORUS-CALCIUM MAGNESIUM -POTASSIUM -0. 00 0. 00 0. 00 0.00 0. 00 0. 00 3. 00 0. 00 O.OP 0.03 _0. 00 o. do" 0.00 0.00 0.00 0.00 0. 00 0. 00 0.00 0.00 0.00 0.00 q.oo_ "b.dd 0. 00 0. 00 _0.00_ 0. 00 0. 00 0. 00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 "0.00" 0.00 0. 00 o.'oo 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0, 00 0. 00 "0.00 0.00" "7j~70"0" 0.00 0.00 0.00 4.08 41.41 4. i.8 "20.65" f ' r T R E E N O . 2*3 S P E C I E S N O . 2 L E N G T H ( M » 17. a O B H ( M ) 0.310 CROWN 01 A(H ) 5.5 C R O W N B U T T L . ( M ) L . I M I 11.5 6.3 •••PERCENT NUTRIENT CONTENT 8Y SAMPLE SPECIFICATION******************»»*****»************^ -WOOD** SECT I ON * * BARK** SEC TI ON * _5 7 9 10 1 3 5 7 9 1 0 * V NITROGEN -PHOSPHORUS-0. 0*8 0.006 CALCIUM MAGNESIUM -POTASSIUM -0. 060 0. 010 0.0*6 0.0*1 _0_._0_0*_ 6.05 7 0.012 0.061 0. 0*2 0. 00* 0. 0*9 0. 010 0.0*7 0.006 0.053 0. 013 0.055 0. 052 0. O i l 0. 062 0.050 0.012 0.055 0. 067 ___P_1_2_ 0. 0** 0. 01* 0. 067 NITROGEN -PHOSPHORUS-CALCIUM MAGNESIUM POTASSIUM 0.301 0.0*3 0. 5*5 0.026 0.085 0.236 0.036 0.629 0.029 0.101 0. 261 0.0*2_ "0.72 5 0.0*0 0. 131 0. 2 88 0.0** 0.5 99 0.038 0.151 0. *01 0.075 6. "76 2" 0.0*5 0.263 0. 522 0.091^ "6". 56"! 0. 0*7 0.271 BRANCH BR AN C H * T WIGS * *j- FOLIAGE » CONES * — - TW_I GS+ F OLIAGE — - * >1 1-1/* (TOP) (MID) (LOW) (TOP) (MID) (LOW) (TOP") " (MID) (LOW) NITROGEN - 0. 127 0.213 0.000 0. (JOO 0.000 0.000 0.000 0.000 0.000 0.852 0.816 0.750 PHOSPHORUS- 0.021 0.032 0.000 0.600 0.000 0. 000 0.000 0.000 0.000 0.110 0.121 0.086 CALCIUM - 0.228 0.25* 0.000 O.dOO 0.000 .0. 000 0. 000 0.000 0.000 0.370 0.339 0.269 MAGNESIUM - 0.02* 0. 03* 0.000 0.600 0.000 0. OOP 0.000 0. 000 O.J)00 0.1 00 0. 087 0_. 067 POTASSIUM - 6.071 0.115 0.000 0.000 0.000 0. 000 0.000 0.000 0.000* 0.3*6" 07311 0.271 • ••NUTRIENT AMOUNTS IN DECAGRAMS BY SAMPLE SPECIFICATION************************************************"**"************'************ -WO00**SECTION- BARK**SECTION-162 36* 566 768 9 10 TOTAL 162 36* 566 768 9 10 TOTAL NITROGEN - 3.*0 2. 10 1.3* 0.72 0.16 0.07 7. 80 NITROGEN - 3.36 3.13 2.17 1.21 0.28 0. 16 10.31 DHOSPHORUS-CALCIUM MAGNESIUM -0.37 *.*6 o.a* 0.21 2.76 0.62 0.21 1.53 0.35 0.12 0. 76 0.17 0.02 0.17 0.0* 0.01 0.05 0.02 0.9* 9. 7* 2. 0* PHOSPHORUS-CALCIUM MAGNESI UM -0.50 7.3* 0.35 0.*9 8.53 0.** 0. 3* 5.23 0.31 0. 21 2.38 0.15 " ""6." 05 0. 53 0.03 0.03 0. 17 0.01 1.61 2*. 18 1.28 POTASSIUM - 4. J9 2.92 1.71 0. B7 0.19 0.07 9.85 POTASSIUM - 1. 17 l.*7 1.12 0. 72 0. 18 0.03 _8R_ANCH_BRANCH ~>V 1-1/'* (TOP) - - T W I G S (MID) " (LOW)" TOtAL (TUP)" • - F O L I A G E ; - ^ - - ; ' " ( "MIDT~'"""L0W» CONES *-NITROGEN *. 29 3. 30 0.00 0.00 0. 00 0.00 0.00 0.00 0.00 TOTAL 0.00 -TWIGS + FOLIAGE— (TOP I (MID) " (LOW) 0.00 5.80 1*.53 *.05 TOTAL" 2*. 38 *. 7* Tf.EE "TOTAL" 50.57 PHOSPHORUS-CALCIUM _MAGNES IUM_-POTASSTUM -0. 73 0. 53 7.71 4.54 0.81__ 0.61 2. 41 2. 06 0.00 0. 00 0. 00 6. 00 0.00 0.00 0.00 0.00 0.00 0. 00 6.00 o.oo 0.00 0.00 0.00 0.00 0.00 0.00 o.ooP.OP 0.00 P. 00 0.00 0.00 0. 00 0.00 o.oo " o.oo" 0.00 0.00 0.00 0.00 0.00^ 0.00 "0.00 ** "o.oo" 0. 75" 2.51 0.68 2.36 2. 16 6.03 1. 56 5.5*" "07*"7~ l.*5 0.36 "1. *6 T7TT 9.99 2. 60 9. 3 6 7.23 56. 16 7.3* 28.41" TREE NO. 2 5 6 S P E C I E S NO. 2 LENGTH ( M ) 2 T . 9 DBH (M) 0 . 4 7 5 CROWN O I A I M ) 0.0 CROWN BUTT L . ( M ) L . ( M ) 2 0 . 3 7.6 •••PERCENT NUTRIENT CONTENT BY SAMPLE S P E C I F I C A T I O N * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * MOOD** SECT ION * * BARK **S ECT I ON * 1 3 5 7 9 10 1 3 5 7 9 10 NITROGEN -PHOSPHORUS-0 . 0 5 5 0. 004 0 . 0 4 0 0 . 0 0 3 0 . 0 4 6 0 . 0 0 5 0. 044 0 . 0 0 8 C A L C I U M MAGNESIUM P O T A S S I U M 0 . 0 5 7 0 . 0 0 9 0. 049 0 . 0 7 0 0 . 0 1 0 0 . 0 4 9 0. 061 0 . 0 1 0 0. 0 5 4 0. 065 0. 0 1 0 0. 0 6 4 0. 0 5 1 _0 . 0 1 3 _ 0."0 57 0 . 0 0 9 0 . 0 6 0 0. 067 0.015 NITROGEN -PHOSPHORUS-0.259 0.03 7 0.217 0.038 0. 048 0. O i l 0. 0 73 C A L C I U M MAGNESIUM -POTASSIUM -0. 4 6 0 0.015 0. 0 9 9 0.348 0.025 0.086 0.226 0. 0 4 1 _ " 0 . 3 4 1 0.029 0.104 0 . 2 8 4 0.04 7 6. 524* 0.035 0. 143 0. 3 1 5 0 . 0 6 2 _ ""0.390 0.030 0. 1 5 7 0.454 _0.0 98 0 . 4 1 6 " 0.031 0.239 NITROGEN -BRANCH BRANCH >1 0 . 0 0 0 1-1/4 0 . 0 0 0 ( T O P ) 0 . 0 0 0 -TWIGS-( M l b ) * 7 L b w ) " o.ooo o.ooo _* --j-FOL I AGE-(TOP) "(MID)'" 0. 0 0 0 0.000 (LOW I 0 . 0 0 0 CONES 0.000 * TWIGS+FOL I AGE * ""( TOP) " ' (MID) (LOW ) 0 . 0 0 0 0.000 0 . 0 0 0 PHOSPHORUS-C A L C I U M MAGNESIUM -0. 0 0 3 0 . 0 0 3 0 . 0 0 0 0 . 0 0 3 0 . 0 0 3 0 . 0 0 0 0 . 0 0 0 0 . 0 0 0 0 . 0 0 0 0. 0 0 0 0 . 0 0 0 0. 0 0 0 0 . 0 0 0 0 . 0 0 0 0 . 0 0 0 0.000 0. 0 0 0 0. 0 0 0 0. 0 0 0 0 . 0 0 0 0. 0 0 0 0 . 0 0 0 0 . 0 0 0 0. 0 0 0 0. 0 0 0 0. 0 0 0 0 . 0 0 0 0 . 0 0 0 0.000 0 . 0 0 0 P O T A S S I U M - 0 . 0 0 0 0 . 0 0 3 0 . 0 0 0 0 . 0 0 0 0 . 0 0 0 0.000 0 . 0 0 0 0 . 0 0 0 0. 0 0 0 0 . 0 0 0 0.000 0.000 0.000 0.000 0 . 0 0 0 0.000 ?5 0 0 0 _ 0 . 0 0 0 • • • N U T R I E N T AMOUNTS IN DECAGRAMS BY SAMPLE S P E C I F I C A T I O N * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * WOOD**SECTIGN * * B A R K * * S E C T I O N * 162 3 64 NITROGEN - 1 3 . 9 5 8. 51 5 66 5.92 768 3.42 0.55 1 0 0.18 TOTAL 3 2 . 5 4 162 3 £4 NITROGEN - 1 3 . 5 2 1 1 . 0 2 566 7.09 7£8 5.55 10 TOTAL 0. 5 0 PHOSPHORUS-C A L C I U M MAGNESIUM -1.31 1 3 . 7 1 2.79 3. 71 1 2 . 9 2 1 .95 0.83 8.25 1.28 0.7 5 4.35 0.7 0 0. 14 0.61 0. 10 0.04 0.13 0.03 3.49 4 4 . 9 8 6.85 PHOSPHORUS-C A L C I U M MAGNESIUM -2.14 2 2 . 94 1.13 "~T79'7" 1 7 . 12 1.35 ~ 1T24"-1 2 . 0 3 0.90 "TToi o r n r 8.45 0.62 0.60 0.05 0.32 3 8 . 0 0 ""OTO 7 6 7 5 3 0.29 6 1 . 4 6 0.02 4. 05 0.17 1 6 . 6 1 P O T A S S I U M - 1 4 . 3 0 1 3 . 1 2 7. 73 4.44 0.65 0.20 3 7 . 4 4 P O T A S S I U M 5.24 4 . 7 3 3.45 2.7"8 0.2 5 BRANCH BRANCH * TWIGS- - F O L I AGE- CONES *• NITROGEN >1 0. 00 1-1/4 ( T O P ) ( M I D ) (LOW) TOTAL (TOP) (MID) " (LOW) TOTAL 0.00 0.00 0.00 0. 00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 " 0 7 0 0 0700"" T W I G S * F O L I A G E ~ TP. EE (TOP) ( M I D ) (LOW) TOTAL "TOTAL 0.00 0.00 0. 0 0 0.00 7 0 . 5 4 o o o b o o o o o o o o 0 .00 0.00 0.00 "'"0. 00 0.00 0.00 0. 0 0 "0.00 0.00 0.00 0. 0 0 0.00 10 . 0 2 1 0 6 . 4 3 1 0 . 9 0 " 54". 05 PHOSPHORUS-C A L C I U M MAGNESIUM -0.03 0. 03 0. 00 0. 0 0 0. 0 0 0. 0 0 0.00 0.03 0. 0 0 PO T A S S I U M - 0 . 0 3 0.00 0.00 0.00 0.00 0.00 0.00 0.00 q. qq 0.00 o. bo" 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 o.oo o.oo o. oo_ __o_ q_ 0.00 0 7 0 0 0.00 0.00 A P P E N D I X 7 N U T R I E N T CONCENTRATION AND N U T R I E N T D I S T R I B U T I O N W I T H I N 1 3 SAMPLED T R E E S ON THE HANEY SAMPLE PLOT S P E C I E S 1 - WESTERN HEMLOCK S P E C I E S 2 - WESTERN RED CEDAR S P E C I E S 3 - YELLOW CEDAR TREE NO. 8 SPECIES NO. 3 LENGTH ( Ml 19.0 OBH IMI 0.231 CROWN 01 AIM) 5.8 CROWN L. (M) 13.5 BUTT L. I Ml 5.5 •••PERCENT NUTRIENT CONTENT BY SAMPLE SPECIFICATION****************************************************** 8 * WOOO**SECTION * * BARK**S ECT I ON * 1 3_ 5 7 __9 1_0 1 3 5 7 9 10 NITROGEN -_PHO_SPifOgJAS-„ CALCIUM MAGNESIUM -POTASSIUM -" 0.050 _0, 005_ 6. 107 0.003 0. 052 0.062 _0.006 0.0 94 0.009 0.050 0.05 3 _o.oi?_ 0.091 0. 009 0.057 0. 063 J>_. 014 6. 089 0. 009 0. 069 0.085 0. 076 _0.012 0.01 0_ 0.107 0.095 0.007 0. 007 0.058 0.068 NITROGEN -_£.H° SPHORUS-CALCIUM MAGNESIUM -POTASSIUM -0.323 _P_._0_43_ 2. 570 0.048 0.280 0.270 0. 045_ 2.270" 0.050 0.266 0. 323 _0. 048 2.265 * 0.052 0. 375 0.4 01 0.055 1.705" 0.050 0. 325 0.433 _0o06 8_ 1.471 0.049 0. 350 0.533 0.061 T.2 56" o.04e 0.291 BRANCH BRANCH >1 1-1/4 -TWIGS- -FOL IAGE-(TOP ) (MID) ( LOW) (TOP) (MID) (LOW) CONES _* TWIGS + FOL I AGE-(TOP) (MIO) ( LOW) NITRQG;N !46 0. 243 0. 00 0 0. 000 0.000 0. OOP 0. 000 0.000 0.000 0.727 0. 752 0.728 PHOSPHORUS-CALCIUM _MAGN£S_IUM POTASSIUM 0.023 0.029 0. 747 0. 751 0. 023 0.028 0.000 0.00 0 0.00 0 0. 000 O.OOP 0. 000 0.000 0.000 0.000 0. 000 0. 000 0. 000 0. 000 0.000 0. 000 0.136 0.135 0.000 0.000 0.000 0.000 0.000 0. 000 0.000 0.000 0.000 0.000 0.000 0.000 0.000 0.079 1.142 _0.061 0.314" 0. 072 0.072 1. 196 1.407 0.054 0.066_ 0.237 0.2 81 ***NUTRIENT AMOUNTS IN DECAGRAMS BY SAMPLE SPECIFICATION************************************************************************** * WO0D**SECTI0N * * BARK** SECT I ON * 162 364 5 66 768 10 TOTAL 162 364 566 768 10 TOTAL NITROGEN - 3.00 PHOSPHORUS-CALCIUM MAGNESIUM -0.31 5.37 0.43 2. 38 0.37 3.30 0.33 6. 39 2.66 0.25 " 0.19 1.39 0.12 0.19 0.03 0.23 0.02 0.03 8.37 NITROGEN 0.00 0.04 0.00 1.29 13.48 1.15 PHOSPHORUS-CALCIUM MAGNESIUM -1.45 0.21 11. 86 0.24 _1_36_ 0.2"i" 10.43 0.23 1.05_ "0.15 5.76 0.15 0. 92_ "P."l4 3.49 0.11 POTASSIUM 2.71 2.19 1.36 0.91 0. 13 0.03 7.82 POTASSIUM 1. 34 1.47 1.02 0.74 0. 26_ "o; o"4 0= 88 0.03  21 0.16 5.20 0"."0"2"" "0.7 7" 0.38 32.60 0.01 0.77 0.09 4.87 _B_ANCH BRANCH « TWIGS- _ * *_->'i 1-1/4 "(TOP) ("MID") ("LOW) TOTAL --"-FOL I AGF * JCONES * TWI GS* FOL I AGE--: * (TOP)' (MIDI (LOW) "TOTAL" (TOP) (MIDI " (LOW) TOTAL" TREE "TOTAL" NITROGEN - 1. 03 3. 39 0.00 0.00 0,00 0.00 0.00 0.00 0. 00 0. 00 0.00 2.55 5. 87 2. 98 11 . 40 29.44 PHOSPHORUS- 0. 12 0. 39 0.00 0.00 0.00 0.00 0. 00 0. 00 0. 00 0. 00 0.00 0.28 0. 56 0. 30 i . 14 3. (i CALCIUM 3. 29 10. 28 0.00 0.00 0.00 0.00 0. 00 0. 00 0. 00 0. 00 0. 00 4.00 9. 33 5. 77 19. 10 78.95 MAGNESIUM - 0. 12 0. 38 0. 00 0. 00 0.00 0.00 0.00 0.00 0. 00 0. 00 0. 00 0.21 0.42 0. 27 0. 90 2.34 POTASSIUM - b." 60 1. 87 0.00 "o.oo 0. 00 0. 00 * 0.00 " " 0.00" "0. 00"" "" 0. oo "o.oo "1.10 1.85 " " 1 . 15 " 4. 10 "'" 19.26 f cv r TREE SPECIES LENGTH D8H CROWN CROWN BUTT NO. NO. (Ml (HI DIA(M) L.(M) L.(M) 50 1 17.1 0.160 4.9 14.3 2.9 • ••PERCENT NUTRI ENT CONTENT BY SAMPLE SPECIF i CATION ************* ****^ • WOOD**SECTION * * B ARK**SECT I ON * 1 3 5 7 9 10 1 3_ 5 7 ? lp_ NITR0G5N - 0. 043 0. 039 0.041 0. 049 0.055 0. 056 NITROGEN - 0. 166 0.210 0.233 0.2 82 0. 331 0. 352 PHOSPHORUS- 0. 003 0. 003 0. 01 0 0. 009 0.012 0. 013 PHOSPHORUS- 0.024 0.039 0.041 0. 054 Oc 063 0. 068 CALCIUM 0. 045 0. 043 0.042 0.046 0.048 0. 045 CALCIUM 0.412 0.502 0.295 0.314 0. 349 0. 3 37 MAGNESIUM - 3. 00 7 0. 003 0.008 0. 009 0.009 0. 009 MAGNESIUM - 0.016 0.023 0.025 0. 034 0. 045 0. 047 POTASSIUM - 0, 053 0. 055 0. 061 0. 064 0.0 54 0. 068 POTASSIUM - 0. 086 0.118 0. 184 0.195 0. 223 0. 2 63 BRA NCH BRANCH • —TWIGS-- • *- -FOLIAGE- • CONES * TWIGS*FOLIAGE — * >1 1- 1/4 ( TOP I (MIDI (LOW) (TOP) (MID) (Low) (TOP) (MID) ( LOW) NITROGEN - 0. 117 0. 123 0. 589 0.437 0.515 0. 759 0.709 0.735 0.000 •6-W174 0. 573 0. bj&-PHOSPHORUS- 0. 019 0. 024 0. 080 0. 082 0.071 0. 128 0. 138 0.161 0.000 0.4 fT-*^ s0.396 >. -3C403 CALCIUM 0. 17 3 0. 179 0.305 0. 313 0.275 0. 432 0.479 0. 542 0. 0 00 0.368 0.409 MAGNESIUM - 0. 015 0. 021 0.05 5 0. 050 0.050 0. 105 0. 113 0. 121 0. 000 •m«085 POTASSIUM - 0. 072 0. 083 0.195 0. 168 0. 185 0. 334 0. 289 0.303 0.000 fl<?64 0.229 0?2>4» •••NUTRIENT AMOUNTS IN DECAGRAMS BY SAMPLE SPECIF] -W0OD*«SECTI0N- -BARK**SECTION-162 364 5 66 768 10 TOTAL 162 364 566 768 10 TOTAL NITROGEN - l.OJ 0.72 PHOSPHORUS-CALCIUM MAGNESIUM -0.21 0.16 1.24 0.81 0.19 0.14 0.60 0.13 0.53 0.11 0.34 0.08 0.07 0.02 0.31 0.07 0.06 0.01 0.03 2.85 NITROGEN 0.01 0.02 0.00 0. 59 3.04 Oo 51 PHOSPHORUS-CALCIUM MAGNESIUM -0. 83 0. 14 2.01 0. 09 0.86 0.16 1.55 0.09 0.64 0.12 0.76 0.07 0.40 0.08 0.43 0.05 0.10 0.11 "070_ """D'70"2~ 0„10 0.10 OoOl 0.01 2.94 ""0753" 4. 96 0.33 POTASSIUM 1.45 1.04 0. 33 0.3 3 0.08 0.03 3.82 POTASSIUM - 0.45 0.59 0.47 0.27 Oo 07 0.08 1.93 BRANCH BRANCH *- -TWIGS->1 1-1/4 (TOP) (MID) (LOW I TOTAL -FOLIAGE-(TOP) (MID) (LOW) CONES TOTA'L THIGS*FOLI AGE— (TOP) "" (MID) u o w r TOTAL" TR EE TOTAL" NITROGEN - 0. 16 2.12 0. 88 1.62 0. 93 3.43 0.61 1.49 0. 74 2.83 0.00 1.49 3.11 1.66 "07*59 0.2 9 2.16 1.04 0.42 0.21 1.23 0.64 6.26 1.10 4. 00 0.80 2. 43 14.34 PHOSPHORUS-CALCIUM _MAGNES I_UM„; POTASS"!UM -0.03 0.42 0. 24 3. 08 0.02 0. 36 6. 10 1. 38 0. 12 C. 46 0.J38 0.29" 0.30 0. 13 1.16 0.49 0. IB 0.09 "0.62 "0.33" 0.55 0.10 2.11 0.35 0.36 0. 08 1.25 0.2 7" 0.29 0.16 1.01 0.54 0.24 0^12 0.61 0.30 0. 55 0. 00 1.89 0.00 0.44 0.00 T . 18" "0700" 0.22 0.80 0.17 '0.56* 2.66 15.32 2.03 "9.6'5 1 3 5 7 9 10 1 3 5 7 9 10 NITROGEN -PHOSPHURUS-0.040 0. 015 0.045 0.007 0.035 0.006 0.057 0.020 0.072 0.018 0. 066 0.014 NITROGEN -PHOSPHORUS-0. 171 0. 027 0.130 0.023 0.151 0. 025 0.206 0.027 0.288 0. 048 0. 357 0. 062 CALCIU* MAGNESIUM -POTASSIUM -0.041 0. 010 0. 08 i 0. 053 0.013 0.066 0. 076 0.019 0.069 0. 064 0. 016 0. 074 0.050 0.012 0.081 0. 040 0. O i l 0. 086 CALCIUM MAGNESIUM -POTASSIUM -0.603 0. 019 0, 096 0.624 0.020 0.124 0.482 0.019 0.085 07496 0.019 0.109 0.270 0.037 0. 201 07242" 0.045 0.22e BRANCH 3RANCH -TWIGS-- * -FOLIAGE- CONES * T WI GS.FOL I AGE * >1 1-1/4 (TOP) (MIO) ( LOW) (TOP) (MID) (LOW) ( TOP) (MID) ( LOW) NITROGEN - 0. 114 0. 153 0.484 0. 363 0.323 0. 776 0. 766 0.759 0.000 •6^30 0. 564 o.>w PHOSPHIIRUS-CALCIUM MAGNESIUM -0. 01 7 0. 20 3 0.019 0.024 0. 135 0.022 0.067 0.265 0. 066 0. 052 0. 259 0. 053 0.055 0.227 0.044 0. 100 0. 486 0. 105 0. 114 0. 538 0. 117 0. 155 0.478 0. 133 0. 000 0.000 0. 000 0. 4 2>«^ 0. 376 0.08>-' vO.409, •"0.085 ^-Tj.407 0.352 ^0^088 POTASSIUM - 0. 034 0.093 0.246 0. 162 0.149 0.372 0. 309 0. 302 0.000 0. 236 07?>6» TREE NO. 54 SPECIES NO. 1 LENGTH (M) 22.4 DBH (M ) 0.394 CROWN 01 A( M) 4.7 CROWN L.IMI 14. B BUTT L. (M) 7.6 • ••PERCENT NUTRIENT CONTENT BY SAMPLE SPECIF I CAT I ON ****************************************************************•**»»» -WOOD**SECTION * -BARK*•SECT ION-4 • ••NUTRIENT AMOUNTS IN DECAGRAMS BY SAMPLE SPECIFICATI ON **************************** ********»*~**»~*~*~*^ -WOOD**SECTION * 162 3 64 5 66 768 9 10 TOTAL 162 364 566 768 9 10 TOTAL NITROGEN - 6.34 5.13 4. 31 2.93 0.22 0.06 19.48 NITROGEN - 4.71 3.67 4.11 2.27 0. 72 0. 14 15. 61 PHOSPHORUS-CALCIUM MAGNESIUM -1.77 7 .23 1.75 0.84 8.44 2.02 1. 33 7.29 1. 80 0.86 2.59 0.64 0.05 0.15 0. 04 0.01 0.04 0.01 4. 86 25. 79 6.26 PHOSPHORUS-CALCIUM MAGNESIUM -0.77 19. 20 0.60 " 0763" 14.43 0.50 0.60 0.34 11.25 3.53 0.43 0.26 0 .T2 0 ,67 0, 09 "0 .02 0. 10 0. 02 2. 49 49. 18 1 .91 POTASSIUM - 11.47 8.53 7.46 3.52 0.24 0.08 31.29 POTASSIUM - 3.44 2.73 2.24 1.43 0. 50 0 .09 10.43 BRANCH BRANCH *• •—TWIGS- -FOLIAGE * CONES *- TWIGS+FOLIAGE— TREE NITROGEN >1 4. 05 1-1/4 (TOPI (MID) (LOW) TOTAL (TUP) (MID)(LOW I " TOTAL" "(TOP) (MID) "" (LOW) "TOTAL" 8.24 1. 02 2.03 1.13 4.18 PHOSPHORUS-CALCIUM _MAGNES IUM _-POT ASSIUM -4.81 11.64 5. 39 21. 83 0. 62 1.23 7. 35 9.66 0_. 66 1. 14 1. 99 5.14 0. 14 0. 56 0. 14 0. 52 0.00 5.83 13.67 6. 52 0.29 0. 19 1.45 0.79 0.30 0.15 0.91 0. 52 0.62 0.62 2.80 3. 01 0.59 0.65 1.95 " 2.30 1.73 8. 17 1.7 8 ' 4.69' 1.10 3. 39 0.94 2.15 3.45 14.58 3.37 "9.14 0.00 0. 00 0.00 0.00 0.76 3.57 0.79 2.82 2. 03 9.62 2.07 5.60 T 7 2 9 " 4.1 9 26. 01 ros"" TOTAL" 73.41 4. 17.38 1.10 3.96 2.67 11.09 13.2T" 109.36 13.93 60.93 TREE SPECIES LENGTH OBH CROWN CROWN BUTT NO. NO. (Ml CM) DIA(M) L. (M) L.(M) 60 2 20.4 0.5 33 0.0. 14.3 6.1 • ••PERCENT NUTRIENT CONTENT * : 1 3 BY SAMPLE SPECIFICATION******************************************************************************* — WOOD**SECT ION * * BARK**SECTI ON * 5 7 9 1 0 1 3 5 7 9 1 0 NITROGEN -PHOSPHORUS-CALCIUM MAGNESIUM -POTASSIUM -0. 057 _0_._003_ 0.094 0. 005 0.056 0.081 0. 009 0.095 0.005 0.050 0.048 0.004 0. 129 0.007 0.050 0.072 _C_006_ 6. 1 24 0.003 0. 047 0.051 0.005 " 0. 119 0.009 0. 044 0 . 053 g. oo 5_ "o. n~8 0. 009 0. 04 4 NITROGEN -PHOSPHORUS-CALCIUM MAGNESIUM -POTASSIUM -0.253 0.028_ "~0.'546 0. 018 0.142 0.229 _0.022_ 0.592 ' 0.017 0.088 0. 203 0.019_ 0,590 0. 018 0.07e 0.063 0.004 "0 .11 3 0.007 0.043 0.063 0. 081 0. 004 0.005 0.116 6.099" 0.009 0.007 0.059 0.040 BRANCH BRANCH >1 1 - 1 / 4 (TOPI rJwiGS-rr * CMIDT (LC'KI ^FOL IAGE-(M ID I (LOW I CONES * TWIGS+FOLIAGE * "(TOP) ""'" (MID) (LUW) NITROGEN 0. OOP . 0. OOP 0.000 o.oqo o.ooo o.ooo 0.000 0.000 0. 000 0.000 0. 000 0.000 PHOSPHORUS-CALCIUM MAGNESIUM -POTASSIUM -0. 003 0. 003 0. 033 0. 003 0.000 0.003 0.003 0.000 0. 000 O . O P O 0.000 6.000 0. 003 0.000 0.000 0.000 0. 00 0 0. 000 0. 00 0 0. 000 0. 000 0. 000 0.000 0.000 0.000 0.000 0. 000 0.000 0.000 0.000 0. 000 0.000 0.000 0.000 0.000 0.000 0.000 0.000 0.000 0.000 0.000 0.000 0. 000 0. 000^ 0.000" • ••NUTRIENT AMOUNTS IN DECAGRAMS BY SAMPLE SPEC I F I CAT ION**************************** ************"************"**************** ****** « . . . WOOD**SECT ION * * BARK**SECTI ON * 162 364 566 768 9 10 TOTAL 162 364 566 768 9 10 TOTAL NITROGEN - 14.51 11.41 8.67 3.81 0.70 0.25 39. 36 NITROGEN - 6.61 3.91 1.26 0.00 0.00 0.00 11.78 PHOSPHORUS- 1.65 1.04 0.73 0.3 7 0.08 0.03 3.89 PHOSPHURUS- 0. 68 0.37 0. 11 0.00 oroo- o."co- T.16 CALCIUM 19.93 19. 80 18.23 7. 50 1.63 0.57 67. 63 CALCIUM 15.59 10.70 3.34 0.00 0. 00 coo 29.63 MAGNESIUM - 1.13 1 .09 1.10 0.52 0.12 0.04 3. 99 MAGNESIUM - 0.48 0.32 0.12 0.00 0,00 0.00 0.92 POTASSIUM - 11.21 8.37 7.03 2.84 0.61 0.21 30.77 POTASSIUM - 3. 16 1.51 0. 58 0.00 0. 00 0. 00 5.25 BRANCH BRANCH *- TWIGS * * — FOLIAGE * CONES * TWIGS + FOL IAGE — TF EE >1 1-1/4 {TOPI ( MIDI (LOW I TOTAL (TOP I (MIDI ( LOW) TOTAL (TOP I" (MIO) "(LOW) TOTAL "TOTAL " NITROGEN - 0.00 0.00 0.00 0.00 0. 00 0.00 0. 00 0. 00 0. 00 0. 00 0.00 0.00 0.00 0.00 0.00 51. 14 PHOSPHORUS- 0. 00 0. 00 O.PO 0.00 0. 00 . 0.00 0.00 0.00 0.00 0. 00 0. 00 0.00 0.00 6.00 0.00 5.03 CALCIUM 0 . 03 0. 00 0. 00 0. 00 0. 00 ' 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0 . 00 97.25 MAGNESIUM - 0. 00 0. 00 0.00 0.00 0. 00 0.00 0.00 0. 00 0.00 0. 00 0.00 0.00 0.00 0.00 0.03 . 4.91 POTASSIUM - 0. 30 0 . 00 o. 66" 0.00" 0. 00 " " 0.00 6.00" 6.66 0. 00 "6. 00 " "o; oo" 0.00 0. 00 0.00 0. 00 3 6:. 02 TREE NO. 6) SPECIES NO. 3 LENGTH ( M ) 27.2 DBH < M I 0.401 CROWN OIA(M) 8.3 CROWN BUTT L.(H) L.(H) 15.4 11.8 •••PERCENT NUTRIENT CONTENT BY SAMPLE SPECIFICATION************************************************ • WOOD**SECT ION * * BARK**S ECT I ON * 1 3 5 7 9 10 1 3 5 7 9 10 NITROGEN -PHOSPHORUS-CALCIUM MAGNESIUM -POTASSIUM -0.053 0. 009 0.049 0.005 0. 126 0.123 0.013 0.012 0.04 9 0.046 0.06 0 0.005 0. 069 0. 009 0.073 0.007 0. 072 0. 012 0. 119 0.014 0. 046 0. 053 0.006 0. 027 0.111 0.135 0.017 0.014 0.068 0.056 NITROGEN -PHOSPHORUS-CALCIUM MAGNESIUM -POTASSIUM -0.255 0. 045 0.250 0.032 3.509 2.182 0.041 0.039 0.168 0.113 0.268 0, 043 2. 326 0.050 0.212 0.339 0. 064_ T788 2" 0.055 0.241 0. 365 0^070 1.477 0.069 0.220 0.429 _(K0 31_ 1.490 0.080 0.279 BRANCH BRANCH -TWIGS- -FOLIAGE->1 1-1/4 (TOP ) ( MIU) (LOW) (TOP) (MIO) (LOW) CONES J * r — T J i ! GS+FOL L A G i - : - * (TOP] ~ (MTOT ( LOW)" _NJTRGGEN ^ _ PHOSPHORUS-CALCIUM MAGNES IUM_ Z_ POTASSIUM" -0. 107 0. 013 0. 523 0. 029 0.062 0.250 0.000 0.000 0.000 0. 000 0. 000 0.000 0.000 0.846 0.703 0.841 0.031 0. 305 _9_'_°43_ 0. 154 0.000 0. 00 0 0.000 0.000 0.000 0. 000 0.000 0.000 0.000 0.000 _q._o_oo_ "o.ooo 0.000 0. 000 0. 000 0.000 0. 000 0. 000 0. 000 0.000 0.000 0.000 0.000 0.000 0. 000 0.000 0.000 0.000" 0. 094 1.222 0.092 0. 091 1. 313 0.091 0.476 0. 354 0.091 1.447 0.094 0.476 • .•NUTRIENT AMOUNTS IN DECAGRAMS BY SAMPLE SPECIFICATION*****************************************" -WPOU** SECTION- -BARK**SECTION-162 364 566 768 9 10 TOTAL 162 364 566 768 9 10 TOTAL NITROGEN - 10.63 7.95 7.45 3.98 0.68 0.19 30.84 NITROGEN - 2.45 3.32 2.83 2.96 0. 51 0. 34 12.40 PHOSPHORUS-CALCIUM MAGNESIUM -1.33 26.01 2.64 0.67 17.66 1.94 0. 31 9.94 1.16 0.45 4.59 0.64 0.06 1.03 0. 16 0.03 0.35 0.04 3.41 59. 53 6.57 PHOSPHORUS-CALCIUM MAGNESIUM -0.37 2 7. 60 0.39 0.48 28.85 0.57. 0.49 19.57 0.49 0.56 14.11 0.52 ""O.T'O"" 2.07 0. 10 '• 0.06 ' 1.19 0.06 '2 .07 93.39 2.13 POTASSIUM - 9.91 6.73 4.25 2.67 0.63 0.15 24. 33 POTASSIUM - 1.36 2.08 2.11 1.94 0.31 0.22 8.01 BRANCH BRANCH * - TW I GS- •* - F O L I AGE CONES *- TWIGS*:FOLIAGE « TREE NITROGEN >1 5. 21 1-1/4 (TOP) (MIO) (LOW) TOTAL 5. 93 PHOSPHORUS- 0.87 CALCIUM - 25.31 MAGNESIUM -__ 1.40 ' POTASS IUM - ' "3.05' 0. 73 19. 09 1. 02 "3.65 0. 00 0. 00 0.00 0. 00 0. 00 " 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0. 00 "" 0. 00 ( TOP) (MID) (LOW) " TOTAL "(TOP)' (MID.) '" (LOW) 0.00 0.00 0.00 0.00 0. 00 0.00 4.57 12. 70 0.00 0. 00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0. 00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 '" 0.00 0.51 6.60 0. 50 2.57 15.67 2.04 nre~ 29.28 21.85 2.03 1.42 7.90 7.19 TOTAL 32.94 s~7T2" 57.73 3.95 17.66 TOTAL 87.33 11.00" 255.59 15.OS 56.70 9 L TREE NO. 65 SPECIES NO. 1 LENGTH ( M) 23. 5 DBH ( M) 0. 343 CROWN DIA(M) 6.1 CROWN BUTT L.IM) L.(M) 18.5 4.9 •••PERCENT NUTRIENT CONTENT BY SAMPLE SPECIFICATION**************************************************** • WOOD**SECT ION * * BAPK**SECTI ON * 1 3 5 7 9 10 1 3 5 7 9 10 NITROGEN -PHOSPHORUS-CALCIUM MAGNESIUM -POTASSIUM -• . 045 J2- 01 2_ 0.051 0.00? 0. 06 0 0.042 JJ.-P14_ 0.055 0. Oil 0.035 0.045 _0. 011_ 0.04 7 0.010 0.054 0.047 0.008 0.041 0. 008 0. 061 0.052 0.014 0.045 0.009 0. 069 0. 067 _0. 01__ 0. 043 0. 009 0. 079 NITROGEN -_P>HOSPHORUS^ _ C AlC IUM '"-MAGNESIUM -POTASSIUM -0.151 0.158 0.036 0.020 0.3 f6" 0.423"" 0.013 0.011 0.062 0.069 0. 167 0.027_ 0. 464 0.015 0. 108 0.201 0.032 0.284 0.017 0.137 0. 275 0.311 0. 041 0. 044_ 0.383 "0.3 63 0.024 0.026 0.154 0.180 _BRA'CH BRANCH_ 1-1/4" >1 (TOP) -TWIGSr ( MID) ( LOW) * — --FOL IAGE-(TOPf (MID) (LOW ) CONES * TWIGS+FOL I AGE * "(TOP) (MID) " (LOW)" NITROGEN - 0. 091 0. 161 0.479 0.423 0.423 0. 856 0.814 0.814 0.000 PHOSPHnRUS-CALCIUM MAGNES IUM -POTASSIUM -0. 014 0. 173' 0. 013 0. 059 0.025 0.249 0. 020 0.094 0. 06 5 0.42 3 0.04 6 0. 054 0.419 0.042 0.054 0.419 0.042 0. 106 0. 472 0. 085 0. 124 0. 510 0. 107 0.220 0.219 0.219 0. 467 0.43 2 0.124 0. 510 0. 107 0.432 0.000 0.46 0.000 0.447 _0.000 0.06 0.000" ***NUTPIE~NT AMOUNTS IN DECAGRAMS. BY SAMPLE SPECIFICAT I ON****************************A********************************************* * WOOD**SECTION * * BARK**S EC Tl ON — .NITROGEN PHOSPHORUS-CALCIUM MAGNESIUM -1C2 5.99 1.77 7.3) 1.4J POTASSIUM - 10.01 3C4 4.52 1.32 5. 35 1.08 7.24 5 66 2.72 7t8 1.30 0.28 10 0.12 TOTAL 14. 93 NITROGEN 1C2 4.94 3E4 3.64 5£6 2.30 0.58 2.63 0.52 3.43 0.2 9 1.12 0.22 0.07 0.24 0.05 0.03 0.08 0.02 4.07 16.73 3.28 PHOSPHORUS-CALCIUM MAGNESIUM -0. 89 11.80 0.38 0.53 9.94 0.28 0.3 7 4.67 0.20 7£8 9 1.55 0^33_ 0.24 0. 05 2.17 0.46 0.14 0.03 10 0. 12 "'0.0'2 0. 15 0.01 "TOTAL 12. 89 ro"9~ 29. 18 1.04 1.72 0.37 0.14 22.91 POTASSIUM - 2.09 1.98 1. 53 0.94 0. 13 0.07 6.60 BRANCH BRANCH *->1 1-1/4 (TOP) NITROGEN - 7.32 11.85 2.06 •— TW IGS "(MT6) (LOwf .* * FOLIAGE-TOTAL (TOP) (MID) (LOW) TOTAL 3. 09 2.41 CONES * TW I GS+ FOL I AGE— ""( TOP)"" ( MID) "" ( LOW) 7.56 6.33 10. 34 8.79 25.46 0.00 PHOSPHURUS- 1.10 CALCIUM - 14.34 MAGNESIUM- 1.09 POTASSIUM - 4.77 1. 84 18. 2 5 1. 43 6. 3 9 0. 23 1. 82 0. 20 "0. 95 " 0.39 0.31 3.06 2.39 0.31 0.24_ I'.bO 1.25 0.98 0.79 7.26 3.49 0.74 0.63 3.79 3.46 1.58 6.48 1^36 5.49 1. 34 5.51 JL. 15 4.67 3. 70 15.48 3.14 T3." 6 r 0. 00 0.00 0.00 "0. 00 8.39 13.43 1.06 5.31 0.83 "4". 40" 11.20 T797 1.65 9.54 7.90 1.66 1.39 "7.09" 5*. 91"' "TOTAL" 33. 02 4.68 22.74 3.88 17.40" TREE TOTAL" 80.02 13.79 101.25 10.76 •"58.78 " TREE NO. 69 SPECIES NO. 1 LENGTH t M ) 18.9 OBH (Ml 0.221 CROHN OIA(M) 3.5 CROWN BUTT L.(MI L.(M) 14.0 4.9 • ••PERCENT NUTRIENT CONTENT BY SAMPLE SPECIF I CAT ION*********************************************************************»***»*•»** -WOOD**SECTION- --BARK**SECTI ON-1 3 5 7 9 10 NITROGEN - 0. 042 J . 040 0.047 0. 045 0. 060 0. 06 9 PHOSPHORyJ- 0. 01J 0. 013 0.012 0. 013 0. 014 0. 014 CALCIUM 0. 03 9 0. 049 6. 042 0. 042 0. 038 0. "038 MAGNESIUM - 0. 00 3 0. 013 0.012 0. 012 0. 010 0. O i l POTASSIUM - 0. 06 0 0. 054 0. 055 0. 070 0. 074 0. 073 10 NITROGEN -J>_HOSPHORUS-CALCIUM MAGNESIUM -POTASSIUM -0.150 0. 0 2 4_ 0.2 33 " 0. 019 0. 100 0.185 _0._028_ " 0.222 0.028 0.158 0.201 0. 0 3 3_ "6". 2 28 0.028 0. 179 0.245 0.042_ 0.191' 0.037 0.201 0.288 0.322 0.048_ _0.053 "0.204 "" 0.217 0.038 0.040 0.217 0.212 BRANCH >1 _3R_ANCH_ 1-1/4 -TWIGS- -FOL I AGE-( TOP ) ( M I D I I LOW) (TOP ) (MID) (LOW) CONES •r-rTWIGS*FO_L I AGE — (TO PI (MIDI ( LOW) NITROGEN 0. 105 0. 159 0.472 0. 564 0.345 0. 793 0. 809 0.790 0.000 PHOSPHORUS-CALCIUM _MAGNES IUM_r POTASSIUM -0.02 0. 0. 109 _0.022_ ,074 0. 024 0. 143 _°«027_ 0.100 0.074 0.199 _0. 0_58_ 0.183 0. 072 0. 158 0. 047 0.046 0. 346 0.035 0.169 0.169 0. 122 0. 331 0. 106 0. 134 0.412 0. 128 0.336 0.315 0. 162 0.509 0.108 0.408 •••NUTRIENT AMOUNTS IN OECAGRAMS BY SAMPLE SPEC IF ICAT I ON**************************** **«*'*** + *****"*T'*l>*"********«^irV*>*"i^* ; « WOO0**SECTI0N * * BAR K**SECT ION * 1C2 3C4 . 5&6 7C8 9 10 TOTAL 1&2 3£4 5C6 7E8 9 10 TOTAL _NITROGEN _ - 1.72 1_.38 1.08 0.72 0.16 0.06 5.11 NITROGEN- 1.91 1.49 1.27 0.60 0. 20 0.10 5.77 PHOSPHORUS- 0.47 0.38 O.T§ 0.18 07o4 0.01 1.37 PHOSPHORUS- 0.30 0.24 "0".2'1" 0.13" 0.03 """0.02 0.93 CALCIUM - 1.86 1.44 0.99 0.55 0.10 0.03 4.97 CALCIUM - 2. 59 1.73 1.20 0.59 0.14 0.07 6.32 MAGNESIUM - 0.45 0. 39 0.27 0.15 0.02 0.01 1.29 MAGNESIUM - 0.27 0.21 0.18 0.11 0.03 0.01 0. 81 POTASSIUM - 2.33 1.72 1.47 0.99 0.19 0.06 6. 81 POTASSIUM - 1747 1730 1708 b"7"""3~ 0TT5 0"70"6 4"77b" BRANCH BRANCH * TWIGS >1 1-1/4 (TOO) (MIDI * * ( LOW I TOTAL ( TOP) — FOLIAGE--—-(MIO)"" fLOW) TOTAL JZONES *- TWI GS+ FOL I AGE — (TOP I (MIDI (LOW I' TOTAL" TREE TOTAL" NITROGEN -PHOSPHORUS-CALCIUM MAGNESIUM -POTASSIUM -0. 13 3. 93 0.99 1 .35 0. 24 2.59 4.92 4.69 0.95 10.56 0. 00 5.91 6. 04 0. 02 0.60 0. 13 3.59 0. 03 0. 66 '"o".""09"" 2.50' 0. 16 0,42 0, 12 0. 38 0.17 0.03 0.38 0. 24 0.11 0.02 0.41 " C. 12 0.36 0.75 1.04 2. 05 0.26 0.66 0.91" " 2. 08 0.78 0.19 2. 39 0.61 0.74 0.13 T.82" 0.49" 1.73 5.05 1.53 4.40" 0.00 0.00 0. 00 "0.00 0.91 2.47 0.78 2.47 0.95 2.77 0.85 2.23 1.19 0723"" 0. 85 0. 15 0.61 13.14 6.09 1. 79 5. 31 5.01 21.13 4. 58 19. 40 TREE NO. 72 SPECIES NO. 1 LEN GTH ( Ml 24.2 D8H (HI 0. 396 CROWN 01A(Ml 6.6 CROWN BUTT L.(M) L.(H) 17.3 6.9 •••PERCENT NUTRIENT CONTENT BY SAMPLE SPECIFICATION******************************************************************************* * W0O0**SECT ION * * BARK**SECTI ON * _1 _ J 5 7 9 10 1 3 5 _7 9 10 NITROGEN -PHOSPHCRUS-0. 046 0.._030 0.043 . 0. 0 2_2_ CALCIUM MAGNESIUM POTASSIUM 0.053 0. 014 0. 104 0.057 0. 014 0.153 0. 042 _0.-pl2 0. 042 0.010 0. 03 4 0.042 _0. 006_ 0.048 0. 009 0. 064 0.044 o- o n_ 0.047 0.009 0.073 0. 060 j0.j011_ 0. 038 0. 009 0. 05 5 NITROGEN -PHOSPHORUS-CALCIUM MAGNESIUM -POTASSIUM -0. 159 0.026_ "67332" 0. 015 0. 062 0.118 _0._0_19_ 0.327 ' 0.013 0.083 0. 141 0.023 0.301" 0. 014 0. 109 0. 160 0.031 "6.2 21" 0.015 0.114 0.211 0.302 0. 040 0.045 "0.256"" " 0.255' 0. 019 0. 029 0.165 0.221 BRA NCH BRAN C H_ 1-1/4 NITROGEN >1 0. C77 0. 132 _*r-_- ~~I WIGS- -(TO PI (MI Of" ( LOW I 0.718 0.484 0.444 (TOP) 0. 821 -FOLIAGE-" "(MID) 0. 824 (LOW) 0.820 CONES 0. 000 * TWIGS*FOL I AGE * "(TOP)" (MID) " (LOW) PHOSPHORUS-CALCIUM MAGNESIUM 0. 012 0. 134 0. 014 0.025 0.173 0. 020 0. 075 0.242 0.04 5 0. 068 0.25i 0. 049 0.073 0.246 0.053 0. 112 0. 317 0.072 0.121 0.438 0. 037 0.144 0.430 0.082 0.000 0. 000 0. 000 POTASSIUM - 0.030 0.033 0.131 0.168 0.220 0.431 0.297 0.311 0.000 •••NUTRIENT AMOUNTS IN DECAGRAMS 3Y SAMPLE SPECIFICATION************************************************************************** * WC;OD**SECTION * * BARK*»SECT I ON NITROGEN -1C2 7.70 3£4 4.94 5C6 3.35 7£8 1.52 0.28 10 0.08 TOTAL 17. 86 NITROGEN U 2 7.23 3C4 4.22 5£6 3.24 PHOSPHORUS-CALCIUM MAGNESIUM -4.45 9.45 2.41 1.96 5.76 1.42 0.71 3.65 0.79 0.30 1.68 0.32 0.07 0.2 9 0.06 0.01 0.05 0.01 7.50 20. 88 5. 01 PHOSPHORUS-CALCIUM MAGNESIUM -1 . 1 6 17. 18 0.73 0.69 10.21 0.43 0.5 8 5.60 0.31 7£8 9 2.08 0.38_ 0.40 0. 07 2.67 0.46 0.19 0.03 T P " "TOTAT POTASSIUM - 22.04 13. 73 5,96 2.42 0.46 0.07 44.67 POTASSIUM 3. 80 3.12 2.39 1.56 0, 30 BRANCH BRANCH *- -TWIGS-NITROGEN ->1 4. 31 1-1/4 (TOP) (MIDI (LOWI TOTAL ( TOP) ^FOLIAGE---— ( MIO')" "I LOW I 9.81 4. 09 5.38 3. 19 12.66 7.23 17. 55 11. 73 TOTAL 36. 51 CCNES *- TWIGS•FOLI AGE--(TOP)"~"(MID) (LOW) 0.00 11.32 1.42 4.17 0.89 "4.82" 14.92 2.56 22.93 ~T73"4" 12.11 7.92 2.39 1.56 " "8. 19 6.03 ' TOTAL" 49. 17 TREE "TOTAL" 98.45 —irvr 24. 20 4. 84 "19.04 " 20.30 101.62 13.65 ~ 8 5.41" PHOSPHORUS-CALCIUM _MAGNES IUM__-POTASSTUM -0. 69 7. 49 0.81 4. 49 1.35 12. 81 _1._43_ 5. 93 0, 43 1. 38 0. 26 1. 03" 0.75 2. 78 0.54 1 .86 0. 52 1. 77 0. 38 1. 58" 1.71 5.93 1.18 4.4 8 0.99 2.79 0. 64-_ '3.79 " 2.58 9.33 1. 85_ "6.3 3 2.06 6.14 1.18 4.44 "T76"r 18.27 3.66 TV. 5 6" "1570 0 0 . 0 0 0 . 0 0 0700 TREE SPECIES LENGTH DBH CROWN NO. NO. ( M) (HI DIA(M) 76 3 14.8 0.180 5.1 CROWN BUTT L.(M) L.IM) 9.1 5.7 •••PERCENT NUTRIENT CONTENT BY SAMPLE SPECIFICATION*************************************************** • WOOD** SECT I ON * * BARK**SECT ION * 1 3 5 7 9 10 1 3 5 7 9 10 NITROGEN -__PH OS PHORUS-CALCIUM "AGNESIUM -POTASSIUM -0.043 _0.0.09_ 0. 153 0.011 0. 045 0.053 J>_.JH>9_ 0. 149 0.008 0.050 0.056 0.036 0. 121 0.009 0.044 0. 057 0.006 0.071 0.009 0. 064 0. 011 NITROGEN -PHOSPHORUS-0.261 0. 043 0. 121 0.008 0. 053 0.138 0.008 0.054 0. 140 0. 007 0.054 CALCIUM MAGNESIUM POTASSIUM 2.934 0. 026 0. 162 0.289 0. 048 2.995 0.041 0.173 0.226 0. 047' 2.839 0.032 0. 159 0.276 0.075 2.A43 0.036 0.223 0.348 0.071 0.313 _0.079 2/537 1/9 59 0.047 0.044 0. 329 0.306 BRANCH BRANCH -TWIGS-- * FOLIAGE- CONES >1 1-1/4 (TOP) (MID) ( LOW) (TOP) (MID) (LOW) * TW!GS*FOLIAG_E---*_ (LOW)" ( TOP) (MID) NITROGEN 0.113 0.190 0.000 0.000 0.000 0. 000 0.000 0.000 0.000 0.766 0.675 0.766 PHOSPHORUS-CALCIUM VAGNEJJUM^-" POTASSIUM -0. 316 0. 573 _0.013_ 0. 363 0.025 3. 882 0.023_ 0. 14 7 0.030 0. 30 0 0.000 0.003 0. 000 0.000 0. 000 0.000 0.000 0.000 0.003 0.000 0. 000 0. 000 0. 000 0.000 0.000 0.000 0.000 0.000 0.000 o^ oip o o. o oq_ o.'boo 0.000 0.091 0.082 0.088 0.000 1.180 1. 1 18 1.346 _0.000 0.059 0.05 8 0.060_ 0.000 "b"/"296" 0/310 0. 338 • ••NUTRIENT AMOUNTS IN DECAGRAMS BY SAMPLE SPECIF 1 -WOOD*«SECTION- -BARK**SECTION-NITROGEN PHOSPHORUS-CALCIUM MAGNESIUM -1 £2 3£4 5 £6 7£8 9 1 0 TOTAL 1£2 3£4 5 £6 7£8 9 10 TOTAL 1.53 1. 04 0. 74 0.39 0.06 0.03 3.79 NITROGEN - 0.47 0.33 0.30 0.3 5 0, 09 0.00 1. 55 0.25 4.42 0.26 0.14 2.44 0.15 0.03 1.57 0.11 0.05 0.79 0.05 0.01 0.12 0.01 0.01 0.07 0.00 0. 55 9.42 0.59 PHOSPHORUS-CALCIUM MAGNESIUM -0.08 5. 04 0.06 0.06 3.79 0.05 0.07 3.41 0.04 "0/09 3.2 3 0.05 0. 02 0. 76 0. 01 o.'o'o 0.00 0. 00 0. 33 16.23 0.21 1.34 0.85 0.63 0.32 0.05 0.03 3.25 POTASSIUM - 0.29 0.22 0.23 0.33 0.10 0.00 1.16 . POTASSIUM 8RANCH BRANCH *- -TWIGS- -FOL I AGE- CONE S *- -TWIGS* FOLI AGE- TREE >1 1-1/4 (TOP ) "(MID) ( LOW) TOTAL (TOP) (MIDI (LOW) TOTAL (TOP) (MID) "(LOW) TOTAL TOTAL NITROGEN - 0. 55 2. 11 0. 00 0.00 0.00 0.00 0.00 0.00 0.00 0. 00 0. 00 1.84 3. 51 7. 58 12.93 20.93 PHOSPHORUS- 0. 03 0.29 0. 00 0. 00 0. 00 0.00 0. 00 0. 00 0.00 0.00 0.00 0.22 0.43 0. 87 1.52 2.76 CALCIUM 2.63 9. 79 0.03 0. 00 0. 00 0.00 0. 00 0. 00 0.00 0.00 0.00 2.83 5.81 13.32 21.97 60.09 MAGNESIUM - 0. 06 0. 2 5 0. 00 0.00 0. 00 0.00 0.00 0.00 0.00 0.00 0.00 0.14 0. 30 0. 60 1.04 2. 15 POTASSIUM - 0.32 1.63 0. 00 0.00 " "b.'oo 0.00' "' o. bo " "0. 00 0.00" "0.00" "0.00 0.71 1.61 3. 34 5.67 12.03 TREE NO. 102 SPECIES NO. 3 LENGTH ( Ml 25. 5 DBH (Ml 0.602 CROWN OIAI Ml 9.1 CROWN L.IMI 15.0 BUTT L.IMI 10.5 • ••PERCENT NUTRIENT CONTENT BY SAMPLE SPECIF ICATI0N***************^***^*******^**+****^***^***********+*************^***»»****«**» • WOOD*«SECT ION * * BARK»*SECTI ON * 1 _ 2 5 7 9 10 1 3 5 7 9 L9 NITROGEN -JLHILSPjHOJi!JJi£_ CALCIUM MAGNESIUM -POTASSIUM -0. 052 J)._C0 5_ 0. 105 0.011 0. 0*4 0.054 _0.op3_ 0.095 0.009 0.046 0.051 0.005 0.090 0.01 0 0.04 7 0.067 _0.0 ? a o.io'o" 0.010 0. 049 0.064 0.006 0.100 0.011 0.053 0. 085 0. 009 o".T6i 0. 01 0 0. 056 NITROGEN -_PJjOS_PHORUS-^ _ CALCIUM "'-MAGNESIUM -POTASSIUM -0. 300 0.05_5_ "2.951 0. 039 0.173 0.298 JL'J?73 2.160" 0.045 0. 166 0. 322 0. 086 "2.543 * 0. 050 0.255 0. 365 0.094 2.015" 0. 044 0.300 0. 417 0. 110 1.989"' 0, 064 0.296 0.516 0. 1 04 1.772 0. 062 0. 326 BRANCH BRANCH >1 1-1/4 (TOPI -TWIGS-( MIDI ( LOWI *-- — r f OL I AGE-IT OP) (MIDI (LOW I CONES * TWI GS + FOL I AGE * (TOP) ' (MID) " (LL'W I NITROGEN - 0. 155 0. 259 0.000 0.000 0.000 0.000 0.000 0. OOP 0. OOP 0. 8P7 0.776 P. 665 PHOSPHORUS-CALCIUM MAGNESIUM -0.023 0.037 0.719 0.735 0. 024 p. 032 0. 000 O.POO 0.000 0. 000 0. 000 0.000 O.OPP 0.000 O.OOP 0. 000 0. 000 0.000 O.PPO 0.000 0. 000 0.000 O.OOP 0.000 O.OOP 0.083 0.082 0.091 0.000 1.432 1.697 1.905 0. 000 0.084 0.088 0.114 POTASSIUM - 0. 036 0.138 0.000 0.000 0.000 0. 000 0.000 0.000 0.000 0.394 0.385 0.330 • ••NUTRIENT AMOUNTS IN DECAGRAMS BY SAMPLE S PEC IF ICAT I ON*********************************************"* ***»**«***•«*«*« »*»r***4*"*~ * WOOD^ * SECT ION- * * 8ARK««SECTI0N * 1£2 3(14 5 £6 7E8 10 TOTAL 1£2 3(14 5£6 7E8 10 TOTAL NITROGEN - 21.33 13.22 10.34 5.49 0.73, 0.23 51. 84 NITROGEN 7.77 PHOSPHORUS-CALCIUM MAGNESIUM -2. 32 41.21 4.21 1.65 23. 33 2.45 1.12 16.70 1.80 0.6 0 8.39 0.88 0.07 1.15 0.12 0.02 0.27 0.03 6.28 91.06 9. 50 PHOS'PHORUS*^ "" CALCIUM MAGNESIUM -1. 66 66.44 1. 10 7.96 """2.04 60.44 1.22 5.94 ~"1756" 39.43 0.81 2.66 P.'6"9 12.61 0.37 1.04 "0727" 4. 97 0.16 0.36 25.73 "0707 673*0" 1.24 186.14 0.04 3.70 17.60 POTASSIUM 13.63 11.74 8.39 4.2 3 0.62 BRANCH BRANCH •- -TWI GS->1 0.15 43.81 1-1/4 (TOP) (MID) (LOW) TOTAL POTASSIUM -FOLIAGE-4.40 TTUP) (MID) (LOW) TOTAL 5.40 4.80 2.03 0. 74 CONES *- TWIGS* FOLI AGE— (TOP)~ (MIDI (LOW) 0.23 TOTAL" TP EE "TOTAL' NITROGEN 19.61 13.65 0. 00 0.00 0. 00 0.00 0.00 0.00 0. 00 0. 00 0.00 13.48 38.19 37.88 89.55 200.39 PHOSPHORUS-CALCIUM MAGNESIUM -POTASSIUM 2. 95 90. 73 3.00 10. 34 1.95 41. 29 1.71 7. 25 0.00 COO 0.00 0.00" 0.00 0. 00 0.00 O.OP 0. PO P. 00 0. 00 0.00 0.00 0.00 jO.OO^ 0.00 0.00 0.00 0. 00 O.JO 0. 00 0. OP P.PO P.PO 0.00 0.00 " 0 7 0 0 O 7 0 ' 0 ~ 0. 00 0. 00 0.00. 0.00 0.00 0.00 " " 0 7 0 0 0"70'0" 1.39 23.91 1.41 ""6.57" 4.01 83.51 4.31 "18". 96" 57015 97"41T 83.45 190. 86 4.99 10.70 "14.46 40.00 26.83 600.C9 28. 61 "115.50 " TREE NO. 103 SPECIES NO. 1 LENGTH (Ml 15.5 DBH (M) 0.183 CROWN OIA(M) 2.9 CROWN BUTT L.IM) L.(M) 10.3 5.3 •••PERCENT NUTRIENT CONTENT BY SAMPLE SPECIFICATION****************************************************** • WOOD**SECTION * * BARK**S ECT ION * 1 3 5 7 9 10 1 3 5 7 9 10 NITROGEN -PHOSPHORUS-. CALCIUM MAGNESIUM -POTASSIUM -0.03 7 _0_.-P_39_ 0. 047 0.011 0. 056 0.040 _0_._012_ 0. J43 0.010 0.051 0.036 0.008 0.046 0.015 0.056 0.016 0. 063 0. 014 NITROGEN -PHOSPHORUS-0.200 0. 037 0.217 0.045 0. 047 0.010 0.051 0. 048 0.010 0. 055 0.039 0.011 0.062 0. 04 5 0. 010 0. 062 CALCIUM MAGNESIUM POTASSIUM 0. 574 0.029 0. 120 0.552 0.041 0. 192 0.236 _0.06 0_ 6. 500 0. 059 0. 257 0.268 _0.06 3 0. 399 0.059 0.213 0.342 3.075 0.329 o.jno 0. 41 7 674 00" 0.067 0.060 0. 237 0.232 BRANCH BRANCH * TWIGS- -FOL IAGE-NITROGEN ->1 0. 113 1-1/4 0. 161 (TOP) 0.454 (MID) 0. 514 ( LOW) 0.413 (TOP) 0. 788 (MID) 0. 792 (LOW) 0.779 CONES 0.000 * TWIGS + F0LI AGE-- • (LOW) ( TOP) (MID) PHOSPHORUS-CALCIUM MAGNESIUM -0.013 0.115 0. 025 3.024 0.223 0.031 0.036 0.340 0. 08 7 0.078 0. 350 0. 089 0.065 0.299 0.075 POTASSIUM - 0.079 0.093 0.208 0.221 0.179 0. 124 0. 363 0. 126 0. 341 0.119 0. 49 7 0. 130 0.122 0.537 0.137 0.292 0.246 0.000 0 0.000 0.352 _0.000 0.10 6. oob •••NUTRIENT AMOUNTS IN DECAGRAMS BY SAMPLE SPECIFICATION************************************************************************** •W000**SECT10N- -BARK**SECTION-JNJJROGEN ^_ FHOSPHORUS-CALCIUM MAGNESIUM -162 6.27 3E4 5 C 6 0.76 0.53 7C8 0.41 0.09 10 0.03 TOTAL 2.80 NITROGEN 1C2 0. 90 1.16 0.27 0.20 0.90 0.20 0.15 0.52 0. 13 0.13 0.3 5 0.09 0.03 0.07 0.02 0.01 0.02 0.01 0. 79 3.11 0.71 PHOSPHORUS-CALCIUM MAGNESIUM -0. 18 2. 42 0.15 3G4 0.61_ 0.14 1.42 0.13 5 £6 0.58 "0.14" 1.03 0.14 7£8 9 0.33 0. 13 0.07 0.03 0.45 ' 0.17 0.07 0.03 10 0.14 "O'VO'S" 0. 16 0. 02 TOTAL 2. 68 0.59"" 5.65 0. 54 POTASSIUM 1.36 1 .01 0.63 0. 4 7 0.11 0.03 3 . 66 POTASSIUM - 0. 67 0.61 0.54 0.25 0. 09 0.09 JSRANCHJ3RANCH * I- 1/4 >1 NITROGEN 0. 92 2. 65 (TOP ) 0.68 •--TWIGS --(MID) "(LOW) 1 .08 1. 16 TOTAL 2.92 ( TOP) 0.55 •-FOL I AGE "(MID) (LOW) 0.79 1.01 * "TOTAL" 2.36 CONES *- TWIGS+FOLIAGE— (TOP) (MID) (LOW) 0. 00 1.23 1. 87 2.17 TOTAL 5.27 "0.94 3.53 0. 92 2.13 2.25 TREE "TOTAL" 14.33 2T7T" 17.62 2.90 ' 10.24 PHOSPHORUS-CALCIUM ^MAGNESIUM _-_ POT AS S1UM -3.15 0.39 1.64 3.68 3.21 0.51 0.67 " 1.53 0. 13 0. 51 0. 13 0. 31 0.16 0.16 0.73 0. 84 0.19 _ 0.21 0.46 6. 50" 0.47 0.09 2.08 0.25 0.53 0.09 1.28 " 0.24" 0.12 0.16 0.50 0.70 0.13 0.18 0. 29 0. 32 0.37 0.00 1.45 0.00 0.40_ 0.00_ "0.65 " "0.00 0.22 0.77 0.22 0.55 0.26 0.34 1.23 1.54 0. 32 0.39 0.76 " 0.82 TREE NO. 10* SPECIES NO. 1 LENGTH ( .Ml 17.2 OBH (Ml 0.325 CROWN 01 A(Ml 4.0 CROWN BUTT L.(M) L.(M) 10.4 6.9 •••PERCENT NUTRIENT CONTENT BY SAMPLE SPECIFICATION*************************************** • WOQO**SECT ION * * BARK**SECT 1 ON * 1 3 5 7 9 10 1 3 5 7 9 10 NITROGEN -J_HI3SPM9_RUS:L_ C A L C I U M -MAGNESIUM -P O T A S S I U M -0. 0*3 J). 03 3_ 0.0 5 7 0. 016 0.03 3 0.039 0.014. 0.076 0.015 0.058 0.037 _0.012 0.028 0.00 6 0. 0*9 0.0*5 _Q.011_ 0.03B 0. 009 0. 055 0.053 .0.011 6.0 38 0.009 0.056 0. 07* .0. 01 6_ 6." 0*3 0.011 0. 073 NITROGEN -PHOSPHORUS-CALCIUM MAGNESIUM -POTASSIUM -0.153 0.029 "~0. 5Of" 0.020 0. 081 0.161 _0._032 6.63 9 0.023 0.098 0. 197 0.033 "0.5 25 0. 025 0.101 0.267 0. 050 6.27* 0.036 0.190 0.290 0.313 0.051 0.057 0.260 ' 0.266" 0.035 0.0*2 0.178 0.199 J3RANCH BRANCH 1-1/* NITROGEN ->1 0. 117 0.167 I TOPI 0.633 -JWJGS-( MI Dl 0.536 ( LOW) 0.533 (TOPI 0. 6*3 -FOLIAGE-" (MID) 0. 6*9 CONES (LOW ) 0.632 * TWIGS* FOL I AGE * "(TOP) " (MIO) (LOW) 0. 000 0. 593 PHOSPHORUS-CALCIUM MAGNESIUM -0. 018 0.22* 0. 021 0.032 0.2 0* 0. 02* 0. 09* 0.239 0. 055 0. 073 0.273 0. 052 0.066 0.251 0.050 0. 125 0. *13 0. 131 0. 130 0. 505 0. 1*1 0. 127 0. 5*3 0. 139 POTASSIUM - 0. 090 0.100 0.221 0.192 0. 158 0. *15 0.371 0.368 0.000 • ••NUTRIENT AMOUNTS IN DECAGRAMS BY SAMPLE SPEC IF ICAT I ON************************************************************************** «- WOOD** SECTION * * 8ARK**SECT I ON * 1C2 3t4 5&6 7(18 9 10 TOTAL 1(12 3d* 5&6 768 9 10 TOTAL NITROGEN - 4.09 3. *9 0.7* 0.*5 0.07 0.0* 8. 87 NITROGEN - 2.16 2.52 1.** 0.53 0.12 0. 13 6.89 PHOSPHORUS-CALCIUM MAGNESIUM -2.35 6.66 1.55 1.19 *. 76 0.9* 0.21 0.60 0.13 0.10 0.3 5 0.08 0.01 0.05 0.01 0.01 0.02 0.01 3.87 12. *3 2. 72 PHOSPHORUS-CALCIUM MAGNESIUM -0.*2 7.87 0.30 0.*5 8.20 0.33 0.26 2.*8 0.19 0. 10' 0.51 0.07 0. 02 0, 10 Oo 01 0.02 0. 11 0.02 1.27 19.27 0.92' POTASSIUM - 7.35 *. 90 0.9* 0,51 0.07 0.0* 13.81 POTASSIUM - 1. 23 l . * l 0.90 0.35 0. 07 0. 08 * .o* BRANCH BRANCH *-NITROGEN >1 2.07 1-1/* (TOP) --TWIGS (MIDI (LOW)"' *.99 0.51 0. 97 1 . 4 9 TOTAL 2.96 ( TOP I 1. 09 •j^F OL I AGE— (MIDI (LOW I CONES *-3. 63 4. 11 TOTAL 8. 83 TW I GS* FOL I AGE * (TOP) (MID) (LOW) TOTAL"' 0.00 1.60 4.60 5.60 11.80 TP EE "TOTAL" 3 4 . 6 2 PHOSPHORUS-CALCIUM MAGNESIUM -POTASSIUM -0. 32 0.95 3. 96 6. 07 0.37 0.73 1 , 60 0.03 0. 23 0.0* 2. 99 0. 13 0.13 0.19 0. *9 0.70 .0. 09 0. 1* "0.3 5 0.**" 0.39 0.21 1.42 0.70 JO. 28 0.22 0.97 0.71" 0.73 0.83 2.83 3.53 0.7 9 0.J91_ 2.08" "2.39 7.06 1. 92 5.17 0. 00 0.00 0.00 "0.0*0" 0.29 0.93 0.27 ""0.88"' 0. 86 3. 32 0. 89 "2.42 TTor" 4.23 1.05 "2". 83 ""2776""" 8.49 2.20 "6. 14" 8.57 50.22 6.93 28.58' TREE NO. 108 SPECIES NO. 1 LENGTH (Ml 27.7 OBH (M ) 0.490 CF OWN DIAIM) 7.3 CROWN L.(M) 23.2 BUTT L.IM) 4.6 • ••PERCENT NUTRIENT CONTENT BY SAMPLE SPEC IF I CAT IQN***********»******************************************************************* « WOOO** SECT ION * * BARK **S ECT ION * 1 3 5 7 9 10 1 3 5 7 9 10 NITROGEN PHOSPHORUS-0.049 0.012 0.064 0.02 9 0. 050 0.011 0.065 0.016 0. 063 0. 016 NITROGEN -PHOSPHORUS-0. 146 0. 021 0.155 0.023 CALCIUM MAGNESIUM -POTASS IUM -,03 3 0.059 0.009 0.012 0.059 0.067 0. C46 0.011 0.092 0. 047 0. 010 0. 069 0.038 0. 037 0.009 0.010 0.071 0.076 CALCIUM MAGNESIUM -POTASSIUM -0.235 0.255 0.016 0.015 0.071 0.097 0. 164 _0.J>26_ 0. 23 9 0.012 0. 095 0.191 _0_. 02 8 0.2 30" 0.012 0.098 0.255 _0.04 0_ Oc 176 0.016 0. 145 0.348 _0.054_ 0". 216 " 0.019 0.1 94 BRANCH BRANCH -TWI-GS- -FOLI AGE- CONES * TWIGS+FOLI AGE-NITROGEN ->1 0. 115 1-1/4 0. 146 (TOP) 0.609 (MID) 0.447 (LOW I 0.363 (TOP) 0. 905 (MIO) 0. 834 (LOW) 0.813 ( TOP) (MIO) ( LOW) 0.000 PHOSPHORUS-CALCIUM _MAGNES IUM_-POTASSIUM -0.02 3 0.155 _0.015_ 0. 071 0.023 0. 149 _0. 01? 0.098 0.035 0.292 _0.J>4 7_ 0. 218 0. 087 0. 3'i3 0. 043 0.055 0.194 0. 036 0.124 0.282 0. 092 0. 127 0.462 0. 101 0. 182 0.402 0. 123 0.201 0.121 0.234 0.343 0. 334 0.000 0.4 0.000 0.287 _0.000 0.06, 61 odd •••NUTRIENT AMOUNTS IN DECAGRAMS BY SAMPLE SPECIFICATION************************************************************************** -WO0D**SECTICN- -BARK**SECTION-1C2 3C4 5(16 7 £8 9 10 TOTAL 1£2 3 £4 5£6 7£8 9 10 TOTAL NITROGEN - 14.61 9. 55 5. 96 2.27 0.38 0.08 32.85 NITROGEN - 8.92 8.27 5.00 2.67 0.41 0. 14 25.41 PHOSPHORUS- 3.72 3.53 2.12 0.54 0. 10 0.02 10. 03 PHOSPHORUS- 1 . 30 1.26 0.7 5 O.'l ~ "0.06" 6 ;o'2 3.8i CALCIUM 13.66 8.87 4.33 1.69 0.22 0.04 29.37 CALCIUM 14. 54 12.79 6.61 2.43 0.28 0.O9 36.74 MAGNESIUM - 2.91 1.87 1.09 0.3 8 0.05 0.01 6.31 MAGNESIUM - 0.92 0.68 0.33 0.17 0. 03 0. 01 2.13 POTASSIUM - 17.53 13.39 8.43 2.77 0.41 0.09 42.63 POTASSIUM - 4.99 4.97 2.72 1.46 0.23 0.08 14.45 BRANCH BRANCH * - TWIGS * * — F O L I A G E * CONES_ * TWI GS* FOL I A G E - - TP EE >1 1-1/4 (f OPT (MID) (LOW) TOTAL (TOP) (MID) (LOW) "TOTAL ~ (TOP)" (MIO) "' (LOW) TOTAL " "TOTAL NITROGEN - 14.05 10.48 2. 74 4.47 3.49 10.70 9.96 17.34 13.08 40. 38 0.00 12.70 21. 81 16.57 51.08 133.87 PHOSPHORUS-CALCIUM MAGNESIUM-"POTASSIUM -2.77 1.68 18.99 10.67 1 . 9 3 1 . 3 6 3. 69 6. 33 0. 38 1.32 0. 21 "C.98' 0.87 0.53 3.13 1.86 0.43 0. 34 2. 0 1 1 . 16 1.78 1.37 2.63 2.93 6.31 3. 10 9.62 6.47 0.99 1.01 2.09 1.98 "4.15 " 2.57 7. 13 " 5.38 6.93 0.00 19.19 0.00 5.07 0.00 15.08" 0.00 1.75 4.42 1.22 3.55 3.50 3.46 12.74 8.34 2. 52 2. 32 9.14 6.55 8.71 25.50 6. 06 19.24 T 7 7 W 121.28 17.74 91. 34 A P P E N D I X 8 PROGRAM FOR C A L C U L A T I N G MOISTURE CONTENT OF SAMPLES FOR C H E M I C A L A N A L Y S E S P r o g r a m p r e p a r e d b y M r . K e n T w a i t e s .COMPILE C PROGRAM FOR CALCULATING MOISTURE CONTENT AND STANDARD DEVIATION OF UP TO C **120*« UNORYfD TREE SAMPLES CLASSED IN UP TO **9*« SPECIFICATIONS AMD C ORYE D AT TWO TEMPERATURES<TI,T2) C PROVIDE DATA CARCS AS FOLLOWS C 1. 4 INTEGERS ON 1ST CARD- NO. OF SAMPLES(M), NO. OF SPEC I F IC AT IONS ( N ) , C 1ST DRYING TEMP.(Til, 2ND DRYING TEMP.(T2) C 2. N CARDS WITH CCMENTS UP TO 12 CHARACTERS FOR EACH SPEC. - DATA IN QUOTES C_ 3. M DATA CARDS FORMAT A3 ,11 , A2 , Al ,4F7.3 , 13 (TREE NO., SPECIFICATION, "C DISC NO., LOCATICN'WITHIN'CROWN, EM PT Y ROT TLE WE I GHT(A) , BOTTLE•UNDRYE6 C .SAM PL5(BI, ROTTLE+S AMPLE DRYEO AT 1ST TEMP.(C), BOTTLE+SAMPLE DRYED AT C 2ND TEMo. (Ol, BOTTLE NO. •INTEGER M, N,Tl,T?,SPEC(120l,ROTTLE(120),COUNT!9)/9*0/ RE AL* 9 A(l20),B(120),C(120 I,D(120) ,WB(120 I,WC(1201,WD(1201, 1MC T 1 ( 9 I /9* C . 000 /,MC T2 ( 9) /9*C. CDC/, MC 1U 20 )_, MC2 U 20 I , (9J79*6.obd7",bsoRf 1 SUMT1 (9) /'-(*0.0()0/, SUMT CHARACTER CHAR*12(9 I,TREE*3(120 I,DISC*2(120I ,CR0WN*1(120 I INITIAL READ IN CF ALL DATA i i I ! 30 REA0,M,N,T1,T2 DO 30 1 = 1,N RE AO, C H AR ( I) RE All 5 ,1 .END = 50 I ( TRE E ( I I , SPEC ( I 1 .DISC (I) .CROWN (I ), AI l l , IB( I),(. ( I),0( I ), BOTTLE ( I ) , I=1,M) FORMAT! A3, 11, A2 ,41 ,4F7.3, 131 10_ "i l _L*L 13 GO Tl 40 C CONTROL TO H£RE IF END OF DATA ENCOUNTERED DURING READ IN BEFORE M CARDS READ 50 WRITE (6,21 M _ ^ ___ "2" FOR MA T ( • 1' , • E "f6 OF DATA" ENCOUNTERED BEFOR E '7 13, ' CARDS READ"1! 1'. ..RECOUNT CATA CARDS') GO TO 51 14 "1 5 16 1 7 40 . . CLNTI MUE C CALCULATE WEIGHTS DO 100 1=1 ,M LESS BOTTLE WEIGHT AND MC.IN PER CENT FOR EACH SAMPLE W3(11= B(11-A( 11 WC. ( I I = C ( I ) - A ( I ) WD ( I I = D ( I ) - A ( I ) 18 19 20 MCI(I I= (W S(I)-WC(I)l/WBII 1*100.DO MC2( I l = (Wtl ( I l-WCI I I l/WBI I 1*100.00 CONTINUE _ C PR I NT OUT OF "GRIG INAL DATA. WEIGHTS MINUS BOTTLE WEIGHT,-~070 MC FOR EAtH SAPPT" C AT TEM^ 1 ANO 2 C **HE AOI NG ORINTOUT' 100 21 WRITE!6,10 I 22 10 FORMAT!'1' ,T2,' INITIAL PRINTOUT OF DATA READ IN', T69, ^ I >**«• ,T75, 'CALCULATIONS-WEIGHTS, 0/0 MOISTURE CONTENT' I  2 3 ' " 'WRITE (5, 11 I " " " ' ' 24 WRITEI6.12I T1,T2,T1,T2 25 11 FORMAT) 'U' ,T2, ' TREE ', T8,' SPEC ,T14,' PI SC , T20 , ' CP.CWN' ,T29, '*A*', lT3a,'*3*',T47,•*C*',T56,<*D*',T63,'BOT',T69,'***',T75, l'WTS. MINUS EMPTY WT . • , T98 ,' MC (0/0 ) •, T 10 7 , • MC (0/0 ) • ) 26 12 FORMAT!' • ,T3, 'NO. ' ,T15, 'NO. ' ,T21, 'LOC. • ,T28, • EMPTY' ,T36_, "1 'UNJRYET ,'T45, I 3'i ' ~ORY' ,^54,13", • DR Y • , T 63",'' NO. ' ,"T69, •***r7T7"57 1' *H* *C* *D*',T98,13,• DRY' , T107, 13,' DRY') C **P«tNTOi.lT OF ORIGINAL DATA AND CALCULATIONS 27 WRITE (6,13 I 28 13 FORMAT ( • 0' I 29 wR ITE (6,14 ) ( TRE E( I ) ,SPEC( I I ,DI SCI 1 ) , CROWN (I ) ,A( I I ,R( 11 ,CI I I ,D( I ), """ : " 1 S 0 T T L E ! I I, W*< T I ,WC( 11 , WDI I ), MC 1 ( I) , MC2 ( I ) , 1=1, Ml"" 30 14 FORMAT!' ' , T 3 , A 3 , T 10, 11 , T15 , A 2 , T22 , Al , T27 ,F 7 . 3 , T36 , F7 . 3 , T45 , 1F7.3,T54,F7.3,T63, I3,T69,'***',T76,F5.3,2X,F5.3,2X,F5. 3.T9 8, © r 1F7.3,T107,F7.3) C CALCULATING AVERAGE MOISTURE CONTENT o 31 00 99 9 1=1 , M SI 32 COUNT! SPEC ( I ) l=C0UNT( SPEC I 1)1 + 1 33 MC T i l SPEC! I 1 I = MCT1 (SPEC! I ) l+MCK I) o 34 MC T>( SPEC( I ) 1 =MCT? ( SPEC! I) )*MC2 ( I 1 > © 7 35 999 CONTINUE 36 00 10 0 0 1= 1, N o 37 IF ( CJ 'JNTI I 1 . LE. Ot GO TO 1 0 0 0 o 3 3 MC T I ( I ) = MC Tl ! I 1 /COUNT I I ) 3 9 MO. T2( I )=MC T2 ( I 1/COUNT < I 1 o 40 1P00 CONTINUE o. C CALCULATING STANDARD OEVI AT ICN FROM FORMULA... C STO OEV = SORTISUM FROM 1 = 1 TO M OF ( X < I t-A VG( X ) )** 2/( M-11 ) *s 41 DO 1 ) 0 2 1=1, M C1 4 2 S'JMTl ( S P E C I I ) I = SUM T 1 ( SPEC ( I I 1•(MC1( I 1-MCTK SPEC! I ) 1 )**2 43 SUMT2(SPEC I I I t=SUMT2(SPEC( I1 1 + IMC2!I)-MCT2(SPEC( I)I 1**2 -> 4 4 1002 Cf N T I N U t f ' 45 DO 100 3 1=1, N 46 IF(COUNT!Il.LE.1) GO TO 1 0 0 4 o ' 47 SUM Tl ( I l=DSORT( SUM T1 ( I ) /(COUNT! 11-11) <~. 48 SUMT2 ( I l=OSQRT( SUMT2! I ) / (CCUNTI 11-1)1 4 9 GO TO 1 0 0 3 Q 50 1 0 0 4 s i r m ( i )=e .000 O 51 SU MT2 ( I ) =0 .0 DO 52 1003 CONTINUE n C FINAL °R I NT OUT c 53 WRITEI6.2rOITl.T2 54 200 FORMAT!'-' ,T4,'SPFCIFICAT ICN' ,T23,'NUMBER',T33,,**,,13,' DEG. C ', O 1'DRYING**'.T56, •**•', 13. • DEG. C DRYING**') c 55 WR I TE ( 6 » 20 11 56 201 FORMAT!' ',T22,'IN GROUP') O 57 WRITE I 6, 20 2) 53 202 .FORMAT!' ',T35,' AVG MC S TO DEV ' »T58, • AVG MC STD DEV •) 59 WRITE (6, 20 21 ) o 60 2021 FORMAT! ' 0 ' 1 61 00 100 1 1 = 1, N 62 IF ( C O U N T ! Il.LE.0» GO TO 800 D 63 WR I TE ( 6 ,2C 3 1 I ,CHAR ( I) ,COU^( I 1 , M , MCT 1 ( I 1, SUMT1 ( I 1.MCT2I I 1, c 1SUMT2III 64 203 FORMIC ',T2,I3,' - ' , A12 , T23, 13, • / •, I 3, T35, F 7. 3 ,T44, F 7. 3 , T58 , J> 1 F 7 . 3 . T 5 7 . F 7 . 3 ) 65 GO TO 1 0 0 1 6 6 800 WR I TE ( 6 , 2 0 4 ) I ,CHAR ( I I .COUNT! I ) , M , I 67 204 FO RM\T(1 ',T2,I3,' - ' ,A12,T23,13,*/',I3.T75, l>***|f| OAT A GIVEN FOR GROUP ',11,'***') 68 1001 CONTINUE C CALCULATING TOTAL FOR ALL GRCUPS 69 MC T l ! 1 1 = 0. 000 70 MCT?! 1 1=0.000 71 DO 2 0 0 0 1=1, M 72 MCTK I ) = MCT1 ( 1 1 + MC 1(1) 73 MC T2( 1 ) = M 0 T 2 ( 1 1 *MC2 (I 1 v j : 74 2 0 0 0 CONTINUE' 75 MCTK 1 ) = MCT1 ( 11 /M 5 76 MCT2I 1 1 =MC T2( 1 ) /M 77 DO 2 0 0 1 1= 1,M \^ 73 SUMTK 11= 0 . 0 0 0 . 79 SUMT2 ( 1 ) = 0 . 0 0 0 80 SUMTK 1 >=SUMT1( 1H-IMCK I )-MCTl(l)l**2 > 81 SUM T2 ( 1 ) = S UMT2(11•< MC2(I I-MCT2I 1) )**2 82 2001 CONTINUE 83 SUMTK 1 )=OSQRT( SUMTH l l / I M - l l ) 84 SU MT2 ( 1 I =0 SORT ISUMT2(1 I/(M-lI) C PRINTOUT OF TOTALS 8 5 WR irE16.?06l",M, MCTK 11, SUMT1 ( 1 I ,MCT2( 1), SUMT2 I 1)  86 206 FORMATCO' , T 2 , ' * * * * A L L GROUPS****•,T23,13,•/•.13,T35,F7. 3, 1T4*,F7..3,T53 ,F7.3,T67 ,F7.3) 87 51 WRITE ('6,20 51 •_ 8 3 205 FORMAT P I ' I 8 1 RE TURN go__ EN 0 . tDATA 157 A P P E N D I X 9 S P E C I F I C G R A V I T Y PROGRAM P r o g r a m p r e p a r e d b y M r . K e n T w a i t e s C G C 3 < i/) I— Ul < CL C l _J I— > </) Z 3 C iu — ac o I o: O (_J O * - _ J c a. vi _J D < ui c Q c: | o; c ifl 3 c ^ a r <r i/i c c — —-• -* • C —I C- ti- > I- 7> ft. V • ii o u' — I: or i. " ' - « a: -M S — D m -•r . - c > " — a O, t_j • U l/l L> D • UJ 1 •• 3- a. G. i/~ a- •* - it- c- I* •j - fv cc < - ,U: - -• —> f\j • -J- IT. ' > • o -[O — \T\ — I • n-, > PS > > — <r ,_J — > + o — c I o 1 o c — cc •!_J C '•<> "2 *" I J J C ! * > > - ; -o v: u.* i x O a. — j <: u I • -I \f~ 7. • 1 u: . r' c i -—: _ i 'v; <r U. U- - X - — j f i r © c c c A P P E N D I X 10 PROGRAM FOR C A L C U L A T I N G BIOMASS D I S T R I B U T I O N FOR SAMPLED T R E E S SCOMPHE C - M O P E L OP BIOMASS DISTRIBUTION FOR 20 TREES FROM HIGH ELEVATION SAMPLE C - P L O T N E A R S Q U A M I S H . I N T E G E R T R E E , S E C ( 1 0 ) , 8 P , A G E , B U T L 1 , R U T L 2 , C R O P t , C R O D 2 R E A L V O L W ( 1 0 ) , V O L B ( 1 0 ) , S P G W ( 1 0 ) , S P G B ( 1 0 ) , L , l . T 0 6 , L T 0 1 , D B H , B U T L , I C R O D . F K B I , F w T N S ( 3 ) , F W S B l , F W S B U , F W S T N ( 3 ) , F W B N H ( 3 ) , F W H 4 S ( 3 ) , F W C O , 2 F W S C 0 . P W B 1 , B w S B 4 , D W S T ( 3 ) , P W S N ( 3 ) #DWSCO,DWWSC 1 0 ) , D W B S U O ) ,[)WW,DWB, 3 M C 8 l , M C R « , D W B U ( 3 ) . M C T N ( 3 ) , D W T N ( 3 ) , D W T ( 3 ) , D W N ( 3 ) , D W B « T , D w T N T , D W T T , 4P W N T,0W C0,D W H1,M C C0 I T K E E=0 C - C A L C U L A T I O N FOR A T R E E B E G I N C - R E A D I N T R E E P A R A M E T E R S A N D C R O W N C O M P O N E N T S F R E S H W E I G H T S DO 1 0 0 1 = 1 , 2 0 H E A D ( 1 , 1 0 1 ) T R E E . S P ; A G E , L 1 , L 2 > L T 0 6 t , L T 0 6 2 , L T 0 1 1 , L T 0 1 2 , R B H , B U T L 1 » "" ' i J U U T L 2 , C R 0 n i , C R 0 D 2 , F W H 1 , C F W T N S ( M ) , M s 1 , 3 ) , F W S B l , F W S B 4 , ( F W S T N ( K j 2 K = 1 , 3 ) , ( F W B H R ( J ) , J s l , 3 ) , < F W R 4 S ( N ) , N = 1 , 3 ) , F W C O , F W S C O 1 0 1 F Q R M A T { 1 3 L LL I 3_X j j . 3 , 3 ( I J , Hl,±; i , 1 , 4 ( 1 2 ) , 4 X , F 6 . 1 , 3 X , i ( F S . l ) , 5 ( F 3 . 1 ) / 1 3 X , i ( F 5 .1 ) , 3 ( F 4 , 1 ) , F 4 , 1 , F 1 , 1 ) C - R E A D I N D R Y W E I G H T S O F C R O W N C O M P O N E N T S 7 R E A D ( 5 , 1 0 2 ) D W S B 1 , D W S H « , C D W S T ( K ) , K s l , 3 ) , ( D W S N ( J ) , J = 1 , 3 ) , D W S C O F b R M A T ( « X , B ( M , 3 ) , F 3 , 3 ) " ' • " " ' D W « s D W B = D W R U T = P W T N T i D W T T = O W N T = D W C O a D W B l s O , 0 102 5 , 0 0 0 6 , 0 0 0 7 , 0 0 0 8 , 0 0 0 _ ' ' , 0 0 0 1 0 , 0 0 0 1 1 , 0 0 0 1 2 , 0 0 0 1 3 , 0 0 0 1 1 , 0 0 0 1 5 . 0 0 0 _ i 6", 600 1 7 , 0 0 0 1 8 . 0 0 0 1 9 , 0 0 0 " 2 0 , 0 0 0 J ' • ooo 2 2 , 0 0 0 2 3 , 0 0 0 2 U , 0 0 0 " 2 5 , 0 0 0 " 2 6 , 0 0 0 _ * E J t T E N 3 I 0 N * O T H E R C O M P I L E R S MAY NOT A L L O W M U L T I P L E A S S I G N M E N T S T A T E M E N T S • E X T E N S I O N * • E X T E N S I O N * • E X T E N S I O N * • E X T E N S I O N * • E X T E N S I O N * ' E X T E N S I O N * 1 0 1 1 ' 2 — 1 3 O T H E R C O M P I L E R S M AY NOT A L L O W M U L T I P L E A S S I G N M E N T S T A T E M E N T S O T H E R C O M P I L E R S M AY NOT A L L O W M U L T I P L E A S S I G N M E N T S T A T E M E N T S O T H E R C O M P I L F R 3 M AY NOT A L L O W M U L T I P L E A S S I G N M E N T S T A T F M E N T S O T H E R ' C O M P I L E R S ' M A Y * NOT A L L O W M U L T I P L E A S S I G N M E N T S T A T E M E N T S " O T H E R C O M P I L E R S M AY NOT A L L O W M U L T I P L E A S S I G N M E N T S T A T E M E N T S O T H E R C O M P I L E R S M AY NOT A L L O W M U L T I P L E A S S I G N M E N T S T A T E M E N T S DO 2 Ma i ,3 O W T ( M ) = 0 , 0 D W N ( M ) = 0 , 0 1« 1 5 16 1 7 1 8 1 9 2 0 2 1 " 2 2 2 3 2 4 2 5 2 6 2 "* 0 W T N ( M ) 3 O , 0 ' C-DATA T R A N S F O R M A T I O N TO M E T R I C S Y S T E M L = 0 , 3 0 « f l * F L O A T a i ) + 0 , 0 Z 5 « * F L O A T ( L 2 ) L 1 6 6 6 , 3 6 « * * r-L o A t " L"T 0 6 i ) • b , o "2 S a * F L b A T (L T o 6 2 y ~ L T O 1 * 0 , 3 0 4 8 * F L O A T ( L T 0 1 1 ) • 0 , 0 2 5 4 * F L O A T ( L T 0 1 2 ) _ _ _ B U T L = 0 , 3 0 i 4 8 * F L O A T ( B U T L l ) + 0 , 0 2 5 « * F L O A T ( B U T L 2 ) '" D B « i O , 0 2 5 < 4 * O b H : '[ C R n D s ( 0 . 3 0 i 4 B * F L O A T ( C R O D l ) + 0 , 3 0 a 8 * F L O A T ( C R 0 0 2 ) ) / 2 , 0 ; F W H l s 0 , « 5 i 5 9 2 * F W H l ' • " F W S B l = 0 , 4 5 3 5 9 2 " i F " w S B " i ~ ~ F » S B 4 s 0 . 4 5 3 5 9 2 « F ^ S B 4 C R O L = L - H U T L 00 1 0 3 M = 1 , 3 " •" ""' " ' • ' - ' F W T N S ( M ) s 0 , « 5 3 5 9 2 * F W T N S ( M ) F W S T N ( H ) s 0 , 4 5 3 5 9 2 * F W 3 T N ( M ) • ' \- ' -.27 F - H N R ( M ) s O , a S J 5 9 2 * > W H N R ' M ) .. , } 28 103 F w B 4 S ( M ) s 0 , 4 5 3 5 9 . 2 * F W . R 4 S ( M ) 29 F W C P = 0 , u 5 3 5 9 2 * F W C O ' _ -••' 3 0 '" " • F W S C O s 0 . 4 S 3 5 9 2 « F * S C O ""' C - B I O M A S S F O R T R F E T R U N K (WOOD A N D BARK) C - R E A D I N S P E C I F I C G R A V I T I E S O F S E C T I O N S 3 1 R E A D ' ( 3 , t 0 6 ) ( S P G W ( J ) , S P G B ( J ) , J « 1 , 1 0 ) - 3 2 1 0 6 F 0 R M A T ( 2 I X , F 5 , 3 , l i X , F 5 , 3 ) 3 3 0 0 1 0 4 j = !, 1 0 . . . '. ?.' C - R E A D I N V O L U M E S OF S E C T I O N S • ~r;-; ;- ' ~; lU R E A P ( 2 , 1 0 5 ) V O L W ( J ) , V P L B ( J ) -'-3 5 1 0 5 F O R M A T ( 1 5 X , F 5 , 3 , 2 X , F 5 , 3 ) . 36" " " " V O L W ( J ) s i 0 0 0 * v b L W ( J ) . 2 7 , 0 0 0 2 8 . 0 0 0 2 9 , 0 0 0 _ " 3 0 , 0 0 0 3 1 , 0 0 0 3 2 . 0 0 0_ " 3 3 , 0 0 0 3 4 , 0 0 0 3 5 , 0 0 0 3 6 , 0 0 0 3 7 , 0 0 0 3 8 . 0 0 0 3 9 , 0 0 0 4 0 , 0 0 0 4 1 , 0 0 0 4 2 , 0 0 0 4 3 , 0 0 0 4 4 , 0 0 0 "45","0"0"0~ 4 6 , 0 0 0 4 7 , 0 0 0 4 8 , 0 0 0 4 9 , 0 0 0 _ 5 0 , 0 0 0 _ 5 1 , 0 0 0 5 2 , 0 0 0 5 3 . 0 0 0 5 4 , 0 0 0 5 5 . 0 0 0 5 6 . 0 0 0 " S 7 ,0 00"" © © © © © e o 0 0 © o o o o i o o o o o o l o o o o o o o o o © o ; O O O O © OIO o o o o o O O O 0 0 | 0 0 0 O O C I O O O o o o O O O ' . 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U i 4 u. \ O U i <t • UJ < • 1 '< o -» 3i o T~- CO I «* — 31 o or *- r x - »- r »~ -t~ X Q. 1 M c a i n . o w m « *-• rr o - " S O a 1 Or X O »— '-«to. or c o> x o a c o X U . L t C CO X or o o or o »- z • •. V.,; — X O U. » - K H i . O - i U- u. — - JC u . u - s to. eo toi ',,v'''''" 1 X — eg — rvi . -« A. i . < 1 i *- ; i m *»» X CC) o o —• o ru ** J i rg rv o ru a I ! *•> * ~* *n i . i • i i tn x K C 0 O o ru ^ i n « s co a o mm i i. i o « a a t> o r> o a- o- o O ! ' '. - i V j 1 » t • 1 $ * t % » i > 1 1 t A P P E N D I X 11 PROGRAM FOR C A L C U L A T I N G N U T R I E N T D I S T R I B U T I O N P r o g r a m p r e p a r e d b y M r . K e n T w a i t e s iCOMPILE C **NUTR1ENT DISTRIBUTION PROGRAM** C FUR UP TO 99 TREES INTEGER TREE,SPECIE.NUMBER,NTREES REAL L.DBH.CROD.CROL,BUTTL,DWB1,DW84T,DWCO,DWTT,DWNT,DWTNT, 1DWTI3) ,DWN(3),DWTN(3),TRET,DWWSI 10),OWW,DWBS110).DWB,MCDATA( 8) 1. 000 2.000 3. 000 4. 000 5.000 6.000 REAL NPCR!21),PPER(2i),CAPER!211.MGPEP (21),KPER(21 I,WT(6),NUT(6) R = AL NW0HDC7,51,NBARK(7,5 I ,NB1(5),NB4T(5 I,NT(4,5),NN(4,5),NC0(5>, J.NTN! 4,5), PE-RCNT.. TOTAL (5.) , JNN ( 3 (., TN P ( 3 I ,TNCA( 3 )., TNMG ( 3 ), TNK (.3 ) , 1ZER0I 3 l/3»0. 0/ COMMTN N!JM3;R/AREA2/MC0ATA COMMON /AREA! / N°ER . P PEP., CAPE R, MGP ER . K P ER 7. 000 8.000 9 . 000 10.000 11.000 1 2. 000 99 CHARACTERS 1 CHAR(5) / 'NITROGEN 1'MAGNESIUM -',«POTASSIUM -'/ _REAO \ \ OF. NUMBER OF TREES REAM 5 , 99 INTREE S FCRMATI 12 I R£AO IN OF .MOISTURE CONTENT DATA -• , • PHOSPHORUS- ', 'CALC IUM 13,000 14.000 15. 000 16, 0;)0 17, 000 18. 000 10 REVO! 5 ,100) (MCDATA! I) ,1 = 1 , 11 100 FCRMATI3IF5.3.1 X) ) _XZ 0) 1 1 =1 j.N.TREE S 8) READ IN OF BIOMASS DATA FOR ONE TREE L3 READ!5,1011 TRE E,SPEC IE,L,DBH,CROD,CROL,BUTTL,DWB1,DWB4T, DWCO, 1DWTT, OWNT,DWTNT,(OWT! J) , DWNI J ) ,DWTN(J) , J = l ,3), TRE T, DWWS ,DWW, DWBS , 19. 000 20. 000 21,000 '"22.000" 23.000 24. 000 inwB ' 14 101 FORMAT! I 3,1 I ,F4.1 ,8X, F5. 3 , 3F4.1 ,2F6. 1, F4. 1.3F6. 1/7X.9F6.1, F9. 1/ 17X, 10F6. 1 ,F9. 1 /7X.1QF6.1.F9.1) ;  15 NUM3ER=21 ' 16 IF! S°ECIE.E 0.2. OR.SPECIE. EQ. 3 I NUMB ER =1 8 C CALL SJ-iPRUGRAM TO REAP IN NUTRIENTS N AND P FOR ONE TREE AND RETURN 25. 000 26. 000 27.000 "28. 000" 29. 000 30.000 PERCENTAGES FOR EACH SPECIFICATION ( EG. WOOD , BAR K. . . ) 17 CALL SNP(NPER.PPER) C CALL SJBPRUGRAM TO READ IN NUTRIENTS CA ,MG,K FOR ONE TREE AND RETURN C • PERCENTAGES FOR EACH SPECIFICATION 18 CALL SCAMGK (C AP ER ,MGP ER, KPER ) 19 1 F( SPECIE.EQ. 2. OR.SPECIE.EQ.3 ICQ TO 1004 31. 000 32.000 _3_3_. 000_ 34. 000 35. 000 36. 000 ?0 C CALCULATING PERCENT NUTRIENTS FOR T WIGS CFOLI AGE FOR SPECIES 1 BY AVERAGING 37.000 C THE PERCENTS FOR TWIGS AND PERCENTS FOR FOLIAGE 38.000 N1 = 14;N2 = 17 • • 39.000 22 23 24 00 29 K=l,3 TNNIK ) = ( NPER1K + N1 ) «-NP ER ( K+N2 ) )/2 T.NP I <) = ( PPER (K>N1 l+NP£R(K»N2) 1/2 40.000 41.000 42. 000 25 26 27 TNCAIK) = (C*PER(K + NlH-CAPER(K + N2))/2 TNMG! K 1= ( MGPERI K+Nl ) +MGPER (K+N2) ) / 2 TNK!<)=(K0ER!K+N11.KPERIK+N2)1/2 28 29 CONTINUE 29 1004 CCNTINUE C *««CALCULAT1NG NUTRIENT CONTENT BY SAMPLE SPECIFICATION*** C SSPEC - WOOD C CALCULATES NUTRIENT CONTENT FOR WOOD FOR THE 5 NUTRIENTS AND STORES IN ARRAY C NWOOO!7,51.COMBINES BIOMASS WOOD DATA FOR SECTIONS 1C2.3C4,5tb,7C8,9,10 . 30 31 32 DO 2 J = 1,4 WT! )l =DWWS(2*J-1)*DWWS(2*J) CONTINUE ' . . ": 43.000 44.000 4 5. 000_ "4 67 000 47. 000 48. OOP 49.000 50. 000 51. 000 52.000" 53. 000 54.000 33 WT!5)=DWWSI91 34 1T( 6) = DWWS( 10) C GETS NUTRIENT PERCENTS FOR APPROPRIATE SEC TlONS,AVER AGES,PUTS IN ARRAY NUT(6) 3 5 "00~i J=l,5 36 NWOOO! 7, J I =0.0 37 N = l - - "55. 000 56. 000 57.000 5 8.000" 59.000 60. 000 f 38 GO TOt 10,20,30,40,501 , J 61.000 39 10 CALL CALC(N,NPER,NUT,£60) 62.000 40 20 CALL CALC(N,PPER,NUT, £60) 63.000 41 30 CALL CALCIN,CAPER,NUT,660) 64.000 42 40 CALL CALCIN,MGPER,NUT,660) 65.000 y- 43 50 CALL CALCIN,KPER,NUT,660) 66. 000 44 60 00 4 K=l,6 67. 000 < 45 NWOOO IK,J )=WT IK ) *NUT 1 K ) 68. 000 46 NWOOO( 7, J)=NW00D(7, J) «-NWOOO(K,J) 69.000 47 4 CCNTINUE 70.000 48 3 CONTINUE 71. 000 © C SSPEC - BARK 72.000 C CALCULATES NUTRIENT CONTENT FOR BARK FOR THE 5 NUTRIENTS AND STORES IN ARRAY 73.000 © c NBARKI7,5),COMBINES 310MASS BARK DATA FOR SECTIONS 1£2,3£4,5£6,768,9,10. 74. 0 00 49 DO 5 J = l ,4 75. 000 50 WTI Jl = DW3S~< 2*J-1) *DWBS(2*J ) 76.000 51 5 CCNTINUE 77. 000 52 •Ul 5) = D*3S( 9) 78. 000 53 WTISI=DWBS(10) 79.000 C GETS NUTRIENT PERCENTS FOR APPROPRIATE SECTIONS,AVERAGES,PUTS IN ARRAY NUT I 6) 80. 000 0 54 no •> .1 = 1,5 81.000 55 N6AUI7, J) = 0.O 82. 000 56 N = 7 e3.000 57 GO TlI 11,21,31,41,51),J 84.000 59 11 CALL CALCIN,NPER,NUT,£61> 85. 000 © 59 21 CALL CALCIN,PPER,NUT,£61) 86. 000 60 31 CALL CALCIN,CAPER,NUT,£61) .87.000 61 41 CALL C ALC I N, MGPER ,.NUT , £61 ) 88. 000 62 51 CALL CALC(N,KPER,NUT,£61) 89. 000 63 61 DO 1 K=l,6 90.000 64 NBARKIK,J) =WT I K)*NUTIKI . 91. 000 65 NiOAR< I 7, J )=NBARK( 7, J) tNBARKIK.J) 92.000 66 7 •CONTINUE 93. 000 67 6 : CCNTINUE 94. 000 C SSPEC - DWB1 95.000 68 K=0 - 96.000 69 N = 13 97. 000 70 DO 3 J = 1, 5 98.000 71 CALL c IXI N, J,K, PERCNT ) 99. 000 72 NdllJ)=0W91*PERCNT 1 00. 000 73 8 CONTINUE 101.000 C SSPEC-JW34T 102.000 74 K = 0 103.000 U 75 N = 14 1 04.000 76 • 0 9 J = l ,5 105. 000 77 CALL =IXIN.J.K,PERCNT) 106.000 ' • ~ - • - - — to 73 N34TI J ) = DW34T*PERCNT 107.000 79 9 CCNTINUE 1 C8. 000 C SSPEC-D^T 109.000 80 DO 12 K=l,4 110.000 «» 81 DO 10 J=l ,5 111. 000 82 NT! K, J ) = 0. 0 112.000 • ' " -83 13 CONTINUE 113.000 • 84 12 CCNTINUE 114.000 35 I F( SPECIE.EJ.2.0R.SPECIE.E0.3 IGO TO 1000 115.000 ; 86 N = 14 116.000 87 DO 1H K=l,3 117.000 • 88 56 15 J=1,5 118.000 -89 CALL F IXI N,J, K, PERCNT ) 119.000 • 90 NT[ K, J ) =DWTIK)*PERCNT 120.000 & i 9 j i • j 91 NTI4, J1 = NTI4,JI*NT(K, J l 121.000 92 15 CONTINUE 122.000 93 14 CONTINUE 123.000 94 1000 CONTINUE 124.000 C SSPEC OAN 125.000 95 00 16 K=l,4 ; 126. OOP 96 97 98 17 00 17 J = l,5 KN( K, J 1=0.0 CONTINUE 127. 000 128.000 129.000 99 100 101 16 •CONTINUE IFISPECIE.EQ.2. OR. SPECIE. EO. 3 ) GO TO 1001 H = 17 130. 000 131.000 132.000 102 103 • 104 DO 13 K=l,3 DO 19 J=l,5 CALL PIXIN.J.K, PERCNT 1 133. 000 134.000 135.000 105 'MNK,J) = OWN(K»*PERCNT 136. 000 106 107 19 MMI4, J ) = NN ( 4 » J) +NNI K, J) CONTINUE 137.000 138.000 ioa 109 18 CONTINUE 1001 CONTINUE C SSPEC—'MCO 1 39. 000 140.000 141.000 110 111 112 N = >1 !F( S?ECIE.EQ.2.QR.SPECIE.EQ.3)N=15 K = 0 142. 000 143.000 144. 000 113 114 DO 22 J = 1 , 5 CALL FIX<N,J,K,PERCNT1 145. 000 146.000 115 NCOIJ)=DWCO*PERCNT 147. 000 116 117 22 C CONTINUE SSPEC-OrfTN DO 23 K=l,4 148.000 149.000 1 5U. 000 118 119 120 24 ~ 00 24 J - l i 5 NTNIK,J) =0.0 CONTINUE 151.000 152.000 1 53. 000 121 122 123 23 CONTINUE IF(S°ECIE.EQ.2.0R.SPECIE.EQ.31 GO TO 1002 DO 25 K=l,4 154.000 155.000 1 56. 000 124 125 DO 26 J=l,5 NTNIK,Jl=NT(K,J)*NN(K,J) 157.000 158. 000 126 26 CONTINUE 159. 000 127 128 129 25 1002 CONTINUE GO TO 1003 N = 15 160.000 161.000 162.000 130 131 132 00 27 K=l,3 DO 23 J=l,5 CALL F I X 1 N, J , K, PERC NT I 163.000 164. 000 165. 000 133 134 NTNIK,J)=OWTN(Kl«PERCNT NTNI4,J)=NTN(4,Jl+NTNtK,J» 166.000 167. 000 135 28 CONTINUE 168. 000 136 137 27 CONTINUE 1003 CONTINUE C CALCULATING TOTALS FOR THE TREE 169.OOO 170.000 171.000 133 139 • 0 32 J=l,5 ' TOTAL I J) = NWOOC( 7 , J ) +N BARK (7, J 1+NB1 (J I+NB4TI J )• 1NC0I I)*NTNI 4,J) 172.000 173.000 174.000 140 141 32 C CONTINUE • ••PRINTOUT*** WRITz16, 200)TRE E.SPECIE,L,D8H,CROD,CROL,BUTTL 175.000 17t. 000 177.000 142 200 FORMAT<• 1'//• ','TREE',3X,'SPECIES',3X,'LENGTH' I'WN1, 3 X,1 CROWN' ,3X,•BUTT'/• ' ,1X,•NO.•,5X,•NO.' ,3X,1DBH' , 3X, ,7X,'(MI',4X, 'CRO' , MM)', 178.000 179. 000 13X.'0IA(MI',2X, 'L.(M) ',3X,'L.(Ml•/• • ,1X, I3.6X, 11 ,7X,F4.i,3X ,F5.3, 180.000 ) 13X ,F3.1 ,5X,F4.1 ,3X,F4.1/////' ' , ••••PERCENT NUTRIENT CONTENT BY •, 1•SAMPLE SPEC I F I C A T I O N ' , 7 9 ( • • • ) / ' •,T16,'*•,151•-• 1,•WOOD**SECTlON' 181.000 182.000 \ 9 1*3 1 , 1 S ( ' - ' I , ' * ' , T 7 9 , ' * ' , 1 5 ( ' - , I , 'BARK**SECTI0N',15<'-• 1 ,'*•/• 12( 16X, '1 • ,7X, '3 • ,7X,« 5',7X,'7' ,7X, '9< , 7X, • 1 0' , 7X) / I W R I T E ( 6 , 2 0 1 ) ( N P E R I J I , J = i , 1 2 1 , ( P P E R ( J I , J = l , 1 2 ) , ( C A P E R ! J ) , J = 1 , 1 2 ) , 1 < MGPER! J I , J = l ,1 2) , ( KPFRI J I , J = l ,1 2 I 183.000 184.000 185.000 186. 000 J 201 FORMAT!' •,2( 'NITROGEN -',6(2X,F6.3),6X)/2('PHOSPHORUS-', 16<2X,F6.31.6X )/2('CALCIUM - • , 6 ( 2X, F6. 3 ), 6X)/2( ' MAGNE S IUM - ' , ] 87.000 188. 000 \ 16(2X,F6.3),6X1/2('POTASSIUM - • , 6 ( 2 X , F 6 . 3 ) , 6 X ) ) 189.000 145 WRITE (6 , 202) 190.000 146 202 FORMAT!•-•,T13,2!• 8RANC H•),3X,'*',7('-'),'TWIGS',7!'-'),'*',3X, 191.000 l'*',6('-'),'FOLIAGS',7('-'),'*',3X,'CONES'i3X,'*',3('-'l,'TW!GS + '» 192.000 1•FOLIAGE',' *•/' ', T17, •>1•,5X,•1-1/4',2(3X,•(TOP)',3X,•( MIDI • , 193.000 13X, • ( LOW I • 1 ,12X ,' (TOP) • ,3X,' ( MIDI • ,3X, • (LOW) • / ) 194. 000 147 . tF(S?ECIE.EQ.2.0R.SPECIE.EQ.3l GO TO 1006 195. 000 143 WRIT£(6,2)3)(NPER(JI,J = ) 3 , 2 1 I , ( T N N ( J ) , J = 1 , 3 ) , ( P P E R ( J ) , J = 1 3 , 2 1 ) , 1 ( TN»( J ) , J = l, 3 ) , (CAPER! J l , J = 13 ,21 ) , (TNCAI J 1 , J=1,3I , 196. 000 197.000 ® 1(MGPER(JI,J=13,21 ),(TNMG!J),J=1,3),(KPER(JI,J=13,21I, 199.000 1 ( TN<( J ) , J = l , 3 ) 1 99. 000 149 203 FOR MA T ( 1 ', 'NITROGEN - • , 8 (2 X, F6. 3 ) ,1 X ,4 I 2 X, F6. 3 1 / • PHOSPHORUS-• , 1 8 ( 2X, F6. 3 1, 1X.4I2X, F6 . 3 ) / 'CALCIUM - • , 8( 2 X, F6„ 3) , IX , 4( 2 X ,F 6= 3) / 200.000 201. 000 1 • MAG'IESIUM -',9(2X,F6.3),1 X, 4 ( 2X , F6. 3 1 / • P OT ASS I UM - •, 8 ( 2X, F 6. 3) , 202.000 U X , 4 ( 2X.F6.3I / / / / / I 203.000 @ 150 G") T l ICO5 204. 000 © 151 1006 WRI T? ( 6, 2031 (NPERU 1 , J = 13 ,14) , ( ZEPOI J l , J = l, 3) , 1 ZERO! J) , J = l , 3) , 1 ( N P 5 1 ( J ) , J = l 5 , 1 3 1 , ( P P E R I J ) , J = 1 3 , 1 4 1 , ( Z E R O ! J I , J = l , 3) , H Z = ,=<3 ( J l , J»l, 3) , (PPER ( J 1 , J = 15 ,18 ) , (CAPER! JI »J-13»14) , 205. 000 206.000 207.000 1 ( ZERO! J l , J = l , 3 1 , ( ZERO I J l , J = l, 3) , (CAPER ( J l , J * l 5,181 , 2C8. 000 1(MGPER!J1.J=13 ,14),(ZERO!J),J = l , 3 ) , ( Z E R O ! J) iJ = l,31, 209.000 1 (MGPc R( J ) , J = 1 5 , 1 8 l , ( K P E R ( J ) , J = 1 3 , 1 4 l , ( Z E R O ( J ) , J = l , 3 l , 210.000 1 1 ZERO ( J) , J = l ,3) , (K°ER ( J) , J = 15 ,18) 211.000 . ^EXTENSION* LIMIT OF 5 CONTINUATION CARDS EXCEEDED 152 100 5 CONTINUE 212.000 153 WRITE(3,2 99)TREE,SPECIE.DBH 213.000 154 299 FORMAT(I 3,11,F5.3I 214.000 @ 155 WRIT?(3,300)INPER!J1,J=l,121 215.000 156 157 WRI Tc ( 3, 300) ( PPER1J 1 , J = l , 12 1 WRITE 1 3, 300i (CAPER! J l ,J=1,121 , • vi' 216. 000 217.000 <i 159 WRIT? ( 3,300) (MG PER( J) , J -1 ,12) ' : ' ' 218.000 159 WRIT?(3,3001(KPER!J),J=l,12) > 219.000 160 300 FORMATI12F5.3I 220.000 161 I F ! S'ECI E. EO. 2. OR, SPECIE. EO. 3 1GOT03000 221. 000 162 WRI TE ( 3, 3 011 (NPERI J I, J = 13,20) 222.000 163 WRITS ( 3,3011 < P P E R U 1, J = 13,201 223. 000 164 WRI TS ( 3, 3011 (CAPER! J) , J=13,20l 224.000 165 WRITE(3i301)(MGPER!J),J=13,20) 225.000 166 WRI T? ( 3, 3011 ( K P E R U I, J = 13,20) 226. 000 * 167 301 FORMAT! 8F5.3) 227. 000 163 GOT03 001 228.000 169 3000 WRIT?(3,3 021 ( N P E R ! J I , J = 1 3 , 1 4 ) , ( N P E R I J l , J = 1 6 , 1 8 ) 22 9. 000 170 WRITE! 3, 302) ( P P E R U ) , J = 13,14) ,(PPER(J),J=16,18) 230.000 171 WRITE( 3, 302 ) ( C A P E R I J ) , J = 1 3 , 1 4 ) , ( C A P E R ! J I , J = 16,18) 231. 000 172 WRITE ( 3,3 02) ( MGPER! J) , J= 1 3 ,14 1 , ( MGPER ( J 1, J = 16,1 8 1 232. 000 173 WRITE ( 3, 302) (KPER! J I, J = 13,1 4 l ,( KPER! J) , J=16,18l 233.000 i 174 302 F0RMATI5F5.3I 234. 000 ; 175 3001 CONTINUE . 235. 000 C •**PRINTJUT OF NUTRIENT AMOUNTS*** 236.000 176 WRI T? ( 6 , 2041 237. 000 177 204 FORMAT!' •,* ***NUTRIE NT AMOUNTS IN DECAGRAMS BY SAMPLE ', 238.000 1•SPEC IFICATION' ,74( '* • I / / ' ',T15,'*',16('-'I,'WOOD**SECTI ON' , 239.000 J f ITS 179 lao 118( •-'1.•*•,T80,•*•,16(»-'),'8ARK**SECTI0N',18('-'),'*'/' ',2<15X, l ' U 2 364 566 768 9 10 TOTAL ' l / l 00 2000 J=l,5 WRITE ( 6, 205) CH AR I J ), ( NWOOOI K, J) ,K = 1 ,7 I ,C HAR ( J ) , ( NBARK IK, J ) , 1K = 1,7) 205 FORMAT!• ' , Al 1,1 X ,6 (1 X, F6. 2 1 , IX, F7. 2 , 3X , A 11 ,1 X, 6( 1X, F6. 2 1 ,1X, F7. 2 I 240.000 241.000 242. 000 243.000 244.000 245. 000 N J f 181 132 183 2000 CONTINUE WRITEI6.206) 206 FORMAT! • 0',T13,2!IX,'BRANCH'l.lX.'+'.lOt'-'l.'TWIGS'.ll!'-'),'*', 246. 000 247.000 248. 000 \ 1IX,'*•,91 •-'),'FOLIAGE',10!'-'),•*•,2X,'CONES',IX,'*•,6!' - • ), 1•TWIGS*F0LIAGE',7!•-•1,'*•,5X,'TREE'/• ',T17,'>1",4X,'1-1/4',2X, 12IMT0P) (MID) (LOW)',' TOTAL '),7X, 249.000 250.000 251.000 184 185 IMTQPI ( M i l l (LOW) TOTAL TOTAL'/) DO 2001 J = l , 5 WRI TE (6,207) CHARI J > ,NP.l 1 J) , NB4T ( J I , (NT ( K , J I , K= 1, 4 1 , INN (K , J 1 , K=l, 4 252.000 253. 000 254. 000 186 I ) ,NC:> ( J 1 , (NTNIK, J 1 ,K = 1,4 1 ,TOTAL( J ) 207 FORMAT!' • , Al 1, 511X , F6. 2 ) , IX , F7. 2 , 3 (IX , F6. 2 ), 1X , F7. 2 ,4! IX, F6. 2 ), II X, F7. 2, 2X.F3.2 ) 255. 000 256. 000 257. 000 187 193 189 2001 CONTINUE 1 CONTINUE WRITE(6,203I 258. 000 259. 000 260.000 190 191 192 208 FORHATI'l'l RETURN ENO 261.000 262.000 263.000 193 SUBROUTINE SNPINPER.PPERI C READS IV NUTRIENTS N AND P FROM FILE SOUAM5,ADJUSTS FOR WEIGHT OF SAMPLE 264. 000 , 265.000 194 105 C AND AVERAGES FOR THE TWO SAMPLES TAKEN FPOM EACH SPECIFICATION INTEGER NUM3ER, SSPEC REAL VDS,W0,NPPM,PPPM,Z,NPER(21) ,PPEP.(21I 266. 000 267. 000 268.000 196 197 198 . COMMON NUMBER - ', , DO 1 1=1,21 . NPERI I l = PPER I I 1 =0.0 269.000 270.000 271.000 •EXTENSION* OTHER COMPILERS MAY NOT ALLOW MULTIPLE ASSIGNMENT STATEMENTS 199 1 CONTI N'JE C READS IN 'NUMBER' RECORDS ACCORDING TO SPECIES -272. 000 ., 273.000 • 200 C SPECIES 1 -NUMBER =21,SPECIES 2 -NUMBER = 18 00 2 1=1,NUMBER C READS IN TWO RESULTS FOR EACH SAMPLE SPECIFICATION ANO AVERAGES 274. 000 275.000 276.000 201 202 203 0 3 3 J = l ,2 REATI 1,1001SSPEC,WOS.WD.NPPM,PPPM 100 FORMAT!3X,11,3X,2F7.5.3X,2F6.2) 277. 000 278.000 279.000 204 C ADJUSTS FOR WEIGHT. OF SAMPLE USED Z=WOS-WD ' C CHECKS TO SEE IF THERE IS DATA FOR THIS CARD 280. 000 281. 300 282.000 205 IFIZ.GT.0.01G0 TO 4 206 NPERIIl = PPER(I I=0.0 •EXTENSION* OTHER COMPILERS MAY NOT ALLOW MULTIPLE ASSIGNMENT STATEMENTS 2e3. 000 284.000 207 208 209 • GO TO 3 4 NPHRI I ) = NPER ( I ) + ( (N PPM/10000 ) *104. 20) 11 P PERI I ) =PPER (!)••( ( P PPM/10000 1*104. 20 )/Z 285.000 286.000 2 87. 000 210 211 3 CONTINUE C AVERAGES THE TWO READINGS NPERI I ) = NPER( I I /2 288.000 289.000 290. 000 212 213 Pi'ERI I ) = PPER 1 I 1 /2 C CALLS SUBPROGRAM TO ADJUST FOR MOISTURE CONTENT CALL MCI SSPEC,NPERI III 291.000 292. 000 293. 000 214 215 216 CALL MCI SSPEC,PPERI III 2 • CONTI NUE RETURN 294.000 295. 000 296. 000 0 i ' 217 ENO 297.000 t n o . 218 > 219 C c SUBROUTINE SCAMGKICAPER,MGPER,KPERI READS M NUTRIENTS CA.MG.K FROM S0UAM6,ADJUSTS FOR WEIGHT OF SAMPLE, ANO AVERAGES FOR THE TWO SAMPLES TAKEN FROM EACH SPECIFICATION INTEGER NUM3ER,SSPEC 298.000 299.000 300.000 301. 000 J Qj 0 i I 1 r» • 220 221 222 HEAL CRU,CRUSA,CAPPM,MGPPM,KRE,KPPM,CAPER(21I,MGPER121),KPERI21),Z COMMC'N NUM3ER 00 1 1=1,31 302.000 303. 000 304.000 s 0 i ! «-i r» 22 3 • EXTENSION* • EXTENSION* CAPER ( I I = .MGPER( I 1 = KPER (11 = 0.0 OTHER COMPILERS MAY NOT ALLOW MULTIPLE ASSIGNMENT STATEMENTS OTHER COMPILERS MAY NOT ALLOW MULTIPLE ASSIGNMENT STATEMENTS 305.000 1 <% 224 1 c c CONTINUE READS IN •NUMRER1 RECORDS ACCORDING TO SPECIES -S°ECIES 1 -NUMBER=21;SPECIES 2 -NUMB ER = 18 206.000 3 07. 000 308.000 Q i i i f» 22 5 22 6 c 00 2 I =1 , NUMBER READS IN TWO RESULTS FOR EACH SAMPLE SPECIFICATION AND AVERAGES DO 3 J=l,2 309.000 310. 000 311.000 © I ] <» .22 7 228 100 c RE AO! 2 ,100ISSPEC,CRU,CRUSA,CAPPM,MGPPM,KRE FORMAT!3X,11,3X.2F6.3 ,11X,F6.1,F6.2,F6.11 ADJUSTS FOR WEIGHT OF SAMPLE USED 312.000 313.000 314.000 © 1 f* 229 230 c Z=CRUSA-CRU CHECKS TO SEE IF THERE IS DATA FOR THIS CARD IF! Z.GT.O.OIGO TO 4 315.000 31 6. 000 317.000 e 231 •EXTENSION* •EXTENSION* CADERI I)=MGPER( I )=KPER(I 1=0.0 OTOER COMPILERS MAY NOT ALLOW MULTIPLE ASSIGNMENT STATEMENTS OTHER COMPILERS MAY NOT ALLOW MULTIPLE ASSIGNMENT STATEMENTS 318. 000 : <» 232 23 3 234 4 GO TO 3 CAPER!I)=CAPER(11 •((CAPPM/100001*100.76)/Z MGPER< I 1=MGPER(I)+<(MGPPM/10000)*100.76)/Z 319. 000 320.000 321.000 f 235 236 237 3 KPPM= 3. dl 50+0.4605* KRE KPERI I ) = KPER ( I 1 «•( (KPPM/100001*100. 76l/Z CCNTI NUE 322. 000 323.000 324. 000 233 239 C AVERAGES THE TWO READINGS C AP;R 1 I 1 =CAP ERI 11/2 MGPER( I 1 =MGPER< 1112 325.000 326.000 327. 000 ' 240 241 C KPERI I l = KPER( I 1/2 CALLS SUBPROGRAM TO ADJUST FOR MOISTURE CONTENT CALL MCI SSPEC ,CAPER! I I > 328.000 329. 000 330. 000 • cj» ' 242 243 244 2 CALL MC! SSPEC,MG»ER 1 I 1 1 CALL MC! SSPEC ,KPERI III CONTI NUE 331.000 332. 000 333. 000 t# 245 246 RETURN E NO 334. 000 335. 000 Qi 247 248 249 SUBROUTINE MC1SSPEC,DATAI INTEGER SSPEC REAL DAT A , MCDAT Al 8 1 336.000 337.000 338. 000 y 250 C c COMMON /AREA2/MCDATA . RECEIVES SAMPLE SPECIFICATION AND PERCENT NUTRIENT FOR THAT SPEC. AND ADIUSTS FOR MOISTURE CONTENT USING ARRAY MCDATA 339.000 340. 000 341.000 w • 251 252 253 DATA=DATA*(1-MC0ATA(SSPECI/100. 1 RETURN END 342.000 343.000 344. 000 254 255 SUBROUTINE CALC(N,XPER,NUT,*» REAL XPERI21),NUT(6) 345.000 346. 000 • 256 257 ^ 258 DO 1 1=1,4 NUT(I)=(XPER(N)+XPER(N+1))/2 N = N*1 347.000 348.000 349. 000 • 1 259 1 CONTINUE 350.000 260 NUTI5)=XP5RIN) 351.000 261 NUT(6I=XPERIN*1) 352.000 262 RETURN1 353.000 263 END 354.000 264 SUBROUTINE FI X( N, J, K, PERCNT) 255. 000 265 REAL PERCNT,NPER(21 I,PPER(21),CAPER(211,MGPER(21),KPER(21) 356.000 _266 COMMON. ,/AREAl /._NP_ERjPPJ_P-» CAPER,_MG_P_ER,KP_ER 357. 000 267 GO TO ( 1, 21 3,4,5 ( , J 358. 000 263 1 PERCNT = NPER(K*N1 359.000 269 R-TUR N 360. 000 270 2 PERCNT^PPER I K .N 1 361.000 271 RETUfN 362.000 272 3 P£RCNT = CAPERI K + N! , , 363. 000^ _ 273 RETJRN 364.000 274 4 PERCNT=MGPERIK+Nl 365.000 _27_5 RETURN 366.000 276 5 PERCN T-K P ER I K +N I 367.000 277 RETURN 368. 000 278 ENID • ' 3 69. 000 tDATA 370.000 A P P E N D I X 12 PROGRAM FOR C A L C U L A T I N G WEIGHTED CONCENTRATIONS OF N U T R I E N T S W I T H I N D I F F E R E N T COMPONENTS OF SAMPLED T R E E S P r o g r a m p r e p a r e d b y M r , K e n T w a i t e s t C O M P I L c 1 . 0 0 0 c * « * p * r i G W M T O C A L C U L A T E W E I G H T E D C O N C E N T R A T I O N S F O R N S A M P L E D T R E E S 2 . OOO 1 H E A L 0 3 0 1 ? 0 1 , T T , F T , T F T . T I 0 ) ,F ( 3 I , T F ( 3 ) .WOOD! 1 0 1 , WOODT , BAP.K ( 1 0 ) 3 . 0 0 0 2 R E A L B A R K T , M E A N ( 2 0 ) , N U T C O N < 2 0 , 5 ) , N U T S U M I 6 , 5 ) , N S U M ( 6 , 5 , 2 0 1 / 6 0 0 * 0 . 0 / 4 . 0 0 0 3 P. j ;AL r 3 H 1 5. 0 0 0 _ 4 _ : _ J N T E R_ TR E E 1 2 0 ) . S P E C I E ( 2 0 ) . F L A G ( 2 0 ) . N U M B E R , T R E E l , S P E C 11 6. 0 0 0 • 5 ' I N T C G F . R c f i S S ( 2 0 ) ' 7 . 0 0 0 C R E \ 0 1 4 J c N U M B E R OF T R E E S T O B E P P . O C E S S E D f U P TO 2 0 1 P.. 0 0 0 6 R S A )( 5,< 1 I N _ 9 . 0 0 0 7 ' 9 9 FCP. M\T ( 12 I " 1 C . 0 0 0 8 0 0 I L = 1 . N 1 1 . 0 0 0 T H I S 3 - C T I J N R E A D S I N B I O M A S S D A T A F O R O N E T R E E 1 2 . 0 0 0 9 RL'AOI 3",1 OOTTREE (L I, SPEC IE( L I • O B H ( L 1 , T T , FT ,TFT 1 3 . 0 0 0 1 0 1 0 0 F C n i l I 3 , II ,1 2 X . F S . 3 , 2 8 X , 3 * 6 . 1 I 1 4 . 0 0 0 1 1 _ _ I F ( S>'=CI E (L I . E 0 . 2 . 0 R . SPEC IEIL l . E Q ^ 3 I G O T 0 2 1 5 . 0 0 0 " l 2 R c A >( > ,1 01 I ( T ( I I , P ( 11 ,1=1 , 3 1 ' - - - 1 6 . 0 0 0 1 ? 1 0 1 F O R U T ( 7 X , 2 F 5 , 1 . 6 X . 2 F 6 . 1 , 6 X , 2 F 6 . 1 ) 1 7 . 0 0 0 1 t, G ) T ) l 1 6 . 0 0 0 2 ' R C A O I 3 , 1 3 2 ) ( T F ( I ) ,1 =1 , 3 ) " '"" 1 9 . 0 0 0 1 6 1 0 2 F U R 1 A T U ) X , F 6 . 1 , 1 2 X , F 6 . 1 , 1 2 X . F 6 . 1 I 2 0 , 0 0 0 1 7 3 R E A l t 3 , 1 0 3 I ( WCM 0 ( I I , I =1 , 1 0 1 , W OUOT, ( B A R K ( I K 1 = 1 , l _ 0 l j 3 A R _ K T _ : 2 1 . 0 0 0 Q 1 3 ' 1 0 ? ' F U R I A T ( 7 X , 1 Q F 6 . 1 , F9.1 / 7 X , 1 0 F 6 . 1 , F 9 . i I " " "" ' " •" * " 2 2 . 0 0 0 " ~~~ ~ C T H I S S E C T I O N W R I T E S T H E BIOMASS D A T A 2 3 . 0 0 0 -.;/ ,:• J 9 WRI TJ I 6 , 2 0 0 1 ' 2 4 . 0 0 0 i^Lui!^-2 0 2 0 0 "FOR 1A" f (" • 1 • , • B 1 0 M A S S D A T A ' / / ) 2 5 . 0 0 0 i « ^ s f " 2 1 WR IT.11^,231 I T R E E ( L ) , S P E C I E I L ) , D B H I L I 2 6 . 0 0 0 2 2 2 0 ! F C R I A M * ' , ' T R E E ', 1 3 , 5 X , ' S P E C I E ' , 1 1 , 5 X , ' D B H • , F 5 . 3 / / I 2 7 . 0 0 0 2 3 ' ' W I r i * ( b , 2 12 I ( WOOD ( I I , I =1 ,1 0 ) ,'WOCDT , ( B A R K ("I I V I = 1~»1 0 1 , 3 A R K T • ~ " " Z f t . 0 0 0 2 4 2 0 ? F t R nr ( • ' , O X , 1 1' , 6 X , ' 2 1 , 6 X , • 3 ' , 6 X , , 6 X , • 5* , 6 X , ' 6 1 , 6 X , '7« , 6 X , ' 8 ' 2 9 . 0 0 0 1 , S X , • T , r>X, • i 3' , 6 X , ' S U M ' / • WOOD • , 1 0 ( F 6 . 1 » ! . X I , F 9 . 1 / ' B A R K •, 2 0 . 0 0 0 • i i - . i i =.s". r . i x i , f 9 . rvi 3 i . w 2 5 I c ( i ° E C I E ( L ) . E 0 . 2 . 0 R . S P E C I E I L l . E O . 3 I G 0 T 0 4 3 2 . 0 0 0 2 6 I T - ( 5 , I J 3 > ( T ( I I , I = 1 , ? ) , T T , ( F ( I I , I = 1 , 3 I , F T 3 3 . 0 0 0 2 7 F j v u n • ' , =IX, • 1' , 6 X , >i< , 6 X , ' 3 " , 5 X , • S U M 1 / /' T W 1 G S ' « 4 T F 6 7 1 7 1 X 1 7 3 4 . 0 0 0 l ' F O L ' » 4 ( F j , l , l X I / / / l 3 5 . 0 0 0 2 * G O T l n • 3 6 . 0 0 0 2- i 4 WRI r-: ( 6 , 2 0 " 4 i " ( T F ( T | ' , 1= 173"! , T F T 3 7 7 0 0 0 " 3 0 2 0 4 F C R I A T I 1 • , 3 X , • 1 • , 6 X, • 2 • , 6 X ,' 3 • , 5 X , • S U M • / / • T . F ' , 4 ( F 6 . 1 , 1 X 1 / / / ) 3 8 . 0 0 0 C T H I S S E C T I O N G I V E S P E R C E N T P E R S E C T I O N 3 9 . 0 0 0 3 1 5 0 0 S 1 = 1 , 4 " 4 0 . 0 0 0 3 2 MEA J( I ) = ( WC 0 0 ( 2 * 1 - 1 ) * WOOD ( 2 * 1 ) I / WOODT 4 1 . 0 0 0 3 3 M>'A J( I * 6 I = < R A R K ( 2 * 1 - 1 ) V P A R K ( 2 * I I l / B A P K T 4 2 . 0 0 0 " 3 4 6 C ' l N T I N U E 4"37~0"0"0~ 3 5 M H A I l 5 ) = W r i n O ( 9 ) /WOODT 4 4 . OOC 3 6 Md-V4-( t ) = w r , 0 1 ( 1 0 1 / w n c i D T _ 4 5 . 0 0 0 3 7 " I L V I I l l l = 3 A R < ( 9 l / B A R K T " - - - - ; 4 6 . 0 0 0 3 3 M E A N l 1 2 1 =1'ARK I I 0 ) / B A R K T 4 7 . 0 0 0 C M E A N g.J't 1 3 A N O 1 4 (_QR 1, P.R 4 ) = 1 . 0 4 R . 0 0 0 3 9 '1 •;A "•)("! 3T^E^TnT4"r="r7u : 4~<?7iOTJ"a-C T H I S S E C T I O N C A L C U L A T E S T H E P E R C E N T PER S E C T I O N F O R TWIG,FOL,OR T + F 5 0 . 0 0 0 C I T U S P S A F L A G W H I C H , I F E Q U A L T O 1 , M E A N S T 8 E T R E E WAS S A M P L E D F O R T H E S E 5 1 . 0 0 0 ' " ' " C S P C C I - I C \ f I ON 5 • I F F L A G = 0 , TREE ' ")CI.E "CNL'Y 'WAS ' SAMPLED "SO' MEANSOVO " 5 2 . 0 0 0 • E X T E N S I O N * O T H E R C O * ° I L E R S M A Y N O T A L L O W M U L T I P L E A S S I G N M E N T S T A T E M E N T S 4 0 F L A M L 1 = 1 5 3 . 0 0 0 •"41" ~ " I F ( -,.-> = C IF. ( " L 1 7 F 0 7 " 2 7 0 R . ~ S P E C " T E T L " l T r O T 3 T T ; D T T l 7 : 5 4 T 0 0 0 , 4 2 [ R ( T f , !: 0 . 0 . 0 , 0"- • F T.» E - 0 . 0 . 0 ) GU TOR 5 5 . 0 0 0 . \ - > ) 4 3 - 0 0 » 1 = 1 5 , 1 7 5 6 . 0 0 0 * 'i K j 4 4 Mr-A |( I | = T ( !-! 41 / T T ' " . " ' 5 7 . 0 0 0 " " ' - " " " ' I 4 5 9 C O N T I N U r 5 P. 0 0 0 ; | 4 6 0 0 10 1= 1 8 , 2 0 • ' " • . 5 9 . 0 0 0 ' J c 47 MEAN! I l = F< 1-17) /FT 60. 000 \ 48 CONTINUE 61.000 49 GOTO!1 62.000 50 8 FLAJ(LI=0 63. 000 51 DO 12 1=15,20 64. 000 52 MEAN! I 1 = 0.0 • 6 5 . 000 J > 53 12 CONTINUE • 66. 000 \ 54 GOT .111 67. 000 55 7 I Ft T=T .E0 .O . 01GOT03 6 8 . 0 0 0 56 DO 13 1=15,17 6 9 . 0 0 0 57 MEAN! I l = TF( 1-14 l/TFT 70.000 58 13 CONTI NUE 7 1 . 0 0 0 C THIS SECTION WRITES THE PERCENT PER SECTION 72. 000 59 11 WRITE ( 6 , 2 051 73. 000 60 205 FORMAT!•0','PERCENT PER SECTION') 74. 000 61 WRITE( 6 , 2 0 6 1 TREE!L).SPEC IE(L),DBH!LI,FL AG(L) 75. 000 62 206 FORMAT!' ','TREE 13,5X,•SPECIE ',I1,5X,'0BH • ,F5.3,5X,•FLAG=•, 7 6 . 0 0 0 111//) 77 . 000 63 WRI TE 16, 207)(PEAN! I ), 1=1,14) 78. 000 64 207 FORMA T( ' • , 7 X , • U2' , 4 X , ' 3 £ 4 ' ,4X, ' 5C6' ,4 X, '7te' , 5X ,'9' ,5X,« 10'// 7 9 . 0 0 0 l 'WJOO ' . 6 I F 6 . 3 , lXI/ ' B A R K • , 6 ! F6.3, 1X1//'BR1 • , F6. 3 , 3X, ' RR4 ', 8 0 . 0 0 0 1 Ft. 3/1 . 8 1 . 000 65 IF! S'ECIEIL ). EQ.2.OP. SPEC IE! L ).EQ. 31G0T014 82.000 66 WRITE ( 6 , 2081 (ME AN ( I 1, 1 = 15,20) 8 3 . 0 0 0 67 208 FORiATP ' ,8X ,' 1 • , 6 X , '2 • , 6 X , '3'//'TWIGS ' , 3 ( F6. 3 ,1 X1 / ' FOL ', 6 4 . 0 0 0 1 3 ( P 6. 3 , 1 X 1 / 1 8 5 . 0 0 0 68 GUT )!. 5 e6.000 69 14 WRI TEI 6 ,209)1 ME AN!I 1,1=15,171 87. 000 70 209 FORMAT!' •, 3X, ' 1' ,6X, ' 2 ' , 6 X , ' 3'//• T + F •, 3 ( F 6 . 3 ,1 X ) / ) 88.000 C RE»D IN.OF NUTRIENT CONTENT DATA FROM PEPNUT FILES INTO ARRAY NUTCONI20,51 89. 000 71 15 REAOI1,104)TREE1,SPECI1,OBH1 9 0 . 0 0 0 72 104 FGSM\T(I 3 ,11,F 5 . 3I 91 . 0 0 0 C CHECK TO SEE IF NUTCCN TREE SAME AS BIOMASS TREE 9 2 . 0 0 0 73 IF! R E E l L I . E Q . T R E E 1 IGO TO 16 93.000 74 WRITE(6,210ITREE(L),TREE1 94.000 75 210 FCRIATI'l','TREES 3EING PROCESSED NOT S AM E ' / • BI OM ASS TREE=',I3/ 95. 000 1 ' NIR 13NT TREE= • , 13 1 9fc.000 76 RETURN 9 7 . 0 0 0 77 16 DO 17 J = l,5 - 9 8 . 0 0 0 78 READ!1,1051(NUTCON!I,J),I=1,121 9 9 . 0 0 0 79 105 FCRMATU2F5.3I -• 100 .000 80 17 CCNTI NUE 101 .000 81 NUM3£R=20 102.000 82 IFI SPECIE! LI. EQ.2-.0R.SPECIECL 1. EQ. 3 )NUMBER = 17 103. 000 83 DO 13 J=l,5 104.000 84 "REAOI1,106)(NUTCON!I,J),1=13,NUMBERI 105 .000 . 8 5 106 • FnRM\T(3F5.3) 106. 000 • 86 18. • CCNTI NUE ; • * • . ' " • • • • . , . 107.000 C THIS SECTION WRITES THE NUTCON DATA . 103 .000 87 WRI r. (6,2111 1 09 . 000 83 211 FOR •IA T 1 ' - •, ' NUTRI EN T CONTENT DATA'/) 110 .000 89 " " WRIT1(6,2121TREE1,SPECI1,03H1,FLAG(L1 1 1 1 . 0 0 0 90 212 FCRMUC ' , ' T R E E •, 13 , 5 X , « SPECIE ',I1,5X , 'DBH « , F 5. 3, 5X, • FL AG= ' , 112. 000 111//) 113.000 91 WRIT;i6,213> J 1 4 . 0 0 0 92 213 . . F'IRMA T( • • ,T ! 6, •*• , 15 ( •-• ) , ' WOOD** SECT I ON • ,15 (•-• J , ' *• , T75, •*•, 1 1 5 . 0 0 0 1151 •-' I, 'OAR K** SECT I 0 N • , 1 5 ( • - • ) , ' * • / , ' ,21 16X,'l',7Xi'3" ,7X,'5' , 116 .000 1 7 <, < 7 • ,7X , ' 9 ' , 7 X , ' 1 0 ' ,1X1/1 ' " 117. 000 ~ " 93 .DO 19 J=l,5 " 118.000 94 WRI T^  1 6 , 2 1 4 ) J , 1 NUTCON! I, J ) ,1 = 1,12) : 119 .000 95 214 FORMAT!' ', 11 , • - < ,9 X, 6! 2 X, F6. 31 ,11 X, 61 2 X, F6. 3 II 120.000 96 19 CONTINUE 121.000 97 IF(SPECIEIL). EQ.2.0R. SPECIEIL I. EO. 31 GOT 02 0 122.000 93 WRIT:(6,21.51 123.000 99 215 FORMAT (•-• ,T1 3, 2( • BR ANC H • I , 3X, • * •, 7( '-• I , • TWI GS ' , 7( • - • I , •« • , 3X, 124.000 1 ' «' ,S ( '-' I ,' FCL IAGE ' ,7C-' I . • *'/' ' , T1 7 ,' >1 1 , 5X ,' 1-1 /4', 125. 000 1 2 ( 3<, • ( TOP I ' , 3X , • IMIi) 1 •, 3X, ' ( LOWI • I/) 126.000 100 00 21 J = l , 5 127.000 101 _____ ; WRI H I 6, 2161 J , ( NUTCON ( I, J I ,1=13,201 : 128.000 102 216 FORMAT!' • , 11 , • - • , 9 X, 8 (2 X, F6.31 1 ' "129.000 103 21 •'• CONTINUE 130.000 104 G IT 122 _____ . ; 131. 000 105 20 WRITE(6,217) 132.000 106 217 FC=H\T (• -• , Tl 3,2 ( • BRANCH'I,3X,'*',3('-' 1,'TWIGS + FOLIAGE *'/ 133. 000 1' '. Tl 7, •>!•, 5X ,'1-1/4', 3X, •( TOP) • ,3X, • (MIDI ' ,3X, •( LOW) •/) 134.000 107 00 2, J = l,5 " 135. 000 103 WRI TE ( 6, 213) J , ( NUTCON (I,J ) ,1 = 13,171 136. 000 109 218 • FORMAT!' ' . 11 , ' - ' , 9 X, 5 [ 2X , F6. 31 I 13 7. OOP 110 23 CONTINUE 1 38. 000' C THIS SECTION CALCULATES THE WEIGHTED CONCENTRATIONS (WEIGHTED MEAN X CONC.I 139.000 1U __22 NUM>-P. = 20 _ _ . 140.000 112" " fF"VSP'=ClElLi.E0i'2\OR.SPECl'E(LI.EQ.3'lNUMBER = f7 141.000" 113 00 24 .1=1,5 142.000 114 00 >> 1 = 1, NUMBER ; 143. 00 0 115 NUTCUN (,I ,J)=NUTCON! I, JI * ME AN( I I 144. 000 116 25 ' . CONTINUE 145. 000 l_7 24_ _CONriN'JE . • . - 146.000 ~ : C THIS .'SECTION WRITES" THE WEIGHTEO CONCE NT RAT "IONS-; : ' 14 7. 000" 113. WRITS (6,2191 > 143. 000 1]Q 219 FORMAT!' 1','WEIGHTED-CONCENTRATIONS'I 1 49. 000 120 . WRI f i ( 6,2121 TREE1.SPECI1, DBH1 , E L AG ( L I ~ 1'5"0.000 121 . WRITE!6,2131 ]51.000 122 DO 26 J = l,5__- _ _ _• 1 52. 000 123 WRI TS ( 6, 2141 j,"( NUTCON ( I i J I »1 -1»1 2T '. : 1 53. 000 124 26 CONTINUE 1 54. 000 125 J_F( >*rS:l.E 1 L » ' E0.2.HR. SPEC IE(L I. EQ. 3) G0T027 1 55. 000 126 WRI'Tc ( 6, 2T'5T7~ ~ ~ : " i 156.000 127 00 23 J=l,5 ' .:-"' -' 157.000 129_ WRI Ti 16,216)Jj,(NUTC0NJI,J^,1 = 13,20) 158.000 129" 28 "CONTINUE ' "" "" " ' ' ] 15^.000 130 G0T029 160.000 131 27 WRl IE ( S,21 7J 161.000 132 DO 11 J = 1T5 ~~ : T62.00TJ 133 WRITE(6,2131J,(NUTCON(I,J 1,1 = 13, 171 163.000 134 30 CONTINUE 164.000 Cn,HIS'"SECri0N~SUMS~W30D7"~BSRK7""TffI"GS7""F"0L; C1R TWTCS+FOL SEPARATELY 165. 000 135 29 00 Jl 1 = 1 ,6 166.000 136 DO 32 J = l ,5 . . . ' 167. 000 TV? N j T s i m i , J l = 3.0 ' ~ : TE"8TTJ0r0 139 32 CONTINUE 169. 000 139 31 CONTINUE 170.000 !V0"" ~00*'33 J=175 " 171.000"" 14! DO 34 1 = 1 ,6 1 72.000 142 NUT S J M! 1 , J|=NUTSUM( 1, Jt+NUTCON! 1 , J I 173.000 T41 TTOT SIMT27J I^IUrSTfMI 2, J )•NUTCON! I +6 , J I T74T0D0 144 34 CONTINUE 1 75. 000 145 NUT 5 IM( ? ,.I) = NUTSUM( 3, J 1 + NUTCO N( 13 , J) 176.000 _<,_-- • - : NUT ;JM(4,J1 = NUT SUM!4,JI*NUTCON(14» JI T77.000' 147 33 CONTINUE 178.000 148 -\Ft S 3ECIE(L I.E0.2.0P. SPEC IEIL I.EQ. 3IG0T035 179.000 f 149 150 151 152 DO 36 J = l,5 DO 37 1 = 15,17 NUT3JMI 5,J) = NUTSUM(5,Jl+NUTCGN!I,J) NUTSJMI6,J|=NUTSUM(6,J>*NUTC0N(I+3,JI . 180.000 181.000 182.000 183. 000 153 37 CONTINUE 184.000 V—-—- 154 36 CONTINUE 185. 0 0 0 f 155 . GOT 13 3 1 8 6 . 0 0 0 : < 15& 157 35 • DO 3 9 J=l,5 00 4 0 1=15,17 187.000 188.000 15B NUT i i M ( 5 , J 1 = Ni)T SUM! 5 , J I + NUTCON! I , J ) 189. 0 0 0 159 40 CfNTINUE 190.000 160 39 CONTINUE 191. 0 0 0 C THIS SECTION GIVES THE TREE A DBH CLASS:CLASS ES 1-11 = 6-10",10-14",. ..,46"- 1 192.000 161 162 38 )=O.H(L> CALL CL\SSR.(n,M) 193.000 194. 0 0 0 163 CLA~,3(Lt = M 195.000 C THIS SECTION WRITES THE SUMMED WEIGHTED CONCENTRATIONS 196. 0 0 0 164 WRI TE ( 6, 2201 1 97. 0 0 0 165 220 .FORMAT!'-",'SUMMED WEIGHTED CONCENTRATIONS BY SPECIFICATIONS') 198.000 166 WRI TF. (6,221 ) TREE1 ,SPECU ,DBH1 , FLAG! L ), CLASS (L ) 199.000 -167 _221.__ F O R UTI '0' , 'TREE • , 1 3 , 5X,'SPECIE ' »11,5X,'DBH • ,F5.3 ,5X,•FLAG=' 200. 000 i l l , 3 X , ' O B H CLASS=', Ji/7) 201.000 163 I F ( JPFCI E( L) . E0.2.0R. SPEC IE(L ) . EO. 3 1 G0T04 1 202.000 169 WRITE(6,2;2) 203. 000 170 222 FGRU'TC ",4X,'waOO ",3X,'8ARK ' , 3 X , " BR 1 " , 3 X , ' BR4 ',3X, 204.000 171 1•TW13S',3X,• FOL • / 1 DO 4 2 J=1 , 5 205. 000 206.000 172 WRITE(6, 223 1 J,INUTSUM(I,J 1 , 1 = 1,6) 207. 000 173 223 FOR n i l " ",I1, •- • , 6 ( F 5 . 3 , 3 X I ) . 2Ce. 000 174 42 • CONTINUE 209.000 175 176 41 GOT 14 J . WRI T r ( 6 , 2 2 4 ) 210. 000 211. 000 177 224 FORMAT!• " , 4 X , ' W 0 0 0 ",3X,"BARK ',3X,' BR1 ',3X,' BR4 ',3X, 212.000 • 178 1 ' T*F •/ I DO. 4 4 J = l , 5 "" 213.000 214.000 179 WRI TE ( 6, 2251J, < NUTSUM < I, J I , 1 = 1,5) 215.000 190 225 FORMAT(• ',11,'- ',5(F5.3,3XII 216.000 181 , 4 4 CONTI NUE ••• ' • 217.000 C THIS SECTION i>UTS THE ARRAY NUTSUM INTO A FINAL ARRAY INSUM1 FOR ALL TREES 218.000 182 4 3 DO ,b • J=l,5 ""219.000 ie3 0 0 4 6 .1 = 1 ,6 220. 000 194 NSJM I , J , L ) = NUTSUM( I , J) 221.000 . 185 46 CONTINUE 222.000 186 45 CONTINUE 223.000 187 1 CONTINUE 224.000 C END O F PROCESSING FOR THE N TREES "" 225. 000 C INFORMATION IS IN THE F OL LOW I NG ARRAYS : TREE! L 1 , SPECI E ( L 1 , OBH(L), ClASSILI, 226.000 C FLAG IL1, NSUM!I,J,L); L = TP EE PROCESSING ORDER; I=1-6(SPECIFICAT IONSI; 227.000 C J=l -5INJTRIENTS1; CLASS IS ONE OF 11 DBH CLASSES; FLAG=1 IF TPEE COMPLETELY 22H.O0O C SAMPLED, F L A 3 = 0 IF TREE BOLE ONLY SAMPLED 229.000 188 WRITE(t>, 226> 230.000 189 226 FORMAT('1'1• 231. 000 190 WRITE(6,2201 232. 000 191 DO 5 0 L=1,N 233. 000 192 WRITE(o,221ITREE(L),SPECIEIL) ,OtiH( L) , FL AG ( L 1 , CL AS S ( L ) 234.000 193 IF(S»ECIEIL).EQ.2.0R.SPEC IE(LI.EQ.3IG0TO51 235.000 194 WRITE(6,222) 236.000 195 DO 52 J = 1,5 ~ ' " 237.000 196 WRITE(6,2 23)J,<NSUH(I,J,L),I=1,6) 238.000 197 52 CONTINUE 239. 000 o o I o •9 3 •J r 198 GQT05 3 2 4 0 . 0 0 0 \ 199 51 WRITE(6 ,224) 2 4 1 . 0 0 0 200 DO 54 J = l , 5 242. 000 201 WR!TE(6 , 2 2 5 I J , ( N S U M ( I , J , L ) . I = 1 . 5 I 2 4 3 . 0 0 0 202 54 CONTI NUE 2 44. 000 \ 203 5 3 CONTINUE 2 4 5 . 000 ) 204 50 CONTINUE ' . 2 4 6 . 0 0 0 < 205 WRITE(6,2271 2 4 7 . 000 206 227 f n r U T I ' l ' l 2 4 8 . 0 0 0 C PUT AR R A / NSUM ON F ILE 2 4 9 . 0 0 0 207 DO 55 L=l ,N 250. 000 203 WRI TE ( 2 . 2 50) TRtE (L) .SPEC IE (LI ,OBH( L) ,F L AG ( L ) , CL AS S ( L 1 2 5 1 . 0 0 0 209 250 FORMAT(I 3 , 1 1 , F 5 . 3 , 1 1 , 111 2 5 2 . 0 0 0 210 DO 56 J = l , 5 2 5 3 . 000 211 WRI TE ( 2 , 251 ) ( NSUM( I , J , L ) , 1 =1,61 2 5 4 . 0 0 0 212 251 FOR Mi T(6 F5•3 I 2 5 5 . 0 0 0 213 56 CONTINUE 256. 000 214 55 CONTI NUE 2 5 7 . 0 0 0 215 RET H ) 25B. 000 216 E NO 2 5 9 . 0 0 0 217 SUBROUTINE CLASSR(DBH,CLASS 1 2 6 0 . 0 0 0 C SUBROUTINE TO CLASSIFY OBH INTO ONE OF 11 G R O U P S ( 6 - 1 0 " , 1 0 - 1 4 " , . . . , 4 2 - 4 6 " , 46" -1 2 6 1 . 0 0 0 218 ' REAL TBH.C/O. 0254/ 2 6 2 . 0 0 0 219 I NT;jER C L A S S , J 2 6 3 . 0 0 0 220 CLASS =1 2 6 4 . 0 0 0 221 J=6 2 6 5 . 000 22 2 IF0J3M.LT. ( J = CI IRETURN 2 6 6 . 000 ; ; . 2 2 3 . on i 1 = 1 , 1 0 2 6 7 . 0 0 0 224 CLASS = 1 2 6 8 . 000 . 225 IF( J H.GE . I J * C I . ANO. DBH.LT. (( J + 4I*CI IGOT02 269.COO 22 6 J = J*4 2 7 0 . 000 22 7 1 • ' CONTI NUE 2 7 1 . 000 22 8 CLA3S=11 2 7 2 . 0 0 0 229 2 RETURN 2 7 3 . 0 0 0 230 FN ) 2 74. 000 $OATA 2 7 5 . 0 0 0 L_ A P P E N D I X 13 ROGRAM FOR C A L C U L A T I N G BIOMASS FOR 2 5 6 T R E E S ON THE SQUAMISH SAMPLE PLOT ACCORDING TO L O G A R I T H M I C R E G R E S S I O N EQUATIONS SCOMPILE C BIOMASS DISTRIBUTION FOR 256 TREES FROM C NEAR SQUAMISH 1 CHARACTER»3 TRE HIGH ELEVATION SAMPLE PLOT 2 I N T E G E R SP 3 R E A L L _ _ a HW00D=H8ARKsHDwBl »HDWBt|aHTWJJ3«HFOL IBHTWFPSHSTRU • E X T E N S I O N * OTHER C O M P I L E R S MAV NOT ALLOW M U L T I P L E ASSIGNMENT • E X T E N S I O N * OTHER C O M P I L E R S MAY NOT ALLOW M U L T I P L E ASSIGNMENT • E X T E N S I O N * OTHER C O M P I L E R S MAY NOT ALLOW M U L T I P L E ASSIGNMENT "H3CRO*H3TRE"0,0 STATEMENTS STATEMENTS STATEMENTS • E X T E N S I O N * • E X T E N S I O N * • E X T E N S I O N * • E X T E N S I O N * • E X T E N S I O N * • E X T E N S I O N * 5 • E X T E N S I O N * • E X T E N S I O N * • E X T E N S I O N * • E X T E N S I O N * _ * E X T E N S I O N « _ • E X T E N S I O N * • E X T E N S I O N * • E X T E N S I O N * ' " E X T E N S I O N * 6 7 OTHER COMPILERS MAY NOT ALLOW MULTIPLE ASSIGNMENT STATEMENTS OTHER COMPILERS MAY NOT ALLOW MULTIPLE ASSIGNMENT STATEMENTS OTHER COMPILERS MAY NOTJLLOW MULTIPLE ASSIGNMENT STATEMENTS OTHER COMPILERS MAY"NOT ALLOW-MULT I PLE ASSIGNMENT"ST A TtMENT S OTHER COMPILER3 MAY NOT ALLOW MULTIPLE ASSIGNMENT STATEMENTS OTHER COMPILERS MAY NOT ALLOW MULTIPLE ASSIGNMENT STATEMENTS AwOOD = ABARK = AnwHl«ADwBllBATWIG3AFOLI«AT»«FO = ASTHU: OTHER COMPILERS MAY NOT ALLOW MULTIPLE ASSIGNMENT OTHER COMPILERS MAY NOT ALLOW MULTIPLE ASSIGNMENT OTHER COMPILERS MAY NOT ALLOW MULT IPLE ASSIGNMENT OTHER COMPILERS MAY NOT ALLOW MULTIPLE ASSIGNMENT _OTHER COMPILERS MAY NOT ALLOW MULTIPLE ASSIGNMENT 0fH£R COMPILERS MAY NOT ALLOW MULTIPLE ASSIGNMENT OTHER COMPILERS MAY NOT ALLOW MULTIPLE ASSIGNMENT OTHER COMPILERS MAY NOT ALLOW MULTIPLEASSIGNMENT OTHER COMPILERS MAY NOT ALLOW MULTIPLE ASSIGNMENT ILINEsO DO 1 1*1,256 A"SCH"O = ASTHE»O70r STATEMENTS STATEMENTS STATEMENTS STATEMENTS STATEMENTS STAT EMtNTS STATEMENTS STATEMENTS STATEMENTS 8 9 10 1 i " 12 13 TC" 15 "16"" 17 16 T»~ 20 21 "22 23 2a ~?:v 26 27 28 29 30 " J l " 32 33 "3<r 35 36 "37" 38 HE AD(6,11) TRE,SP,0,H,BUTL,CW1,CW2 11 FORMAT (A3, I I , 3<F4, 1),4X,F2,0,F2,0> Cws(CC*UCW2)/2,01*0,3048 _ _ " " 0*6,0254*0 " CL*H"BUTL BAa(3,H159«0«D) /a,0 • ' IFCSP.EQ . i ) GO TO 21 " ~ ~ ~ IF (SP.EQ.2) GO TO 22 C 8PI«HEML0CK,SP2=ABIFS "21""" B W 0 0 D s 2, 319 5 • 0 ,7 4 5 5 * A L"0G10 (D"«D *"H) "" BBARK33,1089*1,0388«AL0G10(BA) BDWB1=0.6467+0.7853*ALOG10(D*0*H*CW) "D'WB«s I,90'ii6 + 0,2089*(BA»H*CW*CLJ ' BTWFOaO,8851+l.5576*ALOG10(D*0*M)*0, BTWIG=-1.2724*1,1630«ALOG10(D*CL*CL) BFOLI=0,5980*|,7174*ALOGI0(D*D»H)»0, WOODslO,0«*Bk.OOD BARK = 10,0»*B!IARK "DwB'l = 1 0", 0 •"•BDWB'l :  TwFO=iO,0«*BTWFO TWIG=IO,0*»RT*IG P O L I " 1 0 , 0 * » 3 F O L I 7 " STRU=WOOO*BARK SCROiOoBl *nWH4*TWFO ' """' S T H E = 3 T R l J t S C R ' 0 : ~ HnOOD=nWOOD+"000 .HBARKSHBAKK+HARK HDwBlsHDwBl+DWfll ' HDWB4 = H0».B4 + DWB4 HT.WIG = HTMG + TWIG "TIF'OC'rBHFOLIVro'CI HTWFOsHUFO + TWFO 0362*D*D*H 0U47*D«D*H 5.000 6,000 7,000 8,000 9,000 10.000 11.000 12,000 13,000 14.000 15,000 16,000 17,000_ "18,000 19.000 _20 , 000_ ?:\~, ooo 22,000 23,000 24,000" 25,000 26,000 "27"; 0 00"" 28,000 29,000 30.000 31,000 32.000 "3370 0 0" 34.000 35.000 "36.000" 37,000 38,000 "3'9700"0"" 40.000 41.000 "42 , 000 43,000 44.000 "457000" 46,000 39 40 41 42 H3TRU»HSTRU*3TRU HSCR03HSCRO+3CRO HSTREsHSTRE+STRE GO TO 29 4 7 , 0 0 0 4 8 , 0 0 0 4 9 , 0 0 0 / 43 2 2 BW00D=2,0472*0,9S26*AL0G10(D*D«H) 51,000 s 44 BBARK=3,0965+1,3268*ALOG10(BA) 52,000 45 B0*B1 = 1 , 2 6 8 8 * 0 ,0525*0 * C L 53.000 © 46 BDv.B4 = 0.6615 + 0,7597*ALOG10(D*CL) 54,000 47 BT»F0=0,8787+1,0376*ALOG10(D »CL ) 55,000 48 WOOD=10,0**BwOOD 56,000 49 BARK=10,0**RHARK 57,000 SO DHB1*10,0*»BDWB1 5B.OO0 51 D*HUal0.0*»BDwB4 59.000 © 52 T*FO=10,0*«BTWFO 60,000 53 TWIGaO.O 61,000 54 FOLI=0,0 62,000 o 5 5 STRUsWOOD+BARK 63.000 56 SCROsQ».Fil+DWB4 + TWF0 64.000 57 STRE=STRU+3CR0 65,000 o 5 8 A*0O0=AK0OD+w0OD 66,000 5 9 A8ARK=A8ARK+BARK 67,000 60 ADwBlsAOWfll+OWBl 66.000 © 61 AD*B4aADI»B4*DWB4 69,000 62 ATWIGSATKIG+TWIG 70,000 63 AFOLIaAFOLI+FOLl 71.000 o 64 ATWFOsAUFO + TWFO 72,000 65 A3TRU=ASTRU+STRU 73.000 66 ASCRO=ASCRO+SCRO 74,000 o 67 A3THESASTRE+3TRE 75.000 68 GO TO 29 76.000 C PRINT AUT RESULTS FOR INDIVIDUAL TREE3 77,000 o 69 29 I F d . E U . l ) GO TO 3i 78,000 70 GO TO 33 79,000 71 31 WRITEC6.32) 80,000 o 72 ~~32 F ORHAT (11', 'TREE 1X, < SP» , IX,'DBH !,?)(,'LENGTH l , lX,"CRLEN6THl, IX, 61,000 1"CRD IAl,4X,tWOOD',OX,'BARK',2X,'BIG BRAN',1X, '3 M BRAN',2X, ' TWIG', 82,000 23X, 'FOLIAGE', IX, 'TW+FOL' ,2X, 'TREE BOLE',2X,'CROWN',4X,'SUM TREE'/ 63,000 o 3T10, ' (M) ' ,T16, ' (M) ' ,T23, ' (M) ' ,T29, ' (M) ' ,T37 i ' (KG) 'VT4S, ' (KG) '",T53» 84,000 4 ' ( K G ) ' , T 6 2 , ' (KG) ' , T 6 9 , ' ( K G ) ' , T 7 7 , • ( K G ) ' , T 8 5 , ' C K G ) ' , T 9 4 , I ( K G ) ' , 85,000 ST104, ' (KG)•,T114, ' (KG) ' / / ) 86,000 o 73 33 WRITE (6,34) TRE, SP, D, H, C L , CW, WOOD"; BARK , DWB1, DWU4, T w IG, F O L I , T WFO, 8 7,000 1STRU,SCR0,STRE 88,000 74 34 FORMAT ( ' 0 ' , A3 ,2X , I I , 1X ,F5 ,3 ,3 (2X , F 4 , 1 ) , 3X ,7 ( F B , 1 ) , 3 (F10 ,1 ) ) 89,000 tf 75 ILINEsILINE+1 90,000 76 I F ( I L I N E , E 0 , 2 7 ) GO TO 40 91 ,000 77 UO TO 1 92.000 7B 40 WRITfcl6,3<iJ 93,000 79 ILINE=0 94,000 80 1 CONTINUE 95,000 81 T>OOD = HWOOD + AWOOD 96,000 8 2 TBARKSHBARK+ABARK 97,000 83 TOWBlaHD»Bl+ADWBl 98,000 8 4 T0*BUaHD>.B4 + ADWB4 99,000 8 5 ' TTWIGaHUIG + ATWlG 100,000 86 TFOLI=HFOLItAFOLI 101,000 87 TTxF0=HT«F0+AT«F0 1 02 , 000 """ ' " 6 8 T3TRU=HSTRU+ASTRU 103,000 89 TSCROsHSCRO+ASCRO 104,000 J o 9 0 TSTRE=HSIRL+ASTKf 105,000 C PRINT OUT SUMARY FOR SAMPLE PLOT 106,000 91 NRITE<6,35) HWOOD,HBARK,HDWB1,HOWB«,HTWIG,HPOLI»HTWFO,HSTRU,HSCRO, 107,000 1HSTRE»AW00D,ABARK,A0WB1, ADWB4, ATWIG, AFOU , ATWFO, ASTRU, ASCRO, A3TRE# 108,00 0 2T*000,TBARK,TDWB1,TDW8«,TTWIG,TFOLI,TTWFO,TSTRU,TSCRO,T3TRE 109,000 <2 35 F ORMATC///j'/,'BIO^ASSjFHEMJ.OC K_ TR EE SJ _T 35,7 tF 8,1 )__3__F 1 0,1)_// 11 0,000 1 "BIOMASS CF ABlE3~fREES~t,"T3~5,7{F^,Or3(F10,l)//V/ISUM OF BIOMASS F 111 ,060 20R STAND',T35,7(F8,l),3CF10,11) 112,000 93 WRITE(<>,36) ; 113.000 94 36 " "FORMAT (' 1 • > " " - - i i i , 0 0 0 95 STOP 115.000 96 END ' 116,000 SDATA 117,000 

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