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Discount rates, small-scale fisheries, and sustainability Teh, Louise S. L. 2011

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DISCOUNT RATES, SMALL-SCALE FISHERIES, AND SUSTAINABILITY  by Louise Siok Ling Teh  B.Comm., The University of British Columbia, 2000 M.Sc., The University of British Columbia, 2006  A THESIS SUBMITTED IN PARTIAL FULFILLMENT OF THE REQUIREMENTS FOR THE DEGREE OF  DOCTOR OF PHILOSOPHY in THE FACULTY OF GRADUATE STUDIES (Resource Management and Environmental Studies)  THE UNIVERSITY OF BRITISH COLUMBIA (Vancouver)  December 2011  © Louise Siok Ling Teh, 2011     ii Abstract  In the face of overexploited and declining fisheries worldwide, a question that is central to the future sustainability of fisheries resources is how willing are fishers to sacrifice their current fishery benefits in order to be able to enjoy higher benefits in the future? Fishers’ rate of time preference, or discount rate, indicates how willing they are to delay current consumption, and is the primary topic of investigation in this thesis. I aim to answer three research questions, focusing on small-scale reef fisheries in developing countries: 1) what is the discount rate of fishers?; 2) what socio-economic conditions predict low discounting behaviour among fishers?; and 3) are discount rates reflective of the exploitation status of fisheries? I use an experimental economics approach to elicit fishers’ discount rates in Sabah (Malaysia) and Fiji, and then use regression analysis to identify the predictors of low discount rates. Further, I integrate economic and ecological concepts to infer fishers’ private discount rates, as well as to explore whether discount rates are representative of fisheries exploitation status. My main findings are that, first, small-scale reef fishers have high discount rates, with a plausible average annual range of 100 to 300%. This appears to apply to fishers in both open access (Sabah), as well as traditionally managed (Fiji) reef fisheries. There is a surprisingly larger proportion of fishers with low discount rates in the open access, compared to the traditionally managed fishery. Second, site and fishery level variables predict low discount rates among fishers, but the effect is different depending on the local socio-economic context. Overall however, it is not clear what specific site level dynamics contribute to the lower observed discount rates in Sabah. Lastly, I find that official discount rates that are used for policy making appear to be too low to fully reflect the short term outlook of reef fishers. Fishers’ higher private discount rates may be more likely to capture the exploitations status, and may be more appropriate to use for evaluating policies that affect fishers’ current and future fishing activities.     iii Preface  Chapter 2 is modified from a publication in Fisheries Research, authored by myself, Lydia CL Teh, and Rashid Sumaila. I led all stages of the research: obtaining the data, analysing the results, and writing the manuscript. LCLT helped with collecting and analysing the results of a prior study on which this chapter is based. LCLT and RS contributed to interpreting the results.  Chapter 3 is modified from a Fisheries Centre working paper authored by myself, Lydia CL Teh, and Rashid Sumaila. It has been submitted to a peer reviewed journal. I led all stages of the research: conducting the interviews, and analysing the results. LCLT assisted with conducting the interviews. RS contributed to interpreting the results and reviewing the manuscript.  Chapter 4 is modified from a publication in Sustainability, authored by myself, Lydia CL Teh, and Rashid Sumaila. I led all stages of the research, conducted the interviews, and developed the logistic regression model. LCLT helped with conducting the interviews. RS contributed to interpreting the results and reviewing the manuscript.  Chapter 5 is authored by myself. William Cheung of the UBC Fisheries Centre wrote the computer code for the algorithm to estimate the intrinsic growth rate of species.  Field interviews conducted in this research were approved by the University of British Columbia Behavioural Research Ethics Board under the following certificate numbers: H07-00750 for Sabah, and H07-02289 for Fiji.  Publications related to thesis chapters:  Teh, L.S.L., Teh L.C.L., Sumaila, U.R., 2011. Quantifying the overlooked socio-economic contribution of small-scale fisheries in Sabah, Malaysia. Fisheries Research 110, 450-458. (Chapter 2)  Teh, L.S.L., Teh, L.C.L., Sumaila, U.R., 2011.Low discounting behaviour among small-scale fishers in Fiji and Sabah, Malaysia. Sustainability, 3, 897-913. (Chapter 4)    iv Table of contents Abstract .............................................................................................................................. ii Preface ............................................................................................................................... iii Table of contents ............................................................................................................... iv List of tables...................................................................................................................... vii List of figures..................................................................................................................... ix Acknowledgements..............................................................................................................x Chapter  1: Introduction .....................................................................................................1 1.1 Objectives.............................................................................................................. 1 1.2 Exploitation of fisheries ......................................................................................... 1 1.3 Time discounting ................................................................................................... 4 1.3.1 Overview ....................................................................................................... 4 1.3.2 Application of discounting to resource management....................................... 6 1.4 Small-scale fisheries .............................................................................................. 8 1.4.1 Coral reef fisheries ......................................................................................... 8 1.5 Thesis overview................................................................................................... 10 Chapter  2: Quantifying the overlooked socio-economic contribution of small-scale fisheries in Sabah, Malaysia..............................................................................................13 2.1 Introduction ......................................................................................................... 13 2.1.1 Fisheries in Sabah ........................................................................................ 14 2.2 Method ................................................................................................................ 16 2.2.1 Data ............................................................................................................. 16 2.2.2 Summary of reconstructed small-scale catches ............................................. 16 2.2.3 Economic indicators..................................................................................... 18 Landed values .......................................................................................... 18 Profitability .............................................................................................. 20 Employment and number of people supported .......................................... 21 Economic impact of small-scale fisheries on Sabah economy ................... 21 2.3 Results ................................................................................................................. 22 2.3.1 Ex-vessel prices and landed value ................................................................ 22 2.3.2 Profitability of small-scale fishing................................................................ 23 2.3.3 Employment and dependents ........................................................................ 23 2.3.4 Economic impact.......................................................................................... 23 2.4 Discussion ........................................................................................................... 24    v Chapter  3: Time preference of small-scale fishers in open-access and traditionally managed reef fisheries.......................................................................................................34 3.1 Introduction ......................................................................................................... 34 3.2 Method ................................................................................................................ 36 3.2.1 Study sites.................................................................................................... 37 3.2.2 Fisher interviews .......................................................................................... 37 3.2.3 Estimating relative poverty........................................................................... 38 3.2.4 Estimating personal discount rates................................................................ 38 3.3 Results ................................................................................................................. 40 3.3.1 Fishery characteristics .................................................................................. 40 3.3.2 Annualised discount rate of fishers ............................................................... 41 3.3.3 Patience proxy.............................................................................................. 42 3.4 Discussion ........................................................................................................... 42 Chapter  4: Low discounting behaviour among small-scale fishers in Fiji and Sabah, Malaysia.............................................................................................................................51 4.1 Introduction ......................................................................................................... 51 4.2 Method ................................................................................................................ 52 4.2.1 Explanatory variables ................................................................................... 52 4.2.2 Analysis ....................................................................................................... 54 4.3 Results ................................................................................................................. 56 4.3.1 Site comparisons .......................................................................................... 56 Between site comparisons, all fishers........................................................ 56 Within site comparisons of low and non-low discounting fishers .............. 56 4.3.2 Logistic regression models ........................................................................... 56 Pooled model ........................................................................................... 56 Sabah and Fiji models .............................................................................. 57 4.4 Discussion ........................................................................................................... 58 Chapter  5: Investigating the economics of fisheries overexploitation ............................65 5.1 Introduction ......................................................................................................... 65 5.1.1 Discount rates .............................................................................................. 66 5.1.2 Reef fisheries ............................................................................................... 66 5.1.3 Intrinsic growth rate ..................................................................................... 67 5.2 Method ................................................................................................................ 69 5.2.1 Estimating intrinsic growth rates (r) ............................................................. 69    vi Estimating the steepness parameter (h) ..................................................... 70 5.2.2 Catch composition of reef fish catches.......................................................... 71 5.2.3 Assessing whether discount rates are indicative of exploitation status........... 73 5.2.4 Inferring fishers’ private discount rates......................................................... 74 5.2.5 Sensitivity analysis....................................................................................... 75 5.3 Results ................................................................................................................. 75 5.3.1 Intrinsic growth rates (r) ............................................................................... 75 5.3.2 Discount rates and reef fishery status............................................................ 77 5.3.3 Private discount rates ................................................................................... 77 5.3.4 Sensitivity Analysis...................................................................................... 78 5.4 Discussion ........................................................................................................... 78 Chapter  6: Conclusion......................................................................................................93 6.1 What are the private discount rates of fishers?...................................................... 93 6.2 Under what conditions do fishers exhibit low discounting? .................................. 94 6.3 Are discount rates reflective of the exploitation status of fisheries? ...................... 94 6.4 Significance and limitations of thesis research ..................................................... 95 6.5 Future directions .................................................................................................. 96 Bibliography ......................................................................................................................98 Appendices.......................................................................................................................130 Appendix A Reconstructed and reported traditional catch (tonnes year-1) and reconstructed and reported number of traditional fishers for 1950-2009.1 ............................................ 130 Appendix B Interview questionnaire for Sabah (English version) .................................. 132 Appendix C Interview questionnaire for Fiji .................................................................. 144 Appendix D Immediate and future payment amounts with associated discount rates for each choice series. ......................................................................................................... 151 Appendix E Estimated intrinsic growth rates (r) for reef fish species ............................. 154 Appendix F Top 5 families making up the catch composition for each fishery case ....... 161     vii List of tables  Table 2.1. Summary of traditional fish catch composition and breakdown of weighted retail price by retail fish grade categories................................................................................ 29 Table 2.2. Average profitability of small-scale fishing across all gears for 2009. ................. 29 Table 2.3. Estimated number of dependents of small-scale fishers as a proportion of Sabah's population 1950-2009.................................................................................................... 30 Table 3.1. Payment amounts for choice series in Fiji and Sabah. Payments for series AF and BF are in FJD, and series AS, BS, and CS are in RM1. ..................................................... 47 Table 3.2. Fishery and socio-economic characteristics of survey sites. Standard errors are provided with mean values. ........................................................................................... 48 Table 3.3. Summary statistics of fishers' annual discount rates (%) and breakdown of choices for payment options in Fiji and Sabah............................................................................ 49 Table 3.4. Mean annual discount rate (%) and breakdown of Sabah respondents according to those who chose to receive the smallest future payment offered (‘patient’), and those who chose the immediate payment for all choices (‘impatient’)............................................. 49 Table 4.1. List of interview villages in Sabah and Fiji.......................................................... 61 Table 4.2. Expected relationship between explanatory variables and low discount rates....... 62 Table 4.3. Between and within site comparison of socio-economic variables for low (DRLOW=1) and non-low (DRLOW=0) discount rate respondents. .............................. 63 Table 4.4. Regression coefficients and standard errors for logistic regression models of socio- economic variables predicting the probability of fishers choosing a low discount rate. Z values are provided in brackets, and italics indicate the odds ratio for each variable. Significance levels are indicated by asterisks. ................................................................ 64 Table 5.1. List of families grouped under periodic, equilibrium, or opportunistic life history strategies, and corresponding range of steepness parameters (h values).......................... 85 Table 5.2.Summary of catch composition case studies by country or territory, with corresponding exploitation and sustainability status as assessed by Newton et al (2007). Under=underexploited, Over=overexploited, Full=fully exploited, S=sustainable, U=unsustainable. ........................................................................................................... 86 Table 5.3. Top and bottom 3 records with highest and lowest fishery level r values, respectively. .................................................................................................................. 87 Table 5.4. Mean fishery r value by region............................................................................ 88    viii Table 5.5 Comparison of official discount rate (with fishery r (r) and corresponding exploitation status. Inferred private discount rates ( are provided in the last column. . 89 Table 5.6. Summary of families which may potentially be overfished (r< for each country or territory. .................................................................................................................... 90 Table 5.7. Private discount rates (annual rate per year) by region and exploitation status. .... 91 Table 5.8. Results of sensitivity analysis of different h assumptions on fishery level intrinsic growth rates for the 3 cases with highest and lowest fishery level r. ............................... 91     ix List of figures  Figure 2.1. Map of Sabah state, showing the interview sites of Pulau Banggi and the Semporna Islands. The inset map shows Malaysia shaded in grey.................................. 31 Figure 2.2. Annual average nominal and real ex-vessel fish prices....................................... 32 Figure 2.3. Annual nominal and real catch value for Sabah’s reconstructed traditional fish catch.............................................................................................................................. 32 Figure 2.4. Annual estimated nominal value of reconstructed traditional fish catch versus value of reported landings. The gap between the two curves show the catch value unaccounted for in reported statistics. ............................................................................ 33 Figure 2.5. Annual nominal and real catch value per fisher. ................................................. 33 Figure 3.1. Map of Fiji with circles marking the islands where interviews were conducted. . 50 Figure 5.1. Map of global coral reef distribution. Source: coral reef distribution data from UNEP-WCMC ( ................................................................. 92 Figure 5.2. Distribution of fishery r values for all cases. The horizontal lines show the mean family intrinsic growth rate for Serranidae (groupers), Lutjanidae (snappers), Acanthuridae (surgeonfish), and Scaridae (parrotfish).................................................... 92     x Acknowledgements  I thank my supervisory committee: Rashid Sumaila, Colin Clark, Mike Meitner, and Dirk Zeller, for their insights and guidance. I am especially grateful to Rashid, my thesis supervisor, for his constant advice, support, and encouragement that has led me through my Masters and PhD.  This thesis would not have been possible without the cooperation of many fishers in Sabah and Fiji who generously shared their time and knowledge with me. Thanks to Professor Ridzwan and the Borneo Marine Research Institute Seaweed Project staff for providing logistical support in Pulau Banggi. Lee Yoke Lee, Voon Yan Yi, Robecca Jumin, and the WWF Kudat team collaborated and helped with fisheries monitoring. As well, Nina Ho and the WWF Semporna team supported fieldwork in Semporna. Fieldwork in Fiji was done jointly with the Fiji Fisheries Department. I thank Nanise Kuridrani and Vily Tuiwakaya for all their help in coordinating logistics, translating, and travelling around Fiji with me.  Funding for fieldwork in Fiji was provided by the Kingfisher Foundation, while fieldwork in Sabah was partially funded by WWF Malaysia and a COSMOS International Graduate Travel Award from the UBC Fisheries Centre. I acknowledge a doctoral scholarship from the Canada Social Science and Humanities Research Council and a PhD Tuition Fee Award from UBC.  The following individuals generously provided data and/or their expert opinion about reef fish catch composition for Chapter 5: S. Anandappa, J.Cuetos–Bueno, T.E. Davies, A. Edwards, J. Gumbs and the Anguilla Department of Fisheries and Marine Resources, A. Johnson, C. Poonian, and the Turks & Caicos School of Field Studies. William Cheung kindly let me use his algorithm for estimating the intrinsic growth rates of fish. My thanks also to all those in FERU and the Fisheries Centre who have helped me along the way.  Finally, special thanks to Lydia Teh for doing everything with me, and to my parents, for supporting my extended stay in school.      1  Chapter  1: Introduction  1.1 Objectives It is generally accepted that fisheries are on a decline worldwide, and there is a need to stem further overexploitation of marine resources and ecosystems (Pauly et al., 2002; Worm et al., 2009). The challenge for fisheries managers is to identify effective strategies that will sustain the livelihoods of people dependent on coastal marine resources, while adjusting exploitation to sustainable levels. There is increasing recognition that successful fisheries management requires understanding why fishers make the fishing effort decisions they do (Wilen et al., 2002; Salas and Gaertner, 2004). A fisher’s discounting behaviour has a potentially large effect on their exploitation rate, and hence future sustainability of marine resources (Clark, 1973; Sumaila, 2004). Yet, how fishers discount the future has received limited attention in the fisheries literature to date. The aim of this thesis is to improve our understanding of the discounting behaviour of fishers, in order to fill this research gap. Specifically, I will address three research questions: i) What are the private discount rates of fishers?; ii) Under what socio-economic conditions do fishers have high or low discount rates?; and iii) Are discount rates reflective of the exploitation status of fisheries? I focus on the small-scale fisheries of developing countries, which are the most significant to society in terms of their social, economic, and food security contributions (Berkes et al., 2001; Béné, 2006).  1.2 Exploitation of fisheries The global problem of overfishing threatens the capacity of fisheries to sustain societal needs into the future. Simply put, overfishing results from removing fish at a rate that is greater than that at which the fish population can replenish itself. If this persists for long enough, the fish population is eventually reduced. In my thesis, I examine overexploitation from the perspective of fishers’ private discount rates. However, this is but one aspect of many other possible drivers motivating fishers to overexploit marine resources. Other reasons, including the absence of property rights, the presence of perverse economic incentives, poor socio- economic conditions, weak or non-existent management, and lack of scientific knowledge about targeted fish populations all contribute to overfishing. Further, the occurrence of    2 natural catastrophic events decreases the capacity for fish populations to recover from fishing, thus accentuating the impacts from overfishing.  A fish stock can be viewed as a form of natural capital that is capable of producing a flow of sustainable yield (fish catch) and resource rent to the resource owner. In a time dynamic setting, fishery resource owners are faced with the task of deciding on what optimal fishing effort to use at each time period. According to economic theory, the owner of a resource will use a level of effort that maximises the total discounted present value of resource rent from the fishery (Clark and Munro, 1975): dttRe t tE )(maximize 0)}({  ∞ −δ                 [1.1] where  is the discount rate, and R(t) and E(t) denote time dependent resource rent and effort, respectively. An essential component of this equation is the discount rate, which indicates the decision maker’s preference for current over future revenue. I will cover discount rates in more detail in a later section.  The economically optimal harvest strategy which maximises discounted present value involves the resource owner using an effort level that adjusts the initial biomass x(t) as quickly as possible towards a long-term target optimal stock level xopt, which is given as: δ= − − )( )()(')(' opt optopt opt xcp xGxc xG                  [1.2] where )( optxG  describes the biological growth function of the fish, and )(' optxG represents the marginal productivity of the fish stock,  is the discount rate, p is the fish price, and )( optxc and )(' optxc are unit and marginal cost of fishing, respectively. The optimal stock level may be larger or smaller than xMSY, the stock level which maximises sustainable yield. Equation [1.2] shows that xopt  is a decreasing function of the discount rate; thus, discounting favours a stock level that is less than xMSY.  In essence, the discount rate indicates, to a degree, the tradeoff the resource owner is willing to make for fishing heavily now, but obtain smaller catches in the future. This is the main premise of investigation for my thesis. However, besides the discount rate, the degree of tradeoff is also influenced by other economic and biological parameters in the model. In my thesis, I focus primarily on the discounting aspect of exploitation in Chapters 3 and 4, and incorporate biological considerations in Chapter 5; however, I do not explicitly consider the    3 model’s economic parameters (e.g., fishing prices and costs). Nevertheless, economic factors can motivate fishers’ decisions for entering or leaving a fishery, and are therefore crucial considerations if we are concerned about sustainable fishing effort.  Economic incentives can be drivers of overfishing. For instance, the provision of government subsidies in the form of fuel, gear, and other capacity enhancing subsidies to the fishing industry encourages excessive effort by reducing fishing costs. Consequently, this has encouraged fishing fleets to continue fishing even when it is not profitable to do so, thereby encouraging the overexploitation of fisheries resources (Clark et al., 2005, Sumaila et al., 2008; Sumaila et al., 2010). Fisheries trade and the globalization of markets has also contributed to overfishing, leading to concerns about food security and the depletion of certain valuable marine species (Sadovy et al., 2003a; ICTSD, 2006; Smith et al., 2010). Management inertia can also lead to, or prevent recovery from overfishing, as political incentives to maintain the status quo fail to curtail excessive fishing effort (Rosenberg, 2003).  Overfishing can also be driven by pressures arising from the larger socio-economic environment. Malthusian overfishing refers to the situation in many developing countries, where the combined effects of poverty, population growth, and unemployment compel poor fishers to overexploit coastal fisheries in an effort to maintain their incomes and source of food (Pauly et al., 1989). A lack of economic prospects in other sectors of society draws people to fishing, which is normally seen as an occupation of last resort due to its relatively low cost and absence of barriers to entry.  As the number of participants in the fishery increase, fishers have to fish harder and turn to destructive gears in order to maintain their catches in the face of dwindling fish stocks (Pauly et al., 1989; Mangi et al., 2007; McClanahan et al., 2008). Since many developing country fisheries lack the management capacity to regulate fishing effort, a vicious cycle ensues, and can ultimately lead to economic and biological overfishing, as catches and income per fisher decrease. An underlying reason why this cycle can continue is because of the free and unregulated nature of many developing country coastal fisheries, which is described by the ‘Tragedy of the Commons’ (Hardin, 1968).  The commons tragedy refers to the fact that unregulated, open access resources such as marine fisheries inevitably suffer degradation because a lack of property rights over these resources means that individual users, acting upon their own self-interest, will ultimately    4 overexploit limited, open fisheries, even though it is in no one’s long term interest for this to occur. Gordon’s (1954) classic economic model describes the motivation fishers have to overexploit in such an open access fishery. Assuming that fishers are profit maximisers who act rationally and individually, Gordon’s model predicts that each fisher will only consider his own benefits when deciding how much fishing effort to exert. Fishers will ignore the fact that increasing one’s catch lowers the returns to other fishers, as well as hurts the future productivity of all fishers by affecting the fish stock. In an undeveloped fishery, resource rent can be gained; however, since the fishery is open to all, Gordon’s static model shows that fishers will enter the fishery until all economic rent is dissipated and a bionomic equilibrium is reached. This situation is known as economic overfishing, and may also coincide with biological overfishing.  Fish population declines can also occur as a result of natural population fluctuations, or as a response to environmental events that change the productivity of a population (Hutchings and Reynolds, 2004). Catastrophic events such as coral bleaching or hurricanes can lead to ecosystem phase shifts and loss of species productivity (Hughes, 1994; Bell et al., 2006). Although these phenomena do not by themselves cause overfishing, they can amplify the effects of fishing. At the same time, uncertainty about fish population dynamics can lead scientists to make misinformed harvesting recommendations that can result in overfishing (Charles, 1998; Clark, 2006). The advancement and development of new fishing technology can also be a major contributor to overexploitation (Kennelly and Broadhurst, 2002). For example, the development of engines, Global Positioning System (GPS) devices, and fish sounders enabled fishing fleets to impose greater fishing mortality, as they were able to find and catch previously protected fish stocks that lived in unreachable habitats, e.g., in the deep sea (Norse et al., 2011).  1.3 Time discounting 1.3.1 Overview Many individual and public decisions involve consequences that occur some time in the future. An individual’s time preference refers to how he/she values goods and services at different points in time. Discounting addresses the issue of time preference through the use of a discount rate, which reflects one’s willingness to trade current for future consumption. Strictly speaking, the concept of time preference and discounting differ in that time preference refers solely to one’s preference for immediate over delayed utility, whereas time    5 discounting is used to encompass any reason for caring less about a future consequence (Frederick et al., 2003). However, for the purpose of my research, I will use both terms synonymously.  Time discounting is a subject that spans both economics and psychology, with each discipline having different perspectives and approaches to studying it. Economic research on time preference tends to focus on the theoretical aspects and application of the discounted utility model (Samuelson, 1937). In contrast, psychological research on inter-temporal choice focuses on finding out why individuals are more likely to value present outcomes more than future ones. Studies have ranged from measuring differences in individuals’ ability to delay gratification to determinants of impulsivity, and cognitive and emotional mechanisms underlying inter-temporal choice (Loewenstein et al., 2003).  The original concept of inter-temporal choice, in fact, had psychological underpinnings. It was first put forward by John Rae in the 1800s, who attributed the difference in levels of savings and investment in a society to a psychological factor he termed “the effective desire of accumulation” – meaning the willingness to delay gratification. A shift towards an economic perspective of inter-temporal choice occurred with the introduction of the discounted utility model by Samuelson in the 1930s.  The discounted utility (DU) model of inter-temporal choice states that an individual’s utility function (U) for a certain outcome or good C (e.g., money, oranges, etc.), is equal to the sum of the discounted utilities of each outcome in period t, such that a person’s inter-temporal utility function (Ut) is described by:  − = += tT k ktTt t cukDccU 0 )()(),...,(                 [1.3] Where       + = ρ1 1)(kD k  In the above formula, u(ct + k) refers to an individual’s utility, or well being, in period t + k. D(k) can be interpreted as the relative weight the individual attaches in period t, to his/her well-being in period t + k, while  refers to the individual’s discount rate. As t increases, the value of the discounted utility becomes less. The simplicity of the DU model made it widely accepted as the model of choice for analysing inter-temporal decisions. However, it is not without controversy; the main contention is that the DU model does not incorporate    6 psychological considerations behind inter-temporal choice, as it condenses all such factors into a single discount rate (Frederick et al., 2003). In addition, there has been little empirical support for the DU model (Soman et al., 2005). Nevertheless, I will adopt the DU model for estimating fishers’ discount rates in the ensuing chapters, as psychological aspects of discounting are not the main thrust of this research, and the discounted utility model continues to be the dominant model used in economic analysis.  Discounting is central for comparing the net economic benefits of different alternatives that incur costs and benefits through time. The standard procedure for cost benefit analysis (CBA) is to compare the net present values of the alternatives by discounting the streams of benefits and costs of each alternative using a discount rate. The option with the largest time weighted benefit is chosen. Discounting the future heavily (i.e., having high discount rates) puts less weight on future consequences when making decisions in the present, such that options with long-term costs but near-term benefits appear more attractive than those that have near-term costs but long-term benefits.  The serious implications of high discount rates for natural resource policies have been extensively highlighted (Ramsey, 1928; Costanza, 1991; Barbier et al., 1994). Discounting environmental costs and benefits also brings up significant ethical issues such as intergenerational equity, as discounting inherently places less value on the preferences of future generations relative to our present one (Padilla, 2002). This has generated considerable debate over the choice of discount rate, discounting technique, and whether discounting should be used at all for evaluating environmental projects or polices (Fearnside, 1989; Kolb and Scheraga, 1990; Markandya and Pearce, 1991; Barbier et al., 1994; Sumaila, 2004; Sumaila and Walters, 2005; Sáeza and Requenab, 2007). To incorporate intergenerational considerations, alternative techniques of discounting have been developed, such as intergenerational discounting (Sumaila, 2004; Ainsworth and Sumaila, 2005; Sumaila and Walters, 2005), gamma discounting (Weitzman, 2001), and a two-step discounting procedure proposed by Rabl (1996).  1.3.2 Application of discounting to resource management While much attention has been focused on the broad environmental and sustainability implications of discounting at the societal level, less attention has been paid to the effects of private discounting on natural resource use and sustainability. Private discounting refers to    7 the intuitive discounting that fishers and other resource users apply in their decision-making processes. It is this aspect of discounting that my research seeks to address, as many decisions about exploiting natural resources are made privately (Markandya and Pearce, 1991). This is especially true for small-scale artisanal and subsistence fisheries, where fishing is still predominantly carried out by individual fishers.  The important role that fishers’ discount rates play in determining the sustainability of a fisheries resource has been well established theoretically (Clark, 1973; 1985; Sumaila and Walters, 2005). Not incorporating fishers’ discount rates when analysing policies that affect fishers (e.g., no-take areas, rebuilding, effort allocation) will likely result in dissatisfaction and non-compliance with the management strategy. For example, Clark (1985) pointed out that disagreement between regulators and the fishing industry over acceptable catch levels were essentially an argument over discount rates. Despite this, empirical studies investigating private discount rates within the context of natural resource use have mainly been done in the fields of agriculture and forestry. For instance, Ethiopian farmers with higher discount rates were less likely to invest in soil conservation practices (Yesuf and Bluffstone, 2008), and were also less likely to recognise soil erosion being a problem (Shiferaw and Holden, 1998). In contrast, upland farmers in the Philippines still adopted soil conservation practices despite having high discount rates (Lumley, 1997). Gunatilake et al. (2007) found that in Sri Lanka, individuals with higher rates of time preference caused forest resource depletion, while Casse et al. (2005) found that observed discount rates of inhabitants in highly deforested areas were significantly higher than those living in other areas.  Only a handful of studies have documented the private discount rates of fishers (Curtis, 2002; Akpalu, 2008; Fehr and Leibbrandt, 2008; Nguyen, 2009). Of these studies, Akpalu (2008) found that fishers with higher discount rates had a higher intensity of violating fishing gear regulations. Similarly, Fehr and Liebbrant (2008) found that Brazilian fishers who were more patient exerted less pressure on fisheries resources. These two studies provide some empirical evidence that high private discount rates appear to be unfavourable for sustainable fisheries resource use.  For fisheries management, it is useful to not only know what the discount rate of fishers is, but also under what conditions high or low discount rates may occur. There is, to my knowledge, no study which examines the correlates of low or high discount rates among    8 fishers. In the broader social science literature, researchers have found that socio-economic and demographic variables such as income, wealth, age, and education level are correlated with discount rates (Pender, 1996; Becker and Mulligan, 1997; Godoy and Jacobson, 1999; Kirby et al., 2002). Poverty is commonly thought to be a driver of high discount rates (Mink, 1993), and this is a particular concern in small-scale fisheries, which predominate in tropical, less developed countries (Béné et al., 2007; 2010).  1.4 Small-scale fisheries Although they differ according to location, small-scale fisheries can generally be characterised as being multi-gear and multi-species, with almost all of the catch being used as food fish. Half of the world’s supply of food fish originates from small-scale fisheries, and it has been estimated that 90% of fishers worldwide are small-scale fishers (Béné, 2006). As 95% of these fishers reside in developing countries (FAO 2004), small-scale fisheries are fundamental for the livelihood, welfare, and food security of coastal communities in some of the world’s poorest countries. An important welfare dimension of small-scale fisheries is their ability to serve as social safety nets, and to absorb, or act as a buffer for excess labour (Béné 2005).  The scattered and unorganised nature of small-scale fisheries makes them difficult to monitor, and this is compounded by the fact that small-scale fishing communities normally consist of the rural poor, who face social and political marginalisation (Pauly, 1997). Thus, despite their importance, small-scale fisheries generally lack management (Berkes et al., 2001; Salas et al., 2007; Charles, 2011), and their socio-economic contributions have largely been disregarded by policy makers (Zeller et al., 2006a). Consequently, most small-scale fisheries exist in a data-poor condition, making it difficult to capture the economic and social contributions of these fisheries, and leading to their undervaluation and under-appreciation in many countries (Gillett and Lightfoot, 2002; Zeller et al., 2006a; b; Andrew et al., 2007).  1.4.1 Coral reef fisheries I focus on small-scale fisheries which occur in the vicinity of coral reefs, and target the large variety of fish and invertebrate species that live in coral reefs and reef associated habitats. These fisheries are dominated by small-scale subsistence and artisanal fishers, who use gears ranging from hook and line, gillnet, traps, and spears, to destructive techniques such as dynamite and cyanide (Dalzell, 1996; Dalzell et al., 1996). It has been estimated that coral    9 reef fisheries make up approximately 2 to 5% of global marine fisheries catches (Pauly et al., 2003). While the absolute yield from reef fisheries may not count as significant, their relative importance to the subsistence and livelihood of coastal communities is enormous (McManus, 1997; Whittingham et al., 2003; Sadovy, 2005; Loper et al., 2008). This is especially pertinent in parts of Southeast Asia and Oceania, where coastal communities are heavily dependent on fisheries (McManus, 1997; Pauly, 1997; Kronen et al., 2010a).  A recent assessment of coral reef fisheries sustainability by Newton et al. (2007) found that over 50% of the reef fisheries they studied were exploited unsustainably. Although customary marine tenure of coral reef resources is practiced in parts of Oceania, Indonesia and East Africa (Mantjoro, 1996; McClanahan et al., 1997; Johannes, 2002; Cinner et al., 2005), well- managed reef fisheries are rare, with most coral reef fisheries  being essentially unmanaged (Sale, 2008). The management of coral reef fisheries is even more challenging due to factors such as their open access nature, the presence of multiple landing sites and large numbers of participants (Sadovy, 2005). As with small-scale fisheries in general, tropical reef fisheries are often marginalised from mainstream policy makers (Pauly, 1997), leading to less than adequate management attention and resources spent on monitoring these valuable fisheries (Sale, 2008). These factors compound the current lack of data about the dynamics of fishing communities and the behaviour of reef fishers (Clua et al., 2005).  Fishing on coral reefs reduces the abundance of species targeted by the fishery, alters the size and age distribution of target fish, and possibly alters the trophic structure of reef communities and habitats (Jennings and Polunin, 1995; Jennings and Lock, 1996; Dulvy et al., 2004a; Graham et al., 2005). Overfishing also reduces reef species diversity, and can cause local extinctions of both target and non-targeted reef fish species (Roberts, 1995; Dulvy and Polunin, 2004). In recent decades, the relentless expansion of the live reef fish trade has led to serial local depletions of economically valuable, but biologically vulnerable species such as humphead wrasse (Cheilinus undulatus), and coral grouper (Plectropomus spp.) throughout the Indo-Pacific (Sadovy et al., 2003a; Warren-Rhodes et al., 2003; Scales et al., 2007). At the extreme, overfishing leads to the loss of key functional groups, eventually reducing and threatening the resilience of coral reef ecosystems (Roberts, 1995; Nyström et al., 2000; Wilson et al., 2006).     10 To ensure that reef fisheries can continue to contribute to the sustenance of present and future generations, current social and economic dependence on reef fisheries resources must be reduced. There is consensus that bio-physical centred management strategies for conserving coral reef ecosystems are unlikely to be adopted in developing countries (Bell et al., 2006; McClanahan et al., 2009a). Instead, successful management has to fit within the local socio- cultural and economic context (Cinner et al., 2009b; McClanahan et al., 2009a). Recent emphasis has been placed on management policies that can adapt to local socio-ecological conditions and build the capacity of communities to cope with change (Bell et al., 2006; Cinner et al., 2009b; McClanahan et al., 2009a; 2009b). At the root of these strategies is the need for an increased understanding of fishers’ outlook about their future, i.e., how willing are fishers to sacrifice their current fishery benefits in order to be able to enjoy higher benefits in the future? This is the main topic I investigate in my thesis.  1.5 Thesis overview My thesis presents 4 studies that address two research gaps in the socio-economic aspects of fisheries: 1) The chronic underestimation and undervaluation of small-scale fisheries’ socio-economic contribution; 2) The time preference of small-scale fishers.  In the next section, I outline how I will address each of these questions in my thesis chapters (Chapters 2-5). In the final chapter (Chapter 6), I summarise my main findings and discuss my study’s contribution to the current state of knowledge about fishers’ discounting behaviour. I also consider the strengths and limitations of my research, and provide some recommendations for the practical application of my results, and direction for future research.  In Chapter 2, I set the context for why my study focuses on small-scale fisheries. Using the small-scale fisheries of Sabah, Malaysia, as a case study, I illustrate the crucial role small- scale fisheries play in supporting different levels of economy, and also address the chronic data poor state, and resulting undervaluation, of these fisheries. Specifically, I quantify the present and historical (1950-2009) socio-economic contribution small-scale fisheries make in supporting Sabah’s economy and coastal livelihoods. I show that this contribution has been substantially undervalued or even unaccounted for, historically and in present fisheries statistics.    11  In Chapter 3, I first use an experimental economics approach to quantify the private discount rates of fishers in two developing country coral reef fisheries – Sabah (Malaysia), and Fiji. I then compare the discount rates from both study sites, which differ in that Sabah has essentially open access fisheries, whereas Fiji’s marine resources are governed under a customary marine tenure system. In doing so, I test whether a fundamental fisheries economics theory, that fishers in an open access fishery have ‘infinite’ discount rates (Clark, 1990), applies to fishers in Sabah. I find that fishers under both institutional systems have high discount rates; however, those in Sabah are not excessively high as to be considered ‘infinite’. Interestingly, there is no lower tendency to have high discount rates in traditionally managed versus open access fisheries.  In Chapter 4, I attempt to identify the socio-economic predictors of low discounting behaviour among small-scale fishers in Fiji and Sabah. I use the private discount rates obtained from Chapter 3 to define a low discount rate, and then use a logistic regression model to identify predictors of low discount rates. I find that a substantial proportion (42%) of fishers have low discount rates. In a pooled model that includes fishers from both Fiji and Sabah, site and village level variables are significant predictors of low discount rates. In two site-specific models that include observations from Sabah or Fiji only, boat ownership and relative catch differentiate low discounting from non-low discounting fishers, but these variables have contradictory effects in Sabah and Fiji. This suggests that local socio-cultural, economic, and ecological conditions have to be considered in the process of designing management interventions.  In Chapter 5, I move from site specific studies to a global study of discounting in coral reef fisheries. I use the theory of ‘economics of overexploitation’ developed by Clark (1973) to revisit my first research question about what the private discount rates of fishers are, and also address whether discount rates are reflective of the exploitation status of reef fisheries. My results show that the private discount rates of fishers are high, which contrasts with the substantially lower official discount rates used by fishery managers for evaluating projects. The discrepancy between these two sets of discount rates suggests that the future outlooks of those who manage and those who use fisheries resources do not coincide. This divergence may potentially affect the successful implementation of reef fisheries management strategies    12 in the present and future, and suggests that in order to improve the chances of success, policies which affect local fishers may need to be evaluated using private discount rates.      13 Chapter  2: Quantifying the overlooked socio-economic contribution of small-scale fisheries in Sabah, Malaysia  2.1 Introduction Small-scale fisheries occur throughout the world, but predominate in tropical, less developed countries (Berkes et al., 2001). Although they differ according to location, small-scale fisheries can generally be characterised as being multi-gear and multi-species, with almost all of the catch being used as food fish. These fisheries make substantial social and economic contributions to society at the national, local, and individual levels (Béné et al., 2007). Half of the world’s supply of food fish originates from small-scale fisheries, and it has been estimated that 90% of fishers worldwide are small-scale fishers (Béné, 2006). As 95% of these fishers reside in developing countries (FAO, 2004), small-scale fisheries are fundamental for the livelihood, welfare, and food security of coastal communities in some of the world’s poorest countries. An important welfare dimension of small-scale fisheries is their ability to absorb, or act as a buffer for excess labour.  They also serve as social safety nets in the sense that people may temporarily turn to fishing in times of economic stress or disaster (Béné, 2006).  Knowing the macroeconomic contribution of the fishery sector is essential for understanding the potential of fish stocks to generate wealth and contribute to social welfare (Cunningham et al., 2009). In order to manage small-scale fisheries so that they can continue to contribute to coastal livelihoods, it is thus necessary to understand the magnitude and extent of their current socio-economic contribution. Yet, in spite of their importance, small-scale fisheries generally lack management (Berkes et al., 2001), and their socio-economic contributions have largely been disregarded by policy makers (Zeller et al., 2006b). The scattered and unorganised nature of small-scale fisheries makes them difficult to monitor, and this is compounded by the fact that small-scale fishing communities normally consist of the rural poor who face social and political marginalisation (Pauly, 1997). In addition, policy makers tend to overlook small-scale fisheries in favour of commercial fisheries, such as tuna, which are important foreign exchange earners  (Gillett and Lightfoot, 2002). Consequently, most small-scale fisheries exist in a data-poor condition, making it difficult to capture the economic and social contributions of these fisheries, and leading to their undervaluation and under-appreciation in many countries (Zeller et al., 2006b; Andrew et al., 2007). Even though    14 this situation prevails in developing countries, insufficient or lack of monitoring also occurs in the small-scale fishing sectors of some developed countries (Guyader et al., 2007).  Likewise, small-scale fishers in Sabah, Malaysia are removed from mainstream society due to the mainly rural and poor socio-economic backdrop of coastal fishing communities (Abdul Mannan, 1982; Wood, 2001; Teh et al., 2005). Despite their low socio-economic status, small-scale fishers have historically played a vital role in contributing to food security, trade, and economic activity in Sabah (Sather, 1997; Mohd. Ariff, 1999). Yet, this economic contribution has not been explicitly quantified to date. To address this, my objective is to quantify the present and historical socio-economic contribution small-scale fisheries make in supporting Sabah’s economy and coastal livelihoods. I base my estimation on a previous study which quantified the historical catches of commercial and small-scale fisheries in Sabah (Teh et al., 2009a). In this study, I show the contribution small-scale fisheries make to different levels of the economy by quantifying: i) historical and current landed value of Sabah’s small-scale fisheries catch from 1950-2009; ii) current profitability of fishing to individual fishers; iii) employment and number of dependents supported by the small-scale fishing sector from 1950-2009; and iv) the economic output supported by current levels of small-scale fisheries catch value.  2.1.1 Fisheries in Sabah Sabah is a Malaysian state situated on the northeast corner of Borneo (Fig. 2.1). It is bordered by the South China Sea in the west, the Sulu Sea to the north-east, and the Sulawesi Sea to the south-east (Fig.2.1). Sabah’s history can be divided into two periods for the time scale (1950-2009) considered in this study. Prior to 1963, Sabah was under British colonial rule, and was known as British North Borneo. It then gained independence and became a Malaysian state in 1963, changing its name to Sabah.  Sabah’s marine capture fisheries were mainly small-scale until the introduction of commercial trawlers in the 1960s. In present time, small-scale fisheries continue to provide the main source of income and meat protein for a large proportion of Sabah’s rural coastal communities (Institut Penyelidikan Marin Borneo, 2003; Foo et al., 2006). Small-scale fisheries are concentrated in the inshore area, targeting mainly reef and reef associated estuarine species, and small pelagics (Biusing, 2001; Wood, 2001; Teh et al., 2005). Fishing is done for both subsistence and artisanal purposes, using a variety of manually operated    15 gears, including hook and line, gillnets, traps, spears and spear guns. Sabah’s inshore fisheries resources are extensively exploited, with signs that reef fisheries are overfished in parts of Sabah (Teh and Sumaila, 2007). Meanwhile, destructive fishing techniques using dynamite and cyanide are still common, and have damaged or destroyed unprotected reefs throughout the state (Oakley et al., 1999; Pilcher and Cabanban, 2000).  Local small-scale fishers in Sabah belong mainly to the Bajau, Ubian, Suluk, and Brunei Malay ethnic groups (Biusing, 2001). There is also a large population of migrant fishers originating from the southern Philippines and Indonesia. The number of fishers in Sabah increased rapidly in the mid 1970s to 1980s as refugees fleeing political instability in the southern Philippines settled along coastal areas or outer islands of Sabah, and turned to fishing for a livelihood (Cooke, 2008).  Prior to Sabah’s independence in 1963, British colonial reports provided data only on the quantity of exported fisheries commodities. After independence, statistics on marine fisheries landings, the registered number of vessels, fishers, and fishing gears have been compiled by the Sabah Fisheries Department. However, these data are not complete in their coverage (Cabanban and Biusing, 1999; Teh et al., 2009a). Before 1991, marine fish landings statistics were estimated from fish market surveys, trawler logbooks, fish processing plant reports, and export data.  An improved system was introduced in 1991, which used a stratified random sampling approach to estimate landings of selected fishing gears in Sabah’s 16 coastal districts (Biusing, 2001). However, this has resulted in the underestimation of small-scale landings, and in many cases, the catches from fishers living in offshore islands or rural villages go unrecorded (Teh et al., 2007). Consequently, the reported economic value of Sabah’s small- scale fisheries is also underestimated. In addition, the reported number of fishers is considered to be an underestimate due to the exclusion of many illegal or transient migrant fishers (Biusing, 2001). To address the apparent gaps in fisheries monitoring, a prior study reconstructed Sabah’s historical marine fish catches, and found that from 1950-2006, fish caught from Sabah waters were on average 2.5 times higher than landings data recorded in fisheries statistics (Teh et al., 2009a). In this chapter, I use the reconstructed catches from Teh et al. (2009a) as the basis for quantifying the historic and present economic contribution of Sabah’s small-scale fisheries to society.    16 2.2 Method 2.2.1 Data All fisheries landings and landed value statistics unless stated otherwise were extracted from the Malaysian Department of Fisheries Annual Reports ( Small-scale fisheries are referred to as traditional fisheries in the Malaysia Department of Fisheries statistics, and I use the two terms interchangeably. Data for Sabah were available from 1965 to 2009; however, the reporting of data was not consistent throughout the years, with coarser data available in the earlier years. Due to a change in fisheries statistical recording system in 1991, I compared my results to reported statistics from 1991 onwards in order to work with a consistent data set.  2.2.2 Summary of reconstructed small-scale catches I refer the reader to Teh et al. (2009a) for a detailed description of the methodology used for reconstructing the historical commercial and traditional fish catches of Sabah. Briefly, the authors used published, including grey literature to obtain information on small-scale fishing catch rates and fisher population in order to build a bottom up estimate of the amount of fish caught in Sabah waters from 1950-2006.  For the small-scale sector, the authors explicitly accounted for the large unlicensed fisher population, many of whom were illegal immigrants, and who were not captured in the official statistics. The Sabah Fisheries Department does not know how many of these migrant fishers reside in Sabah (Biusing 2001). Therefore, Teh et al. (2009a) estimated the total number of local and migrant fishers using annual population and demographic data. To summarise, the number of local fishers (inclusive of both commercial and traditional fishers) were estimated from the population of ethnic Bajau living in coastal communities in Sabah, while migrant fishers were estimated from available data on migrant arrivals. All estimated fishers were assumed to be traditional from 1950-1965, as commercial fisheries commenced in the 1960s. From 1991-1999, Biusing (2001) reported that an average of 78.5% of local fishers were traditional, while 40.7% of non-Malaysian fishers were traditional fishers. This proportional breakdown was applied to the estimated number of local and migrant fishers for 1991-2006. The proportion of local fishers considered to be traditional was linearly decreased from 100% in 1965 to 78.5 % in 1991; the same procedure was applied to non-Malaysian fishers. I used the same procedure to update the reconstructed catch and number of small-scale fishers estimates for 2007-2009.    17  Catch rates for 3 periods that roughly coincided with prevailing economic conditions affecting small-scale fishers were estimated based upon existing literature: 1950-1969, 1970- 1999, and 2000-2006. For 1950-1969, an annual catch rate of 3.09 t fisher-1 year-1 was derived from an anthropological study of a Bajau fishing community in 1965 (Sather, 1985;1997). There were no available catch rate data for 1970-1999. However, based upon historic information on the socio-economic development of Sabah, and on fisher interview data, Teh et al. (2009a) assumed that catch rates for 1970-1989 were higher than the ensuing period of 1990-2006. A conservative annual catch rate of 5 t fisher-1 year-1 was applied from 1970-1980; starting from 1981, the catch rate was linearly decreased to 3.68 t fisher-1 year-1 in 2000. The annual catch rate of 3.68 t fisher-1 year-1 used for the period 2000-2006 was based on the average catch rate reported in a case study of small-scale reef fisheries in Sabah (Teh et al., 2007). The times series of annual small-scale catch was calculated by multiplying the estimated number of traditional fishers by the annual catch rate.  I used the reconstructed number of traditional fishers and the reconstructed traditional catch from Teh et al. (2009a) as a starting point for estimating the economic contribution of Sabah’s small-scale fisheries (Appendix A). The reconstructed small-scale catch time series from 1950 to 2006 summed to 9.7 million t, while the number of reconstructed fisher estimates for 1988 to 2006 were 2.5 times higher than the number of traditional fishers reported in the fisheries statistics for the same period. Compared to landings data reported in the Annual Fisheries Statistics from 1965 to 2006, the reconstructed catch was on average 220% larger.  The magnitude of reconstructed traditional catch and number of traditional fishers may even be larger relative to reported statistics than indicated in Teh et al. (2009a), as that study considered commercial gears to consist of trawlers and purse seines only. In this chapter, I included gillnets, which was categorised as a traditional gear by Teh et al. (2009a), as a commercial gear. This was done to be consistent with the Malaysia Department of Fisheries categorisation of commercial gears, which is comprised of trawlers, seine nets, and gill nets. Traditional gears consisted of hook and line, lift and barrier nets, stationary and portable traps, shellfish collection, and miscellaneous gears (inclusive of spears, drive-in nets, push nets, tidal traps, and hand hooks). As a result of allocating gillnets to the commercial sector, the reported traditional landings and number of fishers I used in this chapter was smaller than    18 that used in the Teh et al. (2009a) study. However, this did not affect the reconstructed catch or number of traditional fishers estimated by the authors.  2.2.3 Economic indicators I used 4 economic indicators to show the contribution of Sabah’s small-scale fisheries to different levels of economy: 1) landed value measured the gross direct benefit of the small- scale fisheries catch; 2) profits from fishing was an indictor of fisheries’ benefit to individual fishers; 3) Employment and number of fishers’ dependents were indicators of societal benefits of small-scale fisheries; and 4) economic impact quantified the value of small-scale fisheries output throughout the state economy. Landed values Landed value of reconstructed small-scale catches Landed value for each year i was calculated as Vi = Ci x Pi, Where V = landed value, C= reconstructed traditional catch and P= ex-vessel price. Ex-vessel price is the per unit fish price that fishers receive when they land their catch. Ex- vessel price may be calculated from reported landed values and landings (Sumaila et al., 2007). However, the annual landed values reported in the Malaysia Department of Fisheries Annual Reports are based on wholesale prices, and not ex-vessel prices. Therefore, I took an indirect approach to estimate ex-vessel prices for 2 separate periods as detailed below: 1950-1964: There were no reported landed value data for Sabah available for these years, so I estimated ex-vessel prices from reported retail prices for Peninsular Malaysia. Retail prices were reported by fish grade (Grade I, II, III, manure fish, prawns). I allocated these fish grades to the appropriate fish groups according to the traditional fish catch composition used by Teh et al. (2009a) to obtain an annual average retail price weighted according to traditional catch composition (Table 2.1). To convert Peninsular Malaysia retail prices to ex-vessel prices in Sabah required 2 steps: 1) a factor to account for the higher retail prices in Peninsular Malaysia versus Sabah; and 2) a factor to account for the mark-up in price from fisher to retailer.  1) I had reported annual average retail prices for both Sabah and Peninsular Malaysia for 2006-2009 from the Annual Fisheries Statistics. In addition, a report by Abdul Mannan    19 (1982) had average retail prices for Sabah for 1974-1979. For these years, I was able to calculate a ratio of Peninsular: Sabah retail prices. On average, retail prices in Peninsular Malaysia were 1.37 times higher than those in Sabah for 2006-2009 and 1.81 times higher in the 1970s. It was noted by Sather (1997) that fish prices in Sabah were very low during the 1960s. Therefore, I decided to take the higher ratio (1.81), and assumed that retail fish prices in Peninsular Malaysia were roughly 80% higher than those in Sabah for 1950-1964. 2) I used information from 2 sources to adjust Sabah retail prices calculated in step (1) to ex- vessel prices: i) According to a socio-economic survey of Sabah small-scale fishers done by Abdul Mannan (1982), fish prices were marked up by 200% from producer to retailer; ii) During the late 1960s to 1990s, it was common for fish buyers to buy fish from fishers in bundles of 15-20 individual fish (Mohd. Ariff, 1999). The fish buyer would pay the fisher according to the number of bundles, and then sell the fish at market for the same price, except with a reduced number of fish per bundle. Normally, the number of fish would be reduced by a third to a half per bundle (Mohd. Ariff, 1999). As fish in each bundle were approximately the same size, I assumed that the retail price in terms of weight was approximately 40% to 100% higher than that received by the fisher. Based on (i) and (ii), I conservatively assumed that the estimated average retail price in step (1) was about double the ex-vessel price. 1965-2009: Method 1: I extended the same 2 step procedure used to estimate ex-vessel prices for 1950- 1964 to 1965-2009. Method 2: This method was based on information independent from fisheries statistics. An anthropological study done in Semporna, south-east Sabah, in the 1960s noted that fishermen earned about 30 cents for a kati of fish (equivalent to 50 cents a kg)(Sather, 1997). Ex-vessel prices obtained from field studies in a fishing village in Pulau Banggi, Sabah, were available for 2004, 2005, 2007, and 2009 (L.Teh, unpublished data). I assumed a linear increase in fish prices for the entire period, and linearly interpolated the prices in between 1965 and 2004, and for the intervening years from 2005 to 2009. I used an average of Methods 1 (retail price based) and 2 (statistics independent) prices for all ensuing calculations.  Landed value of reported landings Data on annual landings broken down by commercial and traditional sectors was available for 1982-2009. However, annual landed value data for the same period was not available by fishing sector. To estimate the annual landed value of the traditional sector from 1982-2009, I    20 multiplied reported annual traditional landings by the calculated ex-vessel price described above.  Real prices and landed values Real annual ex-vessel prices and landed values were calculated by adjusting the estimated nominal figures by the Consumer Price Index (CPI). CPI data for 1950-2009 were obtained from the World Bank databank online ( Profitability I defined profitability as the ratio of net monthly fishing income to monthly fishing revenue. This was computed for 2009 only, and was not extrapolated to other years. I based my estimate on information gathered from semi-structured interviews conducted with 75 small- scale fishers in 2009. Interviews were conducted in 11 fishing villages concentrated in 2 geographical areas: Pulau Banggi off the northern coast of Sabah, and the Semporna islands off the south eastern coast of Sabah (Fig. 2.1). These interviews included questions pertaining to the socio-economics of fishing, including each fisher’s fishing income, costs, and fishing effort. I will elaborate on these interviews in Chapter 3.  Fishing villages in both the surveyed locations are rural, with those in Pulau Banggi being particularly remote from the mainland and nearest town centre. Fishers fished for both subsistence and artisanal purposes, and targeted reef and reef associated fish. Fishers along the west coast of Sabah tend to focus more on pelagic species. However, as the gears used by the surveyed fishers represented the main gears used by small-scale fishers throughout Sabah, and I could not find data on fishing costs and revenue for small-scale fishers on the west coast, I found it reasonable to use the interview data to represent profitability for small-scale fishers in all of Sabah. Profitability for each fisher was calculated as: P = NI/R, where NI=net fishing income per month, R=fishing revenue per month, P=profitability; and NI = R – Co - Cf , where Co = operating cost per month, Cf = fixed cost per month.  Fixed cost referred to the depreciation of a fisher’s boat, which was calculated on a straight-line basis to obtain an annual depreciation cost per boat.      21 Employment and number of people supported I made no distinction between full or part time fishers, and so considered employment in the small-scale sector to be equivalent to the reconstructed number of traditional fishers estimated by Teh et al. (2009a) (Appendix A). I estimated the number of people directly dependent on small-scale fisheries income by multiplying the annual estimated number of traditional fishers by the number of dependents per fisher, i.e., for each year i: Di = Fi * di where Di is the number of individuals directly dependent on small-scale fishers in Sabah, Fi is the number of reconstructed traditional fishers, and di is the number of dependents per fisher. I obtained estimates for the number of dependents per fisher for 2 periods: 1) 1950-1981: Abdul Mannan (1982) reported that each traditional fisher had on average 7 dependents in 1981. I applied this number from 1950 to 1981, based on the assumption that the size of fishing households was the same, or larger during this period. 2) 1982-2009: Fieldwork I conducted in 2009 indicated that the average fishing household size in rural and remote fishing villages was still 7, whereas the average household size for rural areas in Sabah was 5.2 in 2009 (Department of Statistics Malaysia, 2009a). I used the average of these two information sources to obtain an average rural fishing household size of 6.1 for 2009. A previous study indicated that there were on average 1.36 fishers per household (Almada-Villela, 1997). I subtracted this from the average fishing household size to obtain 3.49 dependents per fisher in 2009, and then linearly decreased the number of dependents from 7 in 1981 to 3.49 in 2009. Economic impact of small-scale fisheries on Sabah economy I estimated the total output in Sabah’s economy that was dependent to some extent on the output from small-scale fisheries. Counting landed value as the only contribution a fishery offers to economic output is an underestimate of a fishery’s full economic impact because it ignores the variety of secondary activities supported by fisheries (e.g., boat building, fish processing, etc.). To address this, I used the fishing output multiplier developed by Dyck and Sumaila (2010) to estimate how much output is generated by Sabah’s small-scale fishery catches (i.e., landed catch value). The main concept of multipliers is that the different sectors which make up an economy are interdependent. As such, fisheries output multipliers account for the direct, indirect, and induced economic effects of marine fisheries on the global economy. Dyck and Sumaila (2010) used an input-output method to estimate fisheries output multipliers for each maritime country.    22  Input-output analysis is a method used to track the flow of dollars between industries in the production of output for a certain sector. Inter-industry transaction data are used to compute a coefficient matrix which summarises how much output from a certain industry x is required to produce a unit of output for industry y. Dyck and Sumaila (2010) applied the Leontief technological coefficient to estimate total output supported throughout the economy at 2003 fisheries production levels, where production was measured as landed value. The landed value weighted multiplier for Malaysia was estimated to be 2.58. This meant that an extra unit of fisheries demand would generate 2.58 units of output in Sabah’s economy.  Following Dyck and Sumaila’s (2010) methodology, I used landed catch value as a measure for fisheries demand. As multipliers were available only by country level, I used the Malaysian multiplier for Sabah. The total economic impact of small-scale fisheries to Sabah’s economy in 2009 was estimated as: E = V x , where E = economic output, V= landed value of catch in 2009, and  = weighted multiplier.  2.3 Results 2.3.1 Ex-vessel prices and landed value Estimated annual average nominal and real ex-vessel prices were variable, but in general showed an increasing trend from 1950-2009. Ex-vessel prices increased substantially in the 1970s and the early to mid 1990s, reaching the highest of RM 3.9 kg-1 in the most recent year (2009) (Fig. 2.2). Similarly, the nominal and real landed value of small-scale catches showed an increasing trend from 1950-2009, reaching a maximum value of RM 664 million in 2009 (Fig. 2.3). The cumulative nominal landed value of small-scale catches for the entire analysis period totalled RM 9.3 billion, while it was RM 7.2 billion from 1991-2009. In comparison, the estimated value of reported traditional landings for 1991-2009 amounted to RM 2.6 billion, suggesting a cumulative undervaluation of RM 4.5 billion (USD 1.4 billion), or an average of RM 252.9 million (USD 80.5 million) per year. The spread between estimated catch value and reported landed value was greatest in the most recent years (Fig. 2.4). The reconstructed time series of nominal catch value per fisher showed an increasing trend, whereas the real catch value per fisher actually declined following a peak in the late 1970s, and only started to move upwards again around 2007 (Fig. 2.5).     23 2.3.2 Profitability of small-scale fishing I report the profitability of small-scale fishing across all gears because many fishers used more than one gear type. Fishing was the only source of income for the majority (75%) of respondents. The average net monthly fishing income (± standard error) was RM 442±60, with profitability of 0.45 (Table 2.2). Fuel made up the largest cost component, accounting for 73% of all operating costs on average. Although small-scale fishing appeared profitable, monthly net fishing income was still below the 2004 Sabah poverty line income of RM 704 month-1 (UNDP, 2007). Mean household expenditures were approximately RM 276 month-1 (L. Teh, unpublished data); thus, in most cases fishers were making enough just to meet household living expenses. Seventy-five percent of fishers did not have savings.  2.3.3 Employment and dependents On average, the estimated number of fishers employed by Sabah’s small-scale sector was 4.7 times higher than the number of traditional fishers reported in the fisheries statistics for 1991- 2009. In the most recent year (2009), the estimated number of small-scale fishers was 48,833, making up 1.5% of Sabah’s population. In contrast, there were 8,544 traditional fishers recorded in the fisheries statistics for 2009 (Appendix A).  The time series for the number of dependents supported by the reconstructed number of traditional fishers is presented in Table 2.3. On average, small-scale fishers may have supported up to 19% of Sabah’s population during the 1950s, although the proportion decreased so that in 2009, an estimated 48,833 traditional fishers potentially supported up to 170,197 dependents, equivalent to 5% of the state population. In contrast, applying the same number of dependents per fisher to the 8,544 reported number of traditional fishers in 2009 statistics yielded 59,739 dependents, or 2% of the population.  2.3.4 Economic impact Applying a multiplier of 2.58 to the total current (2009) estimated catch value of RM 664 million generated potential economic output amounting to RM 1.71 billion. Of this, up to RM 1.36 billion (79%) may have been due to the unaccounted value of reconstructed traditional catches.       24 2.4 Discussion My findings indicate that Sabah’s small-scale catches may have been undervalued by up to 225% from the early 1990s to the present. This reflects the general disregard of small-scale fisheries’ contribution to economies (Béné, 2006; Zeller et al., 2006b; Andrew et al., 2007). One of the main reasons government agencies tend to overlook the small-scale fishing sector is because of their relatively negligible economic contribution at the macro-level, which typically range from 0.5 to 2.5% (Béné et al., 2007). Yet, this mentality ignores the crucial welfare and livelihood role of small-scale fisheries, as illustrated by this case study.  Sabah’s fishing sector is not a major contributor to the state economy. Among agricultural sectors, the gross value of fisheries output in 2009 was valued at RM 164.4 million, compared to a gross value of RM11.3 billion for crops and livestock (Department of Statistics Malaysia, 2009b). In terms of employment, the 39,300 individuals employed in the fishing sector is a fraction of the 323,000 employed in agriculture, forestry, and hunting in 2006 (Department of Statistics Malaysia, 2007). Nevertheless, the important point of my results is the extent to which the socio-economic contribution of Sabah’s small-scale fisheries has been undervalued. I estimate that direct fisheries value was RM 664 million in 2009, compared to the value of reported traditional landings of RM 136 million. This undervaluation of more than 350% has to be rectified, as it can potentially lead to distorted outcomes if used as the basis for policy decisions.  I estimate that small-scale fishers potentially support up to 170,197 dependents, equivalent to 5% of the state population. In contrast, basing my calculation on the reported number of fishers in 2009 would have resulted in 59,739 dependents, a potential underestimate of 185%. My estimate itself is likely conservative as I only considered dependents within a fisher’s household, and did not include individuals such as fish traders who are also dependent on the output of small-scale fishers. Overall, my estimate implies that a decline in inshore marine resources will have a much more widespread impact than currently anticipated.  Using a fisheries economic multiplier to assess the total economic impact of small-scale fisheries indicates that unaccounted small-scale fisheries value in 2009 can potentially generate RM 1.36 billion in output to Sabah’s economy, which is equivalent to 4% of Sabah’s 2009 gross domestic product (GDP). Accounting for this would have increased the combined GDP contribution of commercial and traditional fisheries from the reported 2009    25 level of 2.1% to 6%, essentially increasing fisheries contribution by 200% and making it comparable in magnitude to the manufacturing industry, which contributed RM 2.4 billion to GDP in 2009 (Department of Statistics Malaysia, 2011). My results are consistent with a study which found that the contribution of small-scale fisheries to GDP was underestimated by a factor of 5 (Zeller et al., 2006b), and imply that a decline in small-scale fisheries will resonate beyond the fishing sector.  One of the fundamental roles of small-scale fisheries is in supporting the welfare of coastal communities (Béné, 2006), but this welfare dimension is under-represented by official statistics in Sabah. For example, Teh et al.’s (2009a) reconstructed number of traditional fishers was 370% higher than reported numbers from 1991 to 2006. This arose from the inclusion of illegal migrant fishers, who are not documented in official statistics. As these migrant fishers cannot easily find legal employment, fishing served as their labour buffer and social safety net; these two crucial functions of small-scale fisheries have been demonstrated in other parts of Asia and Africa as well (e.g., Jul Larsen, 2003; Béné et al., 2010).  Poverty alleviation is one of the central contributions of small-scale fisheries (Béné, 2006). In some countries, small-scale fishing boats are profitable and economically viable (e.g., Adeogun et al., 2009). My profitability analysis shows that small-scale fishing in Sabah functions to prevent further poverty, rather than to alleviate it. Even though small-scale fishing generates positive profits, average monthly fishing income is below the Sabah poverty line income, and is barely enough to cover household expenses. Thus, fishing cannot generate sufficient wealth to lift fishers out of poverty. Rather, it helps to sustain livelihoods and prevent fishing households from falling deeper into deprivation. This aspect of small-scale fisheries contribution is even more important given that Sabah is one of the poorest Malaysian states (Leete, 2008), and social welfare options are limited. Overall however, the real (i.e., inflation adjusted) annual catch value per fisher has been declining temporally, suggesting that the ability of fishing income to support economic needs has decreased from the past. These factors bring up concerns about the vulnerability of fishing households (Allison and Ellis, 2001), especially given the apparent declines in Sabah’s inshore coral reef resources (Cabanban and Biusing, 1999; Scales et al., 2007; Teh and Sumaila, 2007).  Food security is another key contribution of small-scale fisheries (Béné et al., 2007). Sabah’s fisheries were small-scale in nature until the 1960s, but these operations were able to supply    26 enough fish for the domestic market as well as for export during this period (Mohd. Ariff, 1999). Nowadays, with the exception of valuable live reef food fish species, the fish caught by traditional fishers is consumed locally. In contrast, a considerable proportion of commercial trawler landings consist of trash fish which are processed into fishmeal (Biusing, 2001), and hence do not contribute to local food security. Though not directly quantified here, my study brings to light that food security enjoyed by Sabah society at large can be attributed to small-scale fishers. Yet, small-scale fishers often face social marginalisation, and their crucial contribution is not widely recognised.  In contrast to other developing countries where the traditional small-scale sector is omitted from official statistics (e.g., Zeller et al., 2006a), the Sabah Fisheries Department does monitor traditional fish landings and values. However, the problem is that the system fails to capture the entire scale of the small-scale fishing sector, largely due to the social marginalisation of small-scale fishers. In particular, the Bajau Laut, who are the traditional maritime people of this region, have often been shunned by other segments of society for their nomadic sea faring way of life (Saat, 2003; Torres, 2005). In addition, the increasing number of illegal migrant small-scale fishers in the past few decades is difficult to account for, as they tend to settle in rural offshore islands (Piper, 1984; Fisher, 2000; Biusing, 2001). The presence of illegal immigrants is a contentious social and political issue in Sabah (Sadiq, 2009). Nevertheless, it is essential to start documenting this hidden segment of small-scale fishers, not only to obtain a better estimate of small-scale fisheries’ socio-economic contribution, but also to obtain a better understanding of the level of inshore fisheries exploitation.  Teh et al. (2009a) showed that the largest increase in small-scale catches occurred in the 1990s, which corresponded to the period of highest population growth. Sabah’s population doubled from 1990 to 2009 (Department of Statistics Malaysia, 1995; 2009a), in large part due to the arrival of illegal immigrants (Sadiq, 2009). The fisheries impact of population growth spurred by migrant fishers raised concerns in a case study of migrant fishers in neighbouring Indonesia (Kramer et al., 2002), but this issue has not been sufficiently addressed in Sabah.  Although coarse, my time series analysis offers a more encompassing quantification of the historical economic value of Sabah’s fisheries catches from1950-1963, as British records    27 only reported the value of exported processed fisheries products for this period. By doing so, my study provides an approximate baseline by which to assess the socio-economic impact of future changes to Sabah’s inshore marine resources.  My quantification complements existing social economic studies which tend to have focused on the anthropological aspects of Sabah’s traditional fishing communities (e.g., Sather, 1997; Ono, 2010), or have been based on government fisheries data (e.g, Mohd. Ariff, 1999; Biusing, 2001; Noh and Tai, 2001).  My approach for estimating historical and present socio-economic contribution tackles the difficulty of dealing with the data-poor nature of small-scale fisheries, which is a major obstacle to their management (Johannes, 1998). I relied on a combination of formal and informal sources -fisheries data, grey and published literature, and field observations – which are typically publicly available or collectible at minimal cost, to fill in data gaps. At the same time, I acknowledge that this approach entails uncertainty and a certain degree of subjectivity. However, sensitivity analysis shows that the study’s overall trends are consistent even if certain variables change. For example, if the price variable is increased by 10%, estimated reconstructed catch value increases by 5%, but the magnitude by which reported landings are underestimated remains unchanged. Further, the economic impact of Sabah’s small-scale fisheries may be overestimated because the fisheries multiplier calculated by Dyck and Sumaila (2010) was a country average for Malaysia. The rural nature and low economic integration of many Sabah fishing communities means that the multiplier for its small-scale fisheries may be lower than the national average. Using a multiplier that is 25% lower would result in an economic impact of RM 1.28 billion in 2009, and this would lower the current estimated fisheries contribution to Sabah GDP from 4% to 3%.  On the other hand, the national fisheries multiplier may be a conservative estimate because of economic leakage, which lowers a multiplier. Economic leakage occurs when the earnings obtained in one sector do not remain within the sector, but are spent in another sector instead. Kamaruddin and Abdul Rashid (2007) found moderate leakage in the Malaysian fishing industry. However, given the absence of a state specific fisheries multiplier for Sabah, the economic impact calculated based on the national average multiplier is my best estimate. Overall, given the general paucity of data and comparable studies on which to calibrate my findings, I contend that my results present the current best estimate of Sabah’s small-scale fisheries socio-economic contribution with regards to the factors measured. My intent is that these figures be treated as first attempt estimates which can be refined and improved upon.    28  In summary, my study shows that the social and economic contributions of small-scale fisheries to Sabah society have been substantially undervalued or even unaccounted for historically and in the present fisheries statistics. The significant undervaluation of fisheries output implies that fishing pressure on Sabah’s inshore marine resources is probably a lot higher than presently perceived, and illustrates the risk associated with the prevailing indifference towards understanding the ‘true’ magnitude of small-scale fishing. For Sabah, my study strongly suggests the need for more encompassing fisheries monitoring and data collection methods which include the large undocumented population of small-scale fishers. A necessary complementary step may be for Sabah’s Fisheries Department to start addressing the persistent issue of illegal immigrant fishers in tandem with state immigration and other relevant social welfare authorities. Even though my results are specific to Sabah, they are relevant for all small-scale fisheries where monitoring of the sector is minimal or does not exist. Importantly, my results dispel the common notion that small-scale fisheries are of negligible economic importance. Fully accounting for small-scale fisheries contribution shows that although they do not have a heavy impact at the macro-economic level, the socio- economic contribution of small-scale fisheries to the welfare and livelihood of coastal communities is crucial.     29  Table 2.1. Summary of traditional fish catch composition and breakdown of weighted retail price by retail fish grade categories. Traditional fish catch composition1 Fish groups  Percentage of catch Small pelagic (SP)  0.10 Medium and large pelagic (MLP)  0.45 Demersals (DM)  0.31 Crustaceans (CR)  0.06 Molluscs and misc. invertebrates (MIV)  0.02 Trash fish (TF)  0.06 Breakdown of weighted retail price Retail fish grade category Fish group composition  Retail weight Grade I MLP, DM, CR 0.38 Grade II MLP,DM 0.38 Grade III SP, MIV 0.12 Crustaceans CR 0.06 Manure fish TF 0.06 1Source: Teh et al. (2009a).   Table 2.2. Average profitability of small-scale fishing across all gears for 2009. Variable RM month-1 1   (Mean± std. error) Fishing revenue 1141±254.1 Fishing costs 697±219 Net income 442±60.3 Average profitability (%) 0.45 1The currency exchange rate on 5 December 2010 was USD 1 = RM 3.14.      30  Table 2.3. Estimated number of dependents of small-scale fishers as a proportion of Sabah's population 1950-2009. Year Estimated no. of dependents % of Sabah population 1950 67671 0.19 1951 63958 0.19 1952 64528 0.19 1953 65758 0.18 1954 66101 0.18 1955 66577 0.18 1956 67387 0.17 1957 68189 0.17 1958 69436 0.17 1959 70403 0.16 1960 72877 0.16 1961 73326 0.16 1962 73584 0.15 1963 73648 0.15 1964 74205 0.14 1965 74675 0.14 1966 70249 0.12 1967 69401 0.12 1968 68990 0.11 1969 68878 0.11 1970 68968 0.10 1971 69882 0.09 1972 68882 0.09 1973 70169 0.09 1974 74796 0.09 1975 76480 0.09 1976 79245 0.09 1977 81589 0.09 1978 84107 0.09 1979 85537 0.09 1980 88919 0.09 1981 86916 0.08 1982 90183 0.08 1983 93102 0.08 1984 96388 0.08 1985 99632 0.08 1986 102968 0.08 1987 106452 0.08 1988 109408 0.08 1989 113823 0.08 1990 118446 0.08 1991 127456 0.07 1992 133852 0.07 1993 140330 0.07 1994 151291 0.07 1995 154230 0.06 1996 158501 0.06 1997 160447 0.06 1998 161999 0.06 1999 169486 0.06 2000 155850 0.06 2001 157656 0.06 2002 155490 0.06 2003 157243 0.06 2004 159226 0.06 2005 162566 0.05 2006 161509 0.05 2007 163012 0.05 2008 163260 0.05 2009 170197 0.05      31  Figure 2.1. Map of Sabah state, showing the interview sites of Pulau Banggi and the Semporna Islands. The inset map shows Malaysia shaded in grey.      32  Figure 2.2. Annual average nominal and real ex-vessel fish prices.    Figure 2.3. Annual nominal and real catch value for Sabah’s reconstructed traditional fish catch.    33  Figure 2.4. Annual estimated nominal value of reconstructed traditional fish catch versus value of reported landings. The gap between the two curves show the catch value unaccounted for in reported statistics.  Figure 2.5. Annual nominal and real catch value per fisher.     34 Chapter  3: Time preference of small-scale fishers in open-access and traditionally managed reef fisheries  3.1  Introduction The majority of the world’s coral reefs occur along the coasts of tropical developing countries, where socio-economic dependence on reef fisheries is disproportionately large (Whittingham et al., 2003; Bell et al., 2006). Yet, inshore coral reef resources are being overexploited globally, with serious biodiversity, social, economic, and cultural consequences (Bellwood et al., 2004; Newton et al., 2007). Malthusian overfishing is widespread in many tropical inshore fisheries (Pauly et al., 1989; Teh and Sumaila, 2007; McClanahan et al., 2008), which are mostly unmanaged and operate under an essentially open access regime (Sadovy, 2005; Sale, 2008). I define an open access fishery as one in which fishing rights are non-existent or ill-defined, access to the fishery is unrestricted, and there are no regulations governing the use of the fishery resource (Greboval and Munro, 1999). In contrast to open access systems, customary marine tenure is practiced in the Pacific islands and parts of Indonesia and East Africa (Mantjoro, 1996; McClanahan et al., 1997; Johannes, 2002).  Fisheries economics theory suggests that overexploitation occurs when fishers’ discount rates are high (Clark, 1973; Sumaila and Walters, 2005), i.e., they prefer receiving an immediate, certain, smaller benefit over future, uncertain, but potentially larger benefits. Weak fishery institutions provide further incentives for high discount rates. The problem of overfishing stems from the ‘tragedy of the commons’ (Hardin, 1968), in which unrestricted access to a fishery leads to overexploitation because individual fishers have no incentive to conserve, since they have no assurance that the fishery resource will persist into the future. Therefore, in theory, fishers in an open access fishery are compelled to discount the future heavily when making fishing decisions, despite the fact that their individual private discount rates may be low. In essence, the open access environment forces fishers to use a discount rate of infinity (Gordon, 1954; Clark, 1990). Although an ‘infinite’ discount rate is theoretically possible, it is likely not realistic in practical terms. In this chapter, I use ‘infinite’ discount rate to refer to a very high discount rate.  Despite the serious implications the open access theory has for the management of small- scale reef fisheries in developing countries, there has, to my knowledge, been no empirical    35 study investigating this assumption. I fill this gap by examining the discount rates of fishers in two small-scale tropical reef fisheries operating under different institutional environments: 1) an open access regime in Sabah, Malaysia; and 2) customary marine tenure (CMT) in Fiji. I seek to answer the following questions: 1) What are the discount rates of fishers in open access and traditionally managed fisheries? 2) Do fishers in an open access fishery have ‘infinite’ discount rates?  An individual’s time preference refers to how one values goods and services at different points in time, and is expressed through a discount rate, which reflects one’s willingness to trade current for future consumption. The importance of individuals’ time perspective for policy development has been recognised in fields where decisions involving future outcomes are critical, e.g., in health and credit markets (Pender, 1996; Poulos and Whittington, 2000; Amadja, 2008). Recent studies have shown that discount rates elicited for one domain can be applicable to other domains and context (Hardisty and Weber, 2009), as well as behaviour (Chabris et al., 2008).As such, investigating fishers’  time preference is of policy interest because it can inform us about fishers’  tendency towards future behaviour regarding fisheries extraction. As outlined in Chapter 1, I view high discount rates as indicators of unsustainable marine resource use (Clark, 1973; Sumaila, 2004; Akpalu, 2008). Relationships of high discount rates corresponding to less sustainable natural resource use have also been observed in studies of forest clearing and agriculture (Shiferaw and Holden, 1998; Casse et al., 2005; Gunatilake et al., 2007).  Fishery institutions play a key role in influencing how fishers respond to local and global pressures to exploit fisheries resources (Clark et al., 2010a; Smith et al., 2010). The competition inherent in a pure or regulated open access fishery, where fishers have no security to future resources, motivates fishers to engage in myopic behaviour consistent with high discounting. This is evidenced by the ‘race to fish’  mentality that has led to the depletion, and even collapse, of various fisheries around the world (Costello et al., 2008).  In contrast, fisheries managed under customary or other rights-based systems are characterised by a well defined group of users and by the presence of rules or norms which regulate individual use. Rights-based management systems are more likely to provide fishers with an incentive to use fishery resources sustainably (Dietz et al., 2003; Grafton et al., 2006; Costello et al., 2008). Consequently, I expect these fishers to possess a longer term outlook, which is expressed through a lower discount rate. For instance, Asche (2001) shows that    36 discount rates in well developed quota systems are relatively low. At the same time, Clark et al. (2010a) demonstrate that complete privatisation of fisheries may not lead to long-term oriented behaviour.  Coral reef resources have traditionally formed the backbone of fishing economies along the coasts of Sabah and Fiji (Sather, 1997; Gillett and Lightfoot, 2002). Fishing communities in both countries tend to be poor, with high reliance on inshore reef resources for food and income. In the past decade, inshore fisheries resources in Sabah and Fiji have shown signs of overfishing, and fishers in both countries have similarly noted a decrease in catch rates and the size of fish caught (Teh and Sumaila, 2007; Turner et al., 2007; Teh et al., 2009b).  In Sabah, a spatial zoning system is used to regulate fishing activity. The traditional fishing zone is reserved exclusively for small-scale subsistence and artisanal fishing vessels, but is rarely enforced. There are no formal fisheries governance institutions at the village level, especially in rural communities such as those in Pulau Banggi and the Semporna islands, where this study takes place (Teh et al., 2005). Fishers have free access to the majority of fishing grounds around the study areas, although this may change in the future because Marine Protected Areas (MPAs) are in the process of being set up at both sites.  Fiji has an established customary marine tenure system (CMT), in which inshore fishing grounds are divided into customary fishing rights areas called qoliqolis (Cooke et al., 2000). Outsiders wishing to fish within a certain qoliqoli are required to obtain the permission of the qoliqoli owners, who are able to impose fishing restrictions on fishing method, area, season, or species (Veitayaki, 1998). Nevertheless, Fiji’ s traditional system has been undermined in recent decades as villages have become more urbanized and integrated into the market economy (Matthews et al., 1998).  3.2 Method Data for this study were obtained from 122 interviews conducted with fishers in Fiji and Sabah. The interviews were semi-structured and followed a prepared questionnaire, although interviewees had the flexibility to expand on topics of interest.  The discount rate elicitation questions formed part of a longer questionnaire that also covered socio-economics, spatial use of fishing grounds, and perceptions of change to fishery management (Appendix B and C).    37  3.2.1 Study sites Sabah Sabah is a Malaysian state situated on the northeast corner of Borneo, within the Coral Triangle (Fig. 2.1). The population of Sabah was 3.2 million in 2009 (Department of Statistics Malaysia, 2009a). My study was conducted in 11 rural fishing villages grouped into two main geographical areas: 4 villages were located in the Semporna group of islands, off the south-eastern coast of Sabah, while another 7 villages were located in Pulau Banggi, which is off the northern tip of Sabah (Fig. 2.1).  Fiji Fiji is an archipelago situated in a biodiversity rich area of the South Pacific (Fig. 3.1) (WWF, 2003). Fiji’ s population in 2007 was 837,271 (Fiji Islands Bureau of Statistics, 2007). Interviews for this study took place in 9 villages and 2 settlements located on Viti Levu and Vanua Levu, the two largest and most developed islands, as well as on the smaller islands of Yasawa and Kadavu. Interview sites were selected to provide a wide geographical coverage of Fiji, as well as being inclusive of both artisanal and subsistence fishing communities.  3.2.2 Fisher interviews Sabah I interviewed 75 fishers in Pulau Banggi and the Semporna islands in April and May 2009, with the assistance of another graduate student from the University of British Columbia Fisheries Centre. We conducted interviews in Malay, with each interview taking between 45 minutes to one hour to complete. I used opportunistic sampling at each village, walking from house to house to interview fishers after obtaining permission from the village headman. In Sabah, all the fishers I interviewed were men, as they are the fishers in the household. Women may occasionally glean on the reef, but in general do not participate in fishing.  Fiji I interviewed 47 small-scale fishers in Fiji in May and June 2008. My interview team consisted of two research officers from the Fiji Fisheries Department and a graduate student from the University of British Columbia Fisheries Centre. We conducted interviews in English or Fijian, and each interview lasted from half an hour to 45 minutes. Permission from the village chief was obtained prior to conducting our interviews. In some villages, fishers    38 were already gathered in a community hall upon our arrival, and we were able to interview as many fishers as possible from those who were present. Where fishers were not already gathered, we walked from house to house to conduct interviews. Women made up 34% of the respondents.  3.2.3 Estimating relative poverty To assess the relative poverty level in fishing communities in Sabah and Fiji, I calculated a fisher poverty index (Sumaila, 2003). This was measured as the monthly net fishing income divided by the poverty line income in each country.  The 2004 poverty line income in Sabah was RM 704 month-1 (USD 207 month-1) (UNDP, 2007), while the Fiji poverty line income of  USD 254 month-1 was based on a Basic Needs Poverty Line income of FJD125 week-1 (USD 64) in 2003 (Fiji Islands Bureau of Statistics, 2009). These figures were adjusted using the Consumer Price Index to provide a poverty line income of USD 236 month-1 in 2009 dollars for Sabah, and USD 294 month-1 in 2008 dollars for Fiji.  3.2.4 Estimating personal discount rates To estimate each respondent’ s private discount rate, I provided a series of binary choices involving monetary payments that occurred at different points in time. I used hypothetical payments in both Fiji and Sabah due to budgetary and logistical considerations. While other studies have used food or cash payments (Kirby et al., 2002; Fehr and Leibbrandt, 2008), there has been no conclusive evidence that real payments result in more realistic answers (Frederick et al., 2003).  The choice series followed a ‘multiple price list’  format similar to Harrison et al. (2002), and was chosen because it was easy for respondents to quickly grasp how to do the exercise. For example, I asked “Would you prefer $100 today or $100 + x in one month?” The value of each subsequent delayed payment increased while the immediate payment remained constant. I inferred that the respondent’ s discount rate was higher than x% per month if they chose the immediate option and less than x% per month if they chose the delayed payment. Consequently, each payment choice was bounded by upper and lower discount rates (r), which were calculated as r = -ln (x/y)/t, where x and y are the immediate and delayed payment, respectively, and t is the time delay.  I used the midpoint of this range as the discount rate for each choice, and a respondent’ s discount rate was determined by the point at which they switched from choosing the immediate to future payment.    39  Each series had two unbounded ranges. The first unbounded range occurred if the respondent chose to receive the first future payment offered. In this case I assumed that discount rates were non-negative, and used the midpoint between 0 and the discount rate corresponding to the first payment. The second unbounded range occurred if the respondent always chose the immediate option, resulting in a discount rate that was between the last delayed payment and infinity. Although an infinite discount rate is mathematically possible, it is probably not realistic. To remain conservative, I used the discount rate corresponding to the last delayed payment to represent fishers who had ‘infinite’  discount rates. Fishers who chose the first (i.e., smallest) future payment offered were considered ‘patient’ , whereas those with ‘infinite’ discount rates were ‘impatient’ .  Fijian fishers completed two sets of choice series. Series AF had a one-month delay period while Series BF had a one-year delay. Both series had an immediate payment of FJD 400 (USD 204), which was approximately the average monthly income of fishers. Each series consisted of 13 choices, with associated annual discount rates ranging from 29 to 471 % for Series AF, and 27 to 103% for Series BF (Table 3.1). Forty-five out of 47 respondents completed both choice series.  Preliminary inspection of Fiji field data showed that the different discount rate ranges used in series AF and BF   may have affected the analysis of time consistency. Consequently, I used the same discount rate range for all choice series in the next iteration of interviews. Further, I also found that almost half of the respondents chose endpoint options for each series (i.e., always chose the immediate payment or selected the first future payment offered). This may have suggested strategic response behaviour, which other researchers have addressed by using a front end delay in the choice series. I did not get an opportunity to implement these changes in Fiji due to limited resources, but did so for interviews in Sabah.  Fishers completed three sets of choice series in Sabah. Series AS and BS had delays of 1 and 6 months, respectively. Series CS had a front end delay of one year, so that the choice was between payment in one year and one year plus one month. Each series had 14 choices. Delayed payments in all 3 series were calculated so as to correspond to the same annual discount rate range of 29 to 619%. The immediate payment for Series AS and BS was RM 100 (USD 29), while for Series CS it was RM 500 (USD 146) (Table 3.1). These payments were    40 roughly equivalent to a fisher’ s net income for one week, and one month, respectively. Of the 75 participants, 70 completed all three series. The complete choice series with corresponding discount rates for Fiji and Sabah are presented in Appendix D.  To account for the different discount rate ranges in Sabah and Fiji, I computed a ‘patience proxy’ , which was the standardised ratio of each respondent’ s discount rate to the maximum discount rate for each series (i.e., the discount rate corresponding to the largest delayed payment). A high ‘patience proxy’  represented an individual who was relatively more ‘impatient’  than one with a low proxy.  3.3 Results 3.3.1 Fishery characteristics Coral reef fisheries in both Sabah and Fiji are small-scale, making use of multiple gears to catch a wide variety of reef and reef associated species (Table 3.2). In both locations, fishing for international trade commodities such as live reef food fish species and sea cucumbers provided a valuable source of income. As both Sabah and Fiji’ s fisheries lacked catch and effort time series data, I used fishers’  perceptions about temporal changes in fish catch as an indicator of the extent of fisheries exploitation.  Sabah Fishers in Sabah were on average around 40 years old and had been fishing for 20 years. The majority of fishers had no education, and fishing was the only source of income for 75% of respondents, reflecting its importance as a source of livelihood in these villages (Table 3.2). The mean catch per unit effort (CPUE) across all gears in Sabah was 16.0±3.8 kg person-1 trip-1, although seasonal effects might mean this was not representative for the entire year (Teh et al., 2007). Eighty percent of respondents had experienced a temporal decline in fish catch, with the remainder having experienced no change in the amount of fish they caught.  Fiji Similarly, the average age of Fijian fishers was almost 40, and fishing was the only source of income for 65% of respondents. Fishers had been fishing for an average of 14 years, and their mean CPUE of 15.4±1.8 kg person-1 trip-1 was slightly lower than that recorded in Sabah. The majority of fishers (73%) noticed a temporal change in their fish catch. Of these, almost all (97%) had experienced a decline in the quantity and/or size of fish caught. Fijian fishers were    41 generally better educated and had a higher mean net monthly fishing income than those in Sabah (Table 3.2).  Cross-site comparison In general, Sabah and Fiji could be considered similar in terms of the fishery and socio- economic characteristics I measured. Tests of means showed that for fishery characteristics, only the number of years a fisher had been fishing was significantly different between sites (Welch F (1, 110) =14.66, p<0.01), with Sabah fishers fishing on average about 7 more years than Fijian fishers. Mean monthly net fishing income was significantly higher in Fiji (Welch F (1, 70) = 4.96, p<0.05). However, poverty levels at both sites were similar, as there was no significant difference in fishing poverty index between Sabah and Fiji, although the index was higher in Fiji (0.79 vs. 0.59). To assess whether the site characteristics which differed across Sabah and Fiji (number of years fishing and monthly net fishing income) were related to fishers’  discount rates, I examined the correlation between these two factors and fishers’ discount rates. I found no significant relationship in either Fiji or Sabah.  3.3.2 Annualised discount rate of fishers The mean annualised discount rate of fishers in Sabah was high, over 200% for each of the three choice series (Table 3.3). The median and mode discount rates for all series were lower because overall, 31% (n=70) of fishers chose to be ‘patient’ , i.e., always chose the first, smallest future payment option offered. In contrast, 19% were ‘impatient’ , as they chose to receive the immediate payment for all choices. A breakdown of fishers according to their choice of discount rates showed that in series AS and CS, at least half of the respondents preferred to be ‘patient’ , while for series BS, 40% of respondents preferred to be ‘patient’ (Table 3.3). Series CS, which had a front end delay of one year, had a lower but non- significant mean discount rate than series AS.  There was no significant difference in discount rates between male and female respondents in Fiji, so I report the overall average here. Fijian fishers had mean annual discount rates of 208% and 61% for Series AF (1 month delay) and Series BF (1 year delay), respectively.  The median and mode discount rates were substantially lower (Table 3.3). Compared to Sabah, a lower proportion of fishers in Fiji chose the ‘patient’  option, while over 50% of respondents chose non-endpoint options (Table 3.3).  At the same time however, a higher proportion of    42 fishers in Sabah chose the immediate option, which was consistent with having ‘infinite’ discount rates (Table 3.3).  Due to a higher number of respondents at each of the two main study areas in Sabah, I was able to examine cross-area differences in Sabah, but not in Fiji. Compared to Semporna, a larger proportion of fishers in Banggi were ‘patient’ . At the same time, the ratio of ‘patient’ to ‘impatient’  fishers in Banggi for each series was approximately 2 to 1, whereas in Semporna the ratio was less than 1 (Table 3.4). This indicated that some inherent characteristic of the two geographical areas may have been influencing discount rates. Mean discount rates in Semporna were higher than those in Banggi for all choice series (Table 3.4), but the difference was not statistically significant. A further breakdown of responses by village for the two series with different time delays (Series As and Bs) showed that the villages that consistently had the highest proportion of ‘patient’  fishers for both series were situated in Banggi.  These included the villages of Maligu, Manawali, Batu Sireh, and Damaran.  3.3.3 Patience proxy I aggregated individuals’  patience proxies across all choice series to examine if these differed between Sabah and Fiji. The proxy for Sabah was significantly lower at 0.41±0.03, compared to Fiji at 0.51±0.04 (t-test, t=2.07, p<0.05). This suggested that regardless of the absolute value of their discount rates, fishers in Sabah were relatively more willing to delay gratification than fishers in Fiji. In addition, the mode of the proxy for Sabah was 0.05, which corresponded to the lowest discount rate. In contrast, the mode for Fiji was 1, which corresponded to the ‘infinite’  discount rate option.  3.4 Discussion I aimed to determine whether the theory that open access fishers have ‘infinite’ , or very high, discount rates was applicable in real world fisheries. I found that in absolute terms, fishers in both open access and traditionally managed fisheries had high annual average discount rates of 265% and 208%, respectively, for choices involving a one month delay.  Theoretically, each fisher in an open access fishery has an infinite discount rate. Therefore, the average rate of 265% in Sabah could be interpreted as an infinite rate. If that was the case, then we would expect the Fijian discount rate to be a lot lower. The fact that the average rate    43 in Fiji was also over 200% suggests that the observed Sabah rate may not be equivalent to an ‘infinite’  rate. Moreover, when comparing the absolute value of discount rates, 200% is high, but still within the bounds of discount rates of individuals in comparable resource and socio- economic environments. For example, Akpalu (2008) also found a high average annual discount rate of 130% for fishers in Ghana. In terrestrial ecosystems, comparable annual discount rates ranging from 117-208% were estimated for farmers in Madagascar (Casse et al., 2005), while horticulturists in Bolivia had extremely high rates of between 4380-5110% (Kirby et al., 2002). Other studies found lower annual rates ranging from 13-34% for forest users in Sri Lanka (Gunatilake et al., 2007) to 35-61% for farmers in the Philippines (Lumley, 1997). It should be noted that comparing discount rates across studies is complicated due to methodological differences (Frederick, 1999; Cardenas and Carpenter, 2008), and for this study, exacerbated by a lack of studies done in developing countries involving communities dependent on natural resources.  Nevertheless, in light of the above comparisons, it appears that fishers in the open access Sabah fishery, on average, did not have ‘infinite’  discount rates. This may at first appear contrary to theory; however, a clarification of the difference between open access and non open access discount rates should help explain this. The discount rates estimated from the elicitation exercises represented fishers’  private discount rates if they fished in a non open access fishery (regardless of whether they currently fish in an open or restricted access fishery). This is because, as mentioned above, the default open access discount rate is infinity, regardless of the fisher’ s personal discount rate. Thus, a way to interpret my results is that Sabah fishers’  private discount rates are high, but not infinite. However, when faced with fishing decisions, they may be forced to use an ‘infinite’  discount rate, which although is abstract, implies that they would exploit at a rate higher than suggested by the observed 265%. This appears to be supported by the higher occurrence of infinite discounting in the open access fishery, as indicated by 30% of Sabah fishers choosing the ‘infinite’  option, compared to 20% of Fijian fishers for the one month delay option. At the same time, if Sabah’ s fisheries shifted from an open access to limited access regime, fishers may then use a discount rate of 265% when making fishing decisions.  On the whole, my results indicate that the private discount rates of small-scale fishers are high, regardless of whether a fishery is open access or managed under a customary tenure system. This contradicts expectations of a customary marine tenure system coinciding with    44 more sustainable use, and hence, longer-term outlooks for fishers. It suggests that having traditional ownership over fishing grounds does not necessarily equate to greater conservation orientation among resource users. This supports parallel arguments about the limits of using fishing rights as a means of preventing fisheries overexploitation (Clark et al., 2010a). Poverty is commonly thought to contribute to high discount rates (Mink, 1993). Therefore, I speculate the poor socio-economic conditions that exist in Sabah and Fijian fishing communities may be a reason for the high discount rates observed in both locations. I will further investigate the relationship between socio-economic conditions and discounting behaviour in Chapter 4.  Methodological differences in discount rate elicitation experiments, e.g., in the type and amount of payment offered, length of delay, and framing of the question, could have affected respondents’  choices (Frederick et al., 2003). Interviewer credibility is often a concern when conducting surveys, and with time preference surveys, lack of interviewer credibility tends to make people appear more impatient than they really are (Cardenas and Carpenter, 2008).  Other researchers have mitigated the interviewer credibility problem by using a front end delay in time preference surveys. Similarly, I used a front end delay in the third choice series (Cs) in Sabah, but still observed a similar pattern, with 80% and 65% of responses choosing the endpoint options for Series As and Cs, and Bs, respectively. Eliminating the endpoint responses would have resulted in a considerably larger difference in annual discount rates between Sabah and Fiji. For the one month delay series, average private discount rates for Sabah and Fiji would have been 303 and 198%, respectively. Other studies have attempted to control for strategic responses by providing real payments (e.g., Pender, 1996; Kirby et al., 2002). Nevertheless, in a study of horticulturists in Bolivia, Kirby et al. (2002) also found a high proportion of endpoint responses, even though they used real payments and a different binary choice format for eliciting discount rates. Overall, these factors provide some assurance that fishers in Fiji and Sabah were likely revealing their preferences, rather than providing strategic responses in the interviews.  Nevertheless, the potentially confounding methodological factors necessitate that we interpret the absolute value of estimated discount rates with caution. The different discount rate ranges at the two study sites also affected the mean rates obtained. I addressed this effect by calculating a patience proxy to facilitate comparison between Fiji and Sabah. The    45 significantly lower proxy in Sabah suggested that open access fishers did not have higher discount rates than those in the customary managed fishery. When comparing within each location, the proportion of fishers who chose the smallest future payment over the immediate option was substantially larger in Sabah. At least half the interviewed fishers chose the ‘patient’  option in Sabah for the one-month delay series, compared to approximately one- quarter in Fiji. The trend was similar for the other two series in Sabah, though of lesser magnitude for the six months delay. Although this again seemed counter-intuitive in the context of fisheries institutional theory, it may be explained by a similarity among the villages with the highest proportion of ‘patient’  fishers.  Three out of four of these villages - Batu Sireh, Manawali, and Maligu, had taken the initiative to protect the fishing grounds adjacent to their village from outside fishers. The ability to self-organise and collectively implement rules and regulations demonstrates social capital (Pretty, 2003; Grafton, 2005), and this behaviour is consistent with that displayed in community based management systems, where trust building and rule making are important institutional factors that facilitate management of fisheries resources (Jentoft et al., 1998; Lobe and Berkes, 2004; Sekhar, 2007). It may therefore be plausible that social capital is associated with the lower discount rates observed in these three villages. However, I emphasise that my findings do not permit me to infer a causal relationship between discount rates and the prevailing fishery institutional structure.  Differences in fishers’  private discount rates were also likely the result of multiple interacting drivers acting at the individual level. For instance, socio-economic, psychological, and ethical factors affect one’ s time preference (Becker and Mulligan, 1997; Lumley, 1997; Frederick et al., 2003). In particular, factors that are relevant to the marginalised context of small-scale fishing communities, such as wealth, income, and education, have been correlated with discount rates (Pender, 1996; Becker and Mulligan, 1997; Poulos and Whittington, 2000; Kirby et al., 2002). In addition, non-local factors, such as global demand for coral reef resources that undergo ‘boom and bust’  cycles, may influence a short-term outlook among fishers. More in-depth examination of these motivators of time preference could enable managers to further identify intervention policies for encouraging a longer-term outlook among fishers, and is a priority for further analysis.     46 In summary, the open access nature of many developing country reef fisheries is a major hurdle to their management and sustainability. Turning an open access fishery to one that is regulated should, theoretically, lower fishers’  discount rates (Clark, 2006). However, my results indicate that the private discount rates of small-scale inshore fishers are high, even in a non open access environment. Consequently, it suggests that emphasis on understanding what contributes to fishers’  conservation orientation may be just as important as implementing regulations to overcome the open access problem.  My study adds a new perspective to a crucial theory in fisheries economics, which surprisingly, has not been examined empirically within the context of coral reef resource conservation. While theory predicts that fishers in an open access fishery by default have an infinite discount rate, my results suggest that, contrary to expectations, private discount rates are also very high in a fishery with a formal institutional framework in place. Furthermore, it appears that a larger proportion of open access fishers are more patient than those in the customary managed fishery. This is encouraging for conservation because it implies that many open access fishers may be open to more sustainable fishing practices once the competitive nature of open access fisheries is eliminated.  There is therefore a lot of scope to carry out similar studies to further investigate the applicability of the open access-high discount rate theory. Such studies would be most useful if they use a standard methodology for eliciting discount rates.     47  Table 3.1. Payment amounts for choice series in Fiji and Sabah. Payments for series AF and BF are in FJD, and series AS, BS, and CS are in RM1. FIJI (FJD) SABAH (RM) Immediate payment for AF and BF Future payment for AF (1 month delay) Future payment for BF (1 year delay) Immediate payment for AS and BS Future payment for AS (1 month delay) Future payment for BS  ( 6 months delay) Immediate payment for AC (in 1 year) Future payment for AC (1 year +1 month)  400 420   550 100 105    134 500 525 400 435   600 100 110    177 500 550 400 450   650 100 115    231 500 575 400 465   700 100 120    299 500 600 400 480   750 100 125    381 500 625 400 495   800 100 130    483 500 650 400 510   850 100 135    605 500 675 400 525   900 100 140    753 500 700 400 540   950 100 145    929 500 725 400 555 1000 100 150 1139 500 750 400 570 1050 100 155 1386 500 775 400 585 1100 100 160 1677 500 800 400 600 1150 100 165 2017 500 825    100 170 2413 500 850 1  The exchange rate on 27 October 2010 was 1 USD = 1.9 FJD and 1 USD = 3.1 RM.             48  Table 3.2. Fishery and socio-economic characteristics of survey sites. Standard errors are provided with mean values.  Sabah Fiji FISHERY Mean no. of years fishing 21.2±1.4 14.2±1.4 Mean CPUE (across all gears, kg person-1 trip-1) 16.0±3.8 15.4±1.8 Perceived change in fishery resource Declining Declining Fishery management structure Open access Customary management Main fishing gears Handline, gillnet, speargun, traps Handline, spear/speargun, nets, traps Main target fish groups  Serranidae, Lutjanidae, Lethrinidae, Carangidae, Scombridae, Nemipteridae Lethrinidae, Lutjanidae, Serranidae,  Scombridae, Acanthuridae, Mugilidae, Mullidae  SOCIO-ECONOMIC Mean age (years) 39.6±1.4 38.9±1.7 Mean household size (no. people  household-1) 7.0±0.4 6.3±0.5 Education level    Primary 45% 32%    Secondary 7% 68%    Tertiary 1% -    None 47% - Mean net fishing income (USD month-1) 138±19 249±36 Fishing poverty index 0.59±0.08 0.79±0.12 Fishing the only source of income 75% 65%       49  Table 3.3. Summary statistics of fishers' annual discount rates (%) and breakdown of choices for payment options in Fiji and Sabah. Choice series Summary statistics for annual discount rates Proportion of respondents (%) choosing each option   Mean ± SE (%)  Median (%)  Mode (%)  Immediate1  First2  Other Sabah AS   (1 month delay) 265±33 29 29 30 51 19 BS   (6 months delay) 277±31 141 29 25 40 35 CS   (1 yr+1 month) delay) 240±32 29 29 29 53 18 Fiji AF   (1 month delay) 208±27 121 29 36 20 27 53 BF   (1 year delay) 61±5 36 24 24 51 1  Choosing the immediate payment for every choice, equivalent to having an ‘infinite’  discount rate. 2  Choosing the first, smallest future payment offered, equivalent to being ‘patient’ .   Table 3.4. Mean annual discount rate (%) and breakdown of Sabah respondents according to those who chose to receive the smallest future payment offered (‘patient’ ), and those who chose the immediate payment for all choices (‘impatient’ ). Series Annual discount rate % (Mean ± SE) Proportion of ‘patient’ respondents (%) Proportion of ‘impatient’ respondents (%)  Semporna Banggi Semporna  Banggi Semporna Banggi AS 343±57 214±38 38 55 42 24 BS 347±56 229±35 25 48 38 19 CS 318±63 187±35 39 60 48 19      50  Figure 3.1. Map of Fiji with circles marking the islands where interviews were conducted.    51  Chapter  4: Low discounting behaviour among small-scale fishers in Fiji and Sabah, Malaysia  4.1 Introduction The combined pressures of dwindling fish stocks and the need to earn a livelihood have led many fishers to engage in resource destructing activities (Pauly et al., 1989), a behaviour which is consistent with having a high discount rate, or an inability to delay economic gratification. In this context, I adopt the standpoint that low, rather than high discount rates are desirable for achieving the goal of sustainable marine resource use for both current and future generations. Consequently, understanding the socio-economic factors that are associated with a longer-term outlook among fishers may be useful for designing appropriate fisheries management and conservation policies. Pollnac and Poggie (1978) examined factors that affect small-scale fishers’  economic gratification orientation, a parallel concept to discounting. However, I do not know of any study that explicitly deals with the socio- economic predictors of fishers’  discount rates. To address this research gap, my objective in this chapter is to identify the socio-economic characteristics that are associated with fishers who have low discount rates. I continue to use the small-scale reef fisheries of Sabah, Malaysia, and Fiji as my case studies.  Factors such as wealth, income, education, and age are thought to lower individuals’  discount rates (Becker and Mulligan, 1997), although case studies involving communities in comparable socio-economic settings to Sabah and Fiji have provided inconclusive evidence. For example, Pender (1996) and Holden et al. (1998) found that income was inversely related to discount rates, but Poulos and Whittington (2000) found no relationship. Similarly, education and age have been both positively and negatively related to discount rates (Godoy and Jacobson, 1999; Kirby et al., 2002). Differences in individuals’  discount rates are also likely due to neurophysiological features (e.g., impulsiveness), personality, or learning differences (Kirby et al., 1999; Frederick et al., 2003). I focus on the socio-economic correlates of discount rates, as these bear the most relevance in terms of developing management policies for small-scale reef fisheries.  Resources from coral reefs and adjacent mangrove and seagrass ecosystems have traditionally formed the backbone of fishing economies along the coasts of Sabah and Fiji    52 (Sather, 1997; Gillett and Lightfoot, 2002). As outlined in Chapter 3, coral reef fisheries in both locations are small-scale, and make use of various manually operated gears to catch a wide variety of reef and reef associated species (Table 3.2). In addition to fish for domestic consumption, fishing for internationally traded organisms such as live reef food fish species and sea cucumbers provide a valuable source of income for fishers in both locations (Adams, 1992; Teh et al., 2005; Ovasisi, 2006).  Inshore fisheries resources in Sabah and Fiji have shown signs of overfishing, and fishers in both places have similarly noted a decrease in catch rates and the size of fish caught (Chapter 3.3.1). The depletion of coral reef fisheries resources has serious socio-economic consequences for fishing communities in both locations, as these tend to be poor, with high reliance on inshore reef resources for food and income (Teh et al., 2007; Turner et al., 2007). Faced with the pressure of having to meet immediate consumption needs from declining inshore marine resources, reef fishers in Sabah and Fiji can be expected to engage in high discounting. In this chapter, I will first identify those fishers who, contrary to expectations, have low discount rates, and then examine what socio-economic factors may predict their low discounting behaviour.  4.2 Method I used discount rates elicited from fishers in Fiji and Sabah as outlined in Chapter 3.2. Interviews in Fiji were carried out in 4 main geographical areas, whereas those in Sabah were concentrated in 2 areas (Table 4.1). The difference in discount rate ranges used in Sabah and Fiji described in Chapter 3.2.4 did not affect my current analysis because I did not focus on fishers’  absolute discount rate values. Rather, I was interested in low discounting fishers, whom I defined as those who chose the smallest future payment presented to them. I considered all other payment choices to be non-low discounting because they represented some degree of unwillingness to delay consumption.  4.2.1 Explanatory variables My study was exploratory, with the aim of identifying factors associated with low discount rates. As such, the study was not designed to determine the direction of causality between low discount rates and socio-economic variables. I split the explanatory variables into 3 groups as detailed below and summarised in Table 4.2:     53 Demographic Age – The effect of age on discount rates is ambiguous. Becker and Mulligan (1997) suggested a U shape relationship between age and discount rate, indicating that discount rates decrease through childhood, reach some minimum during adulthood, and then increase again as people grow older. In contrast, Kirby et al. (2002) found a positive relationship between age and discount rates. Education – Education is expected to focus attention on, and facilitate planning for the future (Becker and Mulligan, 1997).  It is also assumed that people with higher education levels are more likely to earn larger incomes, which are associated with lower discount rates. Household size – Households with more children may be more forward looking; thus, larger households may be associated with lower discount rates (Holden et al., 1998). On the other hand, having a higher number of dependents may be associated with higher discount rates due to the pressure of having to fulfil larger immediate consumption needs. Therefore, the effect of household size appears to be ambiguous. Socio-Economic Poverty index – People with lower income levels tend to be less patient due to the need to provide for current consumption, which leads to an inverse relationship between discount rates and income or wealth (Pender, 1996; Becker and Mulligan, 1997; Holden et al., 1998). I used a poverty index (Sumaila, 2003) as a standardised measure of income for fishers in Fiji and Sabah. This was calculated as a fisher’ s net monthly fishing income standardised to the monthly poverty line income of Fiji or Sabah. Boat ownership – Owning a boat is indicative of economic security and may thus be associated with a fisher who is more oriented towards delaying gratification (Pollnac and Poggie, 1978); the maintenance that has to be done on a boat also requires a fisher to be future oriented. Boat ownership was coded as a dichotomous variable with 0 and 1 indicating not owning and owning a boat, respectively. Relative catch – A fisher who catches more relative to others may exhibit an unwillingness to delay current consumption, everything else being equal. Relative catch was calculated as the total catch per fisher per week, standardised to the average weekly catch for all fishers in Sabah and Fiji. Presence of an alternate job or income source - Due to the seasonal and periodic nature of fishing, having another source of income or livelihood other than fishing is indicative of increased economic security, which has been found to be positively related with a deferred gratification orientation (Pollnac and Poggie, 1978).    54 Temporal decline in fish catch – I hypothesized that fishers’  perceptions about whether there had been a temporal change in their fish catch would affect their willingness to conserve fisheries resources. Perceiving a temporal decline may incite a fisher to become more aware about the need to conserve for the future. On the other hand, it may also motivate a fisher to fish harder and adopt a shorter term focus. Location specific Site – A dummy variable was used to control for site differences, with Sabah=1 and Fiji=0. Market access–Having good market access may mean that fishers are more certain that their produce will be sold for a reasonable price. This added certainty may decrease discount rates. The accessibility of each fishing village was categorised as ‘good’  or ‘poor’ , based on the distance to the nearest main town or fish market, and how accessible it was transport-wise. In both Sabah and Fiji, 8 villages (73%) were considered to have poor market access. These villages were either situated on outlying islands located away from the mainland or main island, or were isolated villages located far from main fish markets. Protection initiative – Previous findings in Chapter 3 indicated that lower discount rates may be associated with villages where inhabitants had taken the initiative to protect their village fishing grounds from outside fishers. At the study villages, forms of protection included: i) marine reserves in which no fishing was allowed; and ii) semi-protected fishing grounds which outside fishers were prevented from fishing, but villagers were allowed to fish. Three villages in Sabah had protection initiatives, as opposed to one in Fiji.  4.2.2 Analysis I fitted a binary logistic regression model using R software ( to investigate which socio-economic factors predicted the probability that fishers would choose low discount rates. I had 3 regression models:  separate models for Sabah and Fiji, and a pooled model for both Sabah and Fiji. Excluding two missing values in Sabah and Fiji led to sample sizes of 118, 73, and 45 for the pooled, Sabah, and Fiji models, respectively.  The correlations between all independent variables were examined for collinearity. I also checked for excessive multi-collinearity using Variance Inflation Factor (VIF) values. I found that market access and protection initiative (PI) were almost perfectly correlated (r=-0.98), and decided to leave market access out of the regression because my Chapter 3 findings indicated that PI may be associated with low discounting behaviour. The dummy variables for site and boat ownership were also highly correlated (r=0.8). As other studies had found    55 inter-country differences in discount rates or willingness to defer gratification (Pollnac and Poggie, 1978; Poulos and Whittington, 2000), I decided to leave site in the pooled model, and used boat ownership in the individual location models instead.  I used my results from the choice experiments to create a dichotomous indicator variable to identify low discounting individuals. I considered two options for identifying low discounters: i) one who chose the smallest future payment offered in Series A, or ii) one who chose the smallest payment offered in all series. Hyperbolic discounting behaviour may have confounded individuals’  choices for option (ii). Therefore, I decided to use option (i), which involved the same one month period delay for both Fiji and Sabah. The dichotomous dependent variable was defined as Y=1 for those who chose the smallest future payment offered in Series A, and Y=0 otherwise. For each model, I started with a “full” model containing the explanatory variables of interest. The logistic regression equation I tested was: εδγβα ++++= WZXY  [1] where X, Z and W were matrices of demographic, socio-economic and location variables, respectively, as defined in Table 4.2. The term  is a constant and the residual term, , was assumed to be normally distributed  iid ~ N (0,1). I tested this equation using the full sample of fishers from Sabah and Fiji as well as two more models on a subset of observations consisting of only fishers from either Sabah or Fiji. I controlled for site and village level variation by using dummy variables for site and protection initiative, respectively. Nevertheless, I acknowledge that lack of independence in the Y values may be a possibility due to the hierarchical nature of my sampling, and this may have resulted in overestimates of significance for these terms.  As I was undertaking exploratory analysis, it was appropriate to use a stepwise model selection procedure in both directions to identify a model which presented the “best” fit to the data. Akaike information criterion for small samples (AICc) was used to compare between models fitted to the same data. I accepted significance levels of 0.1 due to the exploratory nature of the analysis, and because I was making observations of human behaviour, which may by nature be more unpredictable (Menard, 1995). Following model selection, I assessed the model goodness of fit and predictive accuracy using the log likelihood test and receiver operating characteristic (ROC) curve. Residual and leverage plots were visually inspected to check for severe outliers and for data points which exerted excessive influence on the parameter estimates.    56  4.3 Results 4.3.1 Site comparisons Between site comparisons, all fishers Overall, 42% of fishers chose the low discount rate option in Series A; a substantially higher proportion in Sabah chose this option, compared to Fiji (51% vs. 27%). Fishers in Sabah and Fiji were similar in terms of age, household size, and dependence on fishing as the only source of income (Table 4.3). As reported in Chapter 3 (Table 3.2), the mean catch per unit effort (CPUE) across all gears in Sabah was 16.0±3.8 kg person-1 trip-1. Mean CPUE in Fiji was slightly lower, at 15.4±1.8 kg person-1 trip-1, but the difference was not statistically significant. Poverty levels at both sites were similar, with no significant difference in fishing poverty index between Sabah and Fiji, although it was higher in Fiji (0.79 vs. 0.59). Fishers in Fiji also had higher levels of education, and were less likely to own their own boats. Twenty-seven percent of fishers in Sabah resided in villages with some form of protection initiative, compared to 9% in Fiji. Within site comparisons of low and non-low discounting fishers Low discounting fishers in Sabah tended to be slightly older than non-low discounters, had slightly smaller families, less education, and a higher boat ownership rate (Table 4.3). In contrast, low discounting fishers in Fiji tended to be slightly younger than non-low discounters, achieved a higher level of education, and had a lower boat ownership rate. A larger proportion of low discounting fishers in Fiji had alternative income sources, had noticed a temporal decline in catches, and lived in accessible villages, whereas the proportion for these variables was roughly the same in Sabah. The difference in proportions between low and non-low discounting fishers who lived in PI villages was statistically significant within Sabah (Fisher’ s exact test, p<0.01), but not in Fiji (Table 4.3). There was no other outstanding factor that distinguished low discounting from non-low discounting fishers within sites, as none of the other independent variables were statistically different at the 0.05 level between low and non-low discounting fishers.  4.3.2 Logistic regression models Pooled model The stepwise procedure indicated that the following variables had non-significant relationships with fishers’  choice of a low discount rate: age, education, poverty index,    57 relative catch, temporal decline, and the presence of an alternate job or income source. I removed these from the model and tested the model using the 3 significant regressors: site, protection initiative, and household size (Table 4.4). The location specific variables (site and protection initiative (PI)) were significant predictors of a low discount rate. Although the coefficient estimate for household size was not statistically significant, I left it in the final model because a nested Chi test showed that its inclusion significantly improved the model fit. The significance of the overall model meant that the explanatory variables as a set allowed me to make better predictions of low discounting fishers than I could have made without the explanatory variables.  Holding the other explanatory variables constant, the odds of a low discount rate for a fisher from a PI village was about 5 times higher than that for a fisher from a non PI village. The odds that a Sabah fisher chose a low discount rate was about 2.6 times over that for a Fijian fisher. Having a larger household decreased the likelihood of choosing a low discount rate, although this effect was statistically insignificant. The Nagelkerke R2 value of 0.21 indicated that the model had low usefulness for predicting the probability of fishers choosing a low discount rate. The ROC measure of 0.71 suggested moderate discriminating ability. Sabah and Fiji models The overall model for Sabah was significant, and included protection initiative, boat ownership, and relative catch as significant predictors of a low discount rate. Owning a boat or living in a PI village increased the odds that a fisher would have a low discount rate, whereas high relative catch decreased the odds of choosing a low discount rate (Table 4.4). The Nagelkerke R2 of 0.32 indicated a moderate relationship between the independent and dependent variables, while the ROC measure of 0.73 indicated that the model had moderate discriminating ability.  One data point was removed from the Fiji model due to a high deviance residual, so that the final model was fitted to a sample of 44. The removal did not affect the overall model results. The Fiji model also included boat ownership and relative catch as significant predictors of low discount rates. Interestingly, in contrast to Sabah, boat ownership in Fiji decreased the odds that a fisher would choose a low discount rate, while an increase in relative catch actually increased the odds of a fisher choosing a low discount rate (Table 4.4). The model had moderately high discriminating ability with a ROC measure of 0.79, and the Nagelkerke    58 R2 of 0.26 indicated a moderate relationship between the independent and dependent variables.  4.4 Discussion I find that 42% of the fishers in this study have low discount rates, which is encouraging given the urgency of conserving coral reef resources. It also appears that the common assumption that small-scale fishers have high discount rates (Pollnac, 1985) may not be applicable in all cases. In fact, other researchers have also found that fishers’  discount rates are not excessively high (Asche, 2001; Curtis, 2002), while both Ngyuen (2009) and Poggie (1978) showed that small-scale fishers had lower discount rates compared to workers in other occupations within the same community.  The significance of the site dummy variable in the pooled model is consistent with the few other studies which have compared discount rates across developing countries (Pollnac and Poggie, 1978; Holden et al., 1998; Poulos and Whittington, 2000). It suggests that socio- economic variables I have not controlled for, or some emergent property of local conditions, (e.g., cultural values and traditions (Holden et al., 1998), or local geographical conditions (Moseley, 2001)) may account for the observed difference in the distribution of fishers choosing low discount rates in Sabah and Fiji. The use of certain fisheries management tools may also encourage fishers to develop a longer term planning horizon. For instance, based on two well-developed ITQ (individual transferable quota) systems in Iceland and New Zealand, Asche (2001) estimated that fishers’  discount rates decreased through time.  The protection initiative (PI) variable is significant in the pooled and Sabah models. All PI villages have low accessibility, as they tend to be situated on islands away from the main island or the mainland. Pomeroy et al. (2007) found that social and economic conditions prevalent in small island communities, such as constant face to face interaction among villagers, and sharing similar activities and backgrounds, increased the likelihood of villagers cooperatively undertaking a joint venture such as implementing community marine protected areas. Although I did not explicitly measure these attributes, the findings of Pomeroy et al. (2007) are consistent with studies which suggest that the presence of social capital among community members can lead to more sustainable marine resource management (Ostrom, 1990; Jentoft, 2000; Grafton, 2005). Having said this, I acknowledge that factors which I was not aware of, such as villagers’  exposure to marine education campaigns, or political    59 incentives, could have spurred villagers to initiate protection of their village fishing grounds. In any case, my results highlight the need for future research to explore the linkage between fishers’  time preference and social capital.  The same fishery related variables affect low discounting in both Sabah and Fiji, albeit with contrary effects. While the Sabah model results are consistent with other empirical studies, the Fiji model results counter expectations. Owning a boat increases the chances of a low discount rate in Sabah, which supports another study that found boat ownership to be associated with a deferred economic gratification orientation among small-scale fishers (Pollnac and Poggie, 1978). Boat owning fishers have invested in the fishery, and are therefore more likely to have a longer term perspective about the fishery compared to non- boat owners, who have no investment, hence likely have less interest or stake in the future well-being of the fishery. The fact that boat ownership is lower in Fiji may explain the lack of the positive boat ownership-low discount rate relationship observed in the Sabah model.  An increase in the relative catch variable is associated with a decrease in the probability of choosing a low discount rate in Sabah. This makes sense if a higher fish catch is due to a fisher intentionally catching more in order to gain as much as possible from the fishery today. Pursuing one’ s short-term interests that ultimately leads to outcomes which are in no one’ s long-term interest is a characteristic problem with common pool resources (Ostrom et al., 1999). In Sabah, this type of short-sighted and competitive behaviour is likely exacerbated by the essentially open access nature of small-scale fishing grounds. On the other hand, a higher catch may also indicate a better skilled fisher who catches more relative to others. If this effect is stronger than the competitive effect, the negative relationship between relative catch and low discount rate may not be present. This appears be the case in Fiji, where the presence of a customary marine tenure system may be dampening the competitive mentality that seems to occur in Sabah.  Surprisingly, demographic variables are not significant predictors of a low discount rate, even though age, education, and poverty have been correlated with discount rates in other studies of resource dependent communities in developing countries (Godoy et al., 2001; Kirby et al., 2002; Tanaka et al., 2010). The lack of statistical relationship between poverty and discount rates may be because fishers in my study are not completely cash constrained, as fishers in both Sabah and Fiji are able to obtain credit from fish buyers or from their friends and    60 families, without having to repay interest. The lack of an explicit debt load may play a role in reducing the pressure for fishers in this study to catch as much as they possibly can, but this was not completely captured in using fishing income as an indicator. The weak relationships between independent and dependent variables may also be because my models do not account for risk attitudes, ethical considerations or psychological and neurophysiological traits, which others have found to be related to discount rates (Lumley, 1997; Frederick et al., 2003). Overall, we must bear in mind that due to the relatively small sample size and exploratory nature of this study, my results should be regarded as tentative, rather than conclusive.  In summary, I find that over 40% of the fishers in my study have low private discount rates, although the proportion of patient fishers in Sabah is nearly double that in Fiji. This is encouraging for conservation of the region’ s coral reefs, particularly in Sabah, where marine protected areas are in the process of being established in both Pulau Banggi and the Semporna islands. My results suggest that location and fishery related variables differentiate low discounting from non-low discounting fishers. Policies which aim to motivate a long- term conservation orientation among fishers may therefore wish to focus on the community and fishery level, rather than on individuals. The fact that the same boat ownership and relative fish catch variables have contradictory effects on low discount rates in Fiji and Sabah cautions against implementing blanket solutions without first carefully considering local socio-cultural, economic, and ecological conditions.    61  Table 4.1. List of interview villages in Sabah and Fiji. Site Island(s) Village SABAH Banggi Batu Sireh, Damaran, Dogoton, Maligu, Manawali, Sibogo Air, Sibogo Balak  Semporna Denawan, Hampalan Laut, Mabul, Omadal  FIJI Vanua Levu Galoa, Moata,Tavea,Vunivutu  Viti Levu Nakavita, Natawarau, Seroa  Kadavu Galoa, Namouna  Yasawa Malakati, Nacula                           62 Table 4.2. Expected relationship between explanatory variables and low discount rates. Explanatory variable Variable type Variable definition Expected relationship with a low discount rate Demographic Age   Continuous Age in years +/- Education  Dichotomous 0 if not educated or primary education, 1 if secondary education or higher + Household size   Continuous No. of people in household +/- Economic Poverty index   Continuous Standardised measure of net monthly fishing income + Boat ownership  Dichotomous 1 if own a boat, 0 if do not own a boat + Relative catch  Continuous Standardised measure of catch (Kg) per fisher per week - Alternate income  Dichotomous 1 if have an alternate income, 0 if none + Temporal decline  Dichotomous 1 if perceive decline, 0 if none +/- Location Site  Dichotomous 1 if survey site is Sabah, 0 for Fiji ? Market access Dichotomous 1 if village is accessible, 0 if poor accessibility + Protection initiative  Dichotomous 1 if there is community initiative to protect fishing area, 0 if none +    63  Table 4.3. Between and within site comparison of socio-economic variables for low (DRLOW=1) and non-low (DRLOW=0) discount rate respondents. Variable SABAH (n=73) FIJI (n=45)  Non-low discounting fishers Low discounting fishers All fishers Non-low discounting fishers Low discounting fishers All fishers Age 37.3±1.9 41.9±1.9 39.6±1.4 39.4±2.4  37.0±2.4 38.9±1.7 Household size (people household-1) 8.1±0.9 7.0±0.6 7.0±0.4 6.4±0.6 5.8±0.9 6.3±0.5 No. years fishing 20.3+2.0 22.2+1.9 21.2±1.4 13.6+1.5 14.7+3.1 14.2±1.4 Relative catch  1.0±0.4 1.5±0.7 1.3±0.4 0.9±0.2 1.2±0.4 1.0±0.2 Poverty index 0.6±0.1 0.6±0.1 0.6±0.1 0.8±0.1 0.9±0.3 0.8±0.1 Education None (%) Primary (%) Secondary/Tertiary (%)  37 57 6  61 32 8  47 45 8  0 42 58  0 25 75  0 32 68 Boat ownership =1 (%) 84 95 89 39 17 9 Alternate income =1(%) 24 21 25 33 42 35 Temporal decline = 1(%) 89 89 90 52 67 56 Accessible =1 (%) 24 21 23 30 58 38 Protection initiative =1 (%)1 11 42 27 6 17 9 1 Significant difference between low and non-low discounting fishers within Sabah (Fisher’ s exact test, p<0.01).     64  Table 4.4. Regression coefficients and standard errors for logistic regression models of socio- economic variables predicting the probability of fishers choosing a low discount rate. Z values are provided in brackets, and italics indicate the odds ratio for each variable. Significance levels are indicated by asterisks. Variable Pooled Sabah  Fiji  Intercept -0.55±0.55              (-1.00) -2.83±1.22 (-2.32)** -1.40±0.55 (-2.56)** Site  0.96±0.44                 (2.18)** 2.61 n/a n/a Protection initiative 1.64±0.54                  (3.06)*** 5.16 2.85±0.99 (2.89)*** 17.29 n/a Household size  -0.11±0.07             (-1.45) 0.90 n/a n/a Boat ownership n/a 2.77±1.22 (2.27)** 15.96 -2.91±1.47 (-1.97)** 0.05 Relative catch n/a -0.17±0.10 (-1.75)* 0.84 0.81±0.42 (1.93)* 2.25  2  19.81(df=3)***  19.65 (df=3)*** 8.44 (df=2)** Log-likelihood 68.97 (df=4) -40.71 (df=4) -20.52 (df=3) Nagelkerke R2 0.21 0.32 0.26 ROC 0.71 0.71 0.79 Sample size 118 73 44 * p<0.1      ** p<0.05     ***p<0.01               65 Chapter  5: Investigating the economics of fisheries overexploitation  5.1 Introduction It is well established that fisheries are declining worldwide (Pauly et al., 2002; Worm et al., 2009), as catch rates and abundance of fish populations have decreased across ecosystems. The problem of overfishing may be attributed to the tragedy of the commons (Hardin, 1968), that is, the competition inherent in the open access nature of fisheries leads to overexploitation because no single fisher has the incentive to conserve. Even if access to a fishery was restricted, Clark (1973) showed that, in a non open access fishery, it may still be economically optimal for the sole owner of a fishery resource to drive it to extinction. According to Clark, under the condition that the value of fish exceeds the cost of fishing, a private owner will have an incentive to fish a stock to extinction if the fisher’ s discount rate, , is sufficiently higher than the biological productivity (intrinsic population growth rate, r ) of the targeted fish, such that  > 2r. Further, the incentive to overexploit a fish stock, although not necessarily to extinction, exists any time  > r.  A special condition applies to open access fisheries, in which fishers have no assurance that the fishery resource will persist into the future. This forces fishers, by default, to use a very large, or infinite discount rate when making fishing decisions (Clark 1990). A distinction is made here between the default infinite discount rate and the fisher’ s individual (private) discount rate. The private discount rate can be regarded as the rate used in a non open access condition since there is no competition to compel fishers to entirely discount the future. Subsequently, one can interpret the discount rate from Clark’ s theory as the rate under a ‘best case’  fishery scenario, as it pertains to a non open access fishery.  Although discounting is a crucial factor in understanding the exploitation of fishery resources, there are few empirical studies of fishers’  private discount rates. Clark’ s theory is especially relevant for small-scale fisheries in developing countries, many of which are open access fisheries (Béné et al., 2007). As such, I use this theory in the context of small-scale coral reef fisheries to investigate two questions: 1) How do discount rates relate to the exploitation status of fisheries? 2) What are the private discount rates of fishers?    66  To answer the first question, I compare whether applying Clark’ s theory to reef finfish fisheries worldwide matches a prior assessment of reef fisheries sustainability carried out by Newton et al. (2007). In that study, the authors found that over half of the 49 island country reef fisheries they assessed were unsustainable according to their exploitation status or ecological footprint, which measured how much annual reef fish landings exceeded maximum sustainable yield. I then address the second question by inferring the value of fishers’  private discount rates based on the exploitation status and biological productivity of reef fisheries.  5.1.1 Discount rates In this chapter, I examine Clark’ s theory using two perspectives of discount rates. The first interpretation treats the discount rate as a financial measure of the rate of return from the next best use of capital invested in the fishery. This is consistent with the figure managers apply when assessing the costs and benefits of government intervention to protect fisheries resources or the marine environment. For instance, the U.S. National Oceanic and Atmospheric Administration uses a discount rate of 7% ( Discount rates can also be viewed as an individual’ s time preference, in which case private discount rates measure an individual’ s willingness to delay economic gratification. It is reasonable to expect small-scale fishers in developing countries to have high discount rates, due to their need to fulfil immediate livelihood requirements. As outlined in Section 5.1, the private discount rates of fishers in open access fisheries are, theoretically, expected to be infinite. Consequently, there are 3 forms of discount rates that may apply to Clark’ s theory: i) official discount rate; ii) private, non open access discount rate; and iii) private open access discount rate. As the private open access discount rate is essentially known, I will use official and private non open access discount rates in this analysis.  5.1.2 Reef fisheries Coral reef fisheries occur predominantly in tropical developing countries (Fig. 5.1), and are dominated by small-scale subsistence and artisanal fishers, who use gears ranging from hook and line, nets, traps, and spears, to destructive techniques such as dynamite and cyanide (Dalzell, 1996; Jennings and Polunin, 1996). It has been estimated that coral reef fisheries make up approximately 2 to 5% of global marine fisheries catches (Pauly et al., 2003). While    67 the absolute yield from reef fisheries may not count as significant (but see Zeller et al., 2006a; 2007), their relative importance to the socio-economic well-being of coastal communities is enormous, as coral reef fisheries sustain the livelihoods and food security of millions of people worldwide (Whittingham et al., 2003; Loper et al., 2008). Despite their socio-economic importance, overexploitation is threatening the future of reef fisheries worldwide (Bell et al., 2006).  Fishing on coral reefs reduces the abundance of species targeted by the fishery, alters the size and age distribution of target fish, and possibly alters the trophic structure of reef communities and habitats (Jennings and Polunin, 1995; Jennings and Lock, 1996; Dulvy et al., 2004a; Graham et al., 2005). Overfishing also reduces reef species diversity, and can cause local extinctions of both target and non-targeted reef fish species (Roberts, 1995; Sadovy et al., 2003b; Dulvy and Polunin, 2004). Subsequently, removing the threats from overfishing is key for ensuring resilient coral reef ecosystems that can continue to sustain ecological functions and processes, as well as provide ecosystem goods and services for present and future generations.  In general, as a reef fishery becomes progressively overfished, fish catch compositions shift from being dominated by large, slow-growing predatory fish such as groupers and snappers, to one where the bulk of the catch comprises herbivores such as parrotfish and rabbitfish, or faster growing predatory species (Butler et al., 1993; Jennings and Polunin, 1996; McManus et al., 2000). The relentless expansion of the live reef food fish trade illustrates how incessant targeting of low productivity (r) species, such as humphead wrasse and large groupers, has led to the depletion and local extinction of these species from many coral reef ecosystems (Sadovy et al., 2003a; Warren-Rhodes et al., 2003; Scales et al., 2007).  5.1.3 Intrinsic growth rate The intrinsic rate of population increase (r) provides an estimate of the average population growth rate of a severely depleted stock, and represents a species’  productivity, or resilience. A population with a lower r is less productive and thus less able to tolerate high exploitation rates (Reynolds et al., 2005). The intrinsic rate of population increase can be estimated by the maximum annual reproductive rate ( ~ α ) of a fish population (Myers et al., 1997). Reproduction in fish populations may be represented as a spawner-recruit curve, which shows the relationship between parental population and the number of offspring produced.    68 The slope at the origin of the stock-recruitment relationship () represents the maximum rate of recruit production per unit of spawner abundance (kg of spawner). This slope, , can be used to approximate r. In order to do so, the slope at the origin ( ) is first standardised to the number of spawners produced per spawner per lifetime )( ∧α (Myers et al. 1997): 0= ∧ ⋅= FSPRαα ,                    [5.1] where 0=FSPR is the spawning biomass resulting from each recruit. This is then standardised to an annual rate, i.e., the number of spawners produced per spawner per year, or, the maximum annual reproductive rate ( ~α ): )1()1( 0 ~ sFs pSPRp s −⋅=−= = ∧ ααα , where ps is adult survival.             [5.2] Once the maximum annual reproductive rate ( ~α ) is calculated, the intrinsic population growth rate can then be estimated iteratively from: 0)()( ~1 =−− − αarsar epe  , where ps is adult survival and a is age at maturity.            [5.3]  Steepness parameter The intrinsic population growth rate can be used to measure the replacement rate, or potential recovery rate of exploited fish stocks, which are greatly reduced in size (Dulvy et al., 2004b). Recovery rate is affected by the steepness, or slope, of the stock-recruitment curve. The steepness (h) of the stock-recruitment relationship is defined as the proportion of recruits produced, relative to recruitment at equilibrium with no fishing, when the spawning biomass is reduced to 20% of its unfished level. The steepness parameter h ranges from 0.2 to 1, and can be expressed as a function of the lifetime number of spawners produced per spawner )( ∧α (Myers et al., 1999): ∧ ∧ + = α α 4 h                      [5.4]  A higher value of h represents higher expected productivity (recruitment) at low levels of spawning biomass. Rose et al. (2001) estimated steepness (h) values from the slope at the origin of spawner-recruit curves that had previously been fitted by Myers et al. (1999). The slope at the origin of the fitted spawner-recruit curve was converted from an annual to a lifetime rate as per Myers et al. (1999), and then used to compute a steepness value. The h values of the assessed species were then grouped into three general life-history strategies:    69 i) periodic; ii) opportunistic; and iii) equilibrium, according to Winemiller and Rose (1992). The average h value and description for each of the 3 life-history strategy groups are summarised below: i) Periodic – A periodic life history strategy is associated with fishes that have longer life spans, larger body size, and higher fecundity. These fishes tend to mature later in order to obtain a larger size that improves their chances of producing a large clutch of eggs and increasing adult survival when environmental conditions are suboptimal (e.g., groupers, Serranidae). Periodic spawners spread their reproductive effort over many years and spatial areas, so that high larval or juvenile survivorship in a particular year or spatial zone will offset bad years or zones (Winemiller and Rose, 1992). Mean h= 0.7 (range 0.55-0.87)  ii) Opportunistic – An opportunistic life history strategy is associated with high adult mortality, fast larval growth, early maturation, and an extended spawning season during which reproduction occurs frequently. This strategy is typical of small, short-lived species such as anchovies (Engraulidae) and silversides (Atherinidae). Mean h=0.56 (range 0.56-0.7)  iii) Equilibrium – Equilibrium species produce a small number of large eggs, invest in parental care of their young, and reside in relatively stable environments (e.g., gobies, Gobiidae). Mean h=0.57 (range 0.3-0.85)  5.2 Method To investigate whether discount rates are indicative of the exploitation status of reef fisheries, I assessed whether the condition  > rcorresponded to an overexploited reef fishery as determined by Newton et al. (2007) for each country under consideration. I also assessed whether Clark’ s (1973) condition for depletion,  > 2r, applied to the fisheries considered in this analysis, even though Newton et al. (2007) did not indicate whether any of the overexploited fisheries were actually depleted. I extended Clark’ s formula to the fishery level by developing a fishery level r for reef fisheries worldwide. I then applied Clark’ s framework to infer the private discount rates of fishers for each reef fishery. Estimating a fishery level intrinsic rate of increase involved two steps: i) determining the r of each species caught in the fishery; and ii) developing a fishery level r by weighting the r of all targeted fish according to the catch composition of each fishery.  5.2.1 Estimating intrinsic growth rates (r) I estimated the intrinsic population growth rate (r) for each species by iteratively solving for r in equation [3], under the condition of a steepness value h (equation [5.4]). Since h values    70 range from 0.2 to 1 (Myers et al. 2001), I estimated intrinsic growth rates for h values between this range, in 0.05 increments. I then selected the appropriate h-r estimate based on the resilience of each species, as described in Section  I used a life table to generate the population parameters required for estimating r. A life table is a discrete, age structured model that tracks the age specific reproductive and mortality rates of the female population through time. This required the following life history input parameters for each fish species: maximum age (tmax ), age of maturity (tm), natural mortality rate (M), von Bertalanffy growth parameter (K), a and b length-weight relationship parameters. These data were extracted from FishBase (Froese and Pauly 2011) for reef and reef-associated species (N=642 species). If life history parameters for a specific species were not available, the parameters for a similar species from the same genus with the same geographical distribution were used instead. If no comparable species were available, K, M, tmax and tm were estimated based on the empirical relationships detailed below (Pauly, 1980; Froese and Binohlan, 2000; Froese and Pauly, 2011):  K = -ln(1 - Lm / Linf) / (tm - t0), where t0 = a parameter of the von Bertalanffy growth function, which represents the hypothetical age at which the fish would have a length of zero.  M = 10^(0.566 - 0.718 Log(Linf) + 0.02 T),  where T= average water temperature.  tmax = t0 + 3 / K  tm = t0 - ln(1 - Lm / Linf) / K  I used the life table to estimate the following parameters: i) spawning biomass resulting from each recruit (SPRF=0) in the absence of fishing mortality; and ii) ∧ α  - The number of spawners produced by each spawner over its lifetime at low spawner abundance (assuming no density dependence). These parameters were then utilised to solve for r iteratively. Estimating the steepness parameter (h) An appropriate steepness parameter had to be chosen in estimating r for each fish species. In fishery stock assessments, h is estimated from stock and recruitment data. However, these data are limited for exploited coral reef fish stocks (Sadovy et al., 2007). As h is indicative of    71 resilience or vulnerability (Dulvy et al., 2004b), I selected a h value for each species based on its resilience status as reported in FishBase, and its life history strategy as outlined in Rose et al. (2001) and described in Section 5.1.3. I used the mean steepness values from Rose et al. (2001) to correspond to medium resilience. I then used the 90th and 10th percentile values to correspond to a high and low resilience, respectively (Table 5.1).  Coral reef fish are commonly characterised as being broadcast spawners that release a large number of eggs into the plankton (Sale, 1991). As such, I categorised relatively large, demersal coral reef fish as periodic spawners. Species which invest in parental care were considered to be equilibrium strategists. Parental investment is exhibited in species which bear live young, such as sharks and rays (Carcharhinidae and Dasyatidae). In addition, small species such as damselfish (Pomacentridae) tend to invest more in offspring, e.g., by guarding their demersal eggs, because they have to keep their vulnerable eggs out of the plankton until their young are well developed enough to settle onto appropriate habitat before dispersing. Reef associated small pelagics that are commonly targeted by small-scale reef fisheries were identified as opportunistic strategists. This group included the sardines and herrings (Clupeidae and Engraulidae), and small species of scombrids and carangids, such as Rastrelliger brachysoma and Atule mate.  5.2.2 Catch composition of reef fish catches I covered small-scale fisheries, which target reef and reef associated finfish. Coral reef is broadly defined to include reef and reef associated habitats such as lagoons, channels, mangroves, and seagrass (Dalzell et al., 1996; Unsworth and Cullen, 2010), as fish are able to traverse and inhabit more than one of these zones (Wright, 1993). Further, reef fishers generally move between habitats, depending on target species, environmental factors, and socio-economic considerations. Therefore, fish species in this study consisted of demersal reef species (e.g., groupers (Serranidae), triggerfish (Balistidae), parrotfish (Scaridae)), as well as reef associated nearshore pelagics (e.g., trevallies and scads (Carangidae), small tuna and mackerels (Scombridae)) and estuarine, seagrass, and lagoon species (e.g., mullets (Mugilidae), emperors (Lethrinidae), and rabbitfish (Siganidae)).  I searched the literature, both published and grey, to find case studies which documented the catch composition of small-scale reef fisheries. I did not distinguish between artisanal and subsistence fishing, but considered a fishery to be small-scale if it met the following criteria:    72 i) fish were caught using manually operated gears (hand line, traps, spear/speargun, gill or seine nets); ii) fishing trips lasted one day (no multi-day trips); iii) fishing was done in inshore, shallow areas (except if a steep bank was within a short distance from the fishing port or village). Studies which involved experimental fishing using local gears and were carried out in actively fished fishing grounds were included.  I did not derive reef fish catch compositions from landings statistics recorded by national fisheries departments, as there is generally a lack of, or limited, coverage of small-scale fisheries catches in national statistics (Zeller et al., 2006a). The exceptions were Anguilla and Antigua-Barbuda, for which I used landings data recorded by the Fisheries Division due to the absence of studies documenting the reef fish catch from small-scale fisheries.   Selection of countries and territories: I selected countries and territories primarily based on the study by Newton et al. (2007), and on the availability of case studies. There were 48 countries and territories in total, of which 43 were the same as those included in Newton et al.’ s study. Of the remaining 5 different countries, 3 were in Southeast Asia (Malaysia, Indonesia, and East Timor), as this region was not well represented in the Newton et al. (2007) study. Kenya and Puerto Rico were also added due to the substantial amount of research available on the respective reef fisheries (Jimenez and Sadovy, 1997; McClanahan and Mangi, 2004; Marshak et al., 2008; McClanahan et al., 2008; McClanahan et al., 2010). With the exception of Barbados and the Bahamas, the countries covered in this study were classified as developing countries according to the United Nations Human Development Index (UNDP, 2011).  Selection of time period: I attempted to find studies which took place in the past decade (2000 onwards); however, this was not possible for some countries and territories. Thus, the time period for case studies spanned from 1981 to 2010. I was able to find studies covering different time periods for 7 countries, although the studies were not all conducted at the same location or with the same gear each time. Given the declining temporal trend of world fisheries (Pauly et al., 2002; Worm et al., 2009), and lack of effective management of reef fisheries (Sale, 2008), catch compositions based on the early period case studies likely reflected conditions which have since changed for the worse. Consequently, this would make the estimated fishery level r conservative.     73  In most cases, the catch composition documented in each case study was provided as a percentage of total catch weight. In cases where the catch composition was expressed as a percentage of total number of fish caught, and the lengths of fish were recorded, I converted the number of fish to weight via the length-weight relationship: W=aLb. The a and b parameters were obtained from FishBase. If fish lengths were not recorded, the catch composition was based on the total number of fish caught; however, this only applied to 3 cases.  Case studies broke down catches to species or family level. If the catch composition was broken down to family level only, I obtained a list of the species for each family that occurred in the respective country from FishBase. I then calculated a family level r by averaging the r across the species. In cases where an ‘others’  group was included in the catch composition, I redistributed the portion of ‘others’  equally among all other families listed in the catch. In total, I had 66 case studies documenting fish catches from 48 countries (Table 5.2). Some case studies recorded reef fish catches from multiple locations or multiple gears; each location or gear specific catch composition was considered as a separate case, so that in total, I had 151 cases of fish catch compositions.  5.2.3 Assessing whether discount rates are indicative of exploitation status I tested Clark’ s theory by comparing discount rates to the fishery level r obtained for each catch composition case. A fishery was considered to be depleted if  >2r, and overfished if >r. As outlined in Section 5.1.1, there were 3 types of discount rates applicable to Clark’ s theory: official, open access, and private non open access discount rates. An open access discount rate of infinity (OA automatically implied that all reef fisheries were overexploited, since 	rfor all cases. That left official and private non open access discount rates to assess. Clark (1973) did not specify which type of discount rate to use.  As data on private discount rates were not available on a worldwide basis, I chose to use official discount rates instead. These were proxied by national bank lending rates used to meet short and long term financing needs of the private sector.  Lending rates were used because discount rates represent the rate of return from the next best use of fishing capital. Revenue obtained from fishing could potentially be lent out at the market lending rate. Lending rates were obtained from the International Monetary Fund ( Where lending rates were not available, assumed rates of 7% and 10% were used for developed and developing countries, respectively. The 7% discount rate for developed    74 countries was based on the recommended discount rate by the United States Office of Management and Budget (OMB, 1992), while the 10% was based on the recommended discount rate of 10-12% the World Bank uses for evaluating projects (Belli et al., 1998). It should be noted that in general, small-scale fishers in developing countries rarely have access to regular lending banks. Rather, it is often the case that they have to resort to high interest loans from informal lenders (Béné et al., 2007). Thus, while official discount rates may be relevant to fishery managers for conducting cost-benefit analysis, they are essentially irrelevant to small-scale fishers.  The result obtained from testing Clark’ s theory was then compared to the exploitation status from Newton et al. (2007), who classified the reef fisheries of 49 island countries and territories as being either overexploited, underexploited, or fully exploited (Table 5.2). I classified countries assessed to be ‘under/fully exploited’  or ‘fully/overexploited’  as being fully exploited. The authors also assessed whether each country’ s reef fishery was sustainable or unsustainable based on its ecological footprint (Table 5.2). For the 5 countries in this study which differed from Newton et al., I based the exploitation status on literature. Four of these countries were considered to have overexploited reef fisheries: Kenya (McClanahan and Mangi, 2004; McClanahan et al., 2008), Indonesia (Ainsworth et al., 2008), Malaysia (Biusing, 2001; Teh and Sumaila, 2007), and Puerto Rico (Matos-Caraballo, 2005). East Timor was considered to be underexploited (McWilliams, 2003; FAO, 2009), although other authors have noted that overfishing may already be occurring (Deutsch, 2003; Barbosa and Booth, 2009). As all the overexploited reef fisheries in Newton et al.’ s study also had unsustainable footprints, I found it reasonable to categorise the reef fisheries of Kenya, Puerto Rico, Indonesia, and Malaysia as unsustainable, while East Timor was considered sustainable.  Discount rates were assumed to be indicative of exploitation status if: 1) 	r or 	 r, and the reef fishery was considered overexploited by Newton et al. (2007); 2) r, and the reef fishery was considered under or fully exploited by Newton et al. (2007).  5.2.4 Inferring fishers’ private discount rates I used Clark’ s formula to infer ranges for fishers’  private discount rates (p), using the fishery intrinsic growth rate as a proxy for the status of the fishery. As specified in Section 5.1.1, these were assumed to be non open access private discount rates, and therefore could be    75 considered conservative since many developing country fisheries are open access, and hence operate under an infinite discount rate.  1) For overexploited fisheries, I assumed that fishers’  private discount rates were greater than the fishery intrinsic growth rate, but less than 2r (the point at which depletion occurs), i.e., r < p< 2r. . 2) For underexploited fisheries, r was treated as an upper bound for p.  There were 2 previous estimates of reef fishers’  private discount rates in Fiji and Sabah (Malaysia) (Chapter 3). The lowest average discount rate at both locations was 29%, which I used as a lower bound for p, such that 0.29 < p < r. 3) For fully exploited fisheries, p was set equal to r. I used the mid-point of the ranges in 1) and 2) to represent the private discount rate of fishers in the respective fishery.  5.2.5 Sensitivity analysis I evaluated the sensitivity of species r to changes in the steepness parameter (h). I conducted the sensitivity analysis using steepness values of 0.9 and 0.55. The high h of 0.9 was intended to represent a scenario of environmentally driven recruitment, while values of approximately 0.55 were found to be either a median or best estimate for h in data poor situations (Mangel et al., 2010).  5.3 Results 5.3.1 Intrinsic growth rates (r) Species level r Reef fisheries in this study targeted 642 species from 86 families, with Serranidae having the most number of species (76), followed by Lutjanidae, Carangidae, Scaridae, Acanthuridae, and Labridae, with 50, 36, 35, 35, and 35 species, respectively. The estimated intrinsic population growth rate for all fish species ranged from a minimum of 0.03 for blue spotted ray (Taeniura lymma), to a maximum of 3.58 for oxeye scad (Selar boops) (Appendix E). Averaging across all species, the family level r for common reef food fish targets such as groupers (Serranidae) was 0.44,  0.57 for snappers (Lutjanidae), 1.04 for parrotfish (Scaridae), 0.93 for surgeonfish (Acanthuridae), 0.60 for emperors (Lethrinidae), 2.21 for rabbitfish (Siganidae), and 2.05 for small mackerels or scads (Atule spp., Decapterus spp., Selar spp., Selaroides spp.).    76  Fishery level r (by country/territory) At the fishery level, the mean r across all fisheries was 0.88± 0.02, ranging from a low of 0.28, to a maximum of 1.82. (Fig. 5.2). Using family level r as a baseline, about 8% of cases fell within the r range corresponding to groupers (Serranidae); the majority of fishery intrinsic growth rates fell within the range comparable to surgeonfish (Acanthuridae) (Fig. 5.2). The top 3 records with highest fishery r were from Mauritius, Philippines, and Kenya, while the 3 lowest fishery r values were from the Turks and Caicos, Netherlands Antilles, and Seychelles. The r values for these fisheries, along with the catch compositions are provided in Table 5.3. The catch compositions for the high and low r fisheries differed substantially. Herbivorous reef fish such as rabbitfish (Siganidae), parrotfish (Scaridae), and surgeonfish (Acanthuridae) comprised the major families in high r fisheries. In contrast, carnivorous fish such as snappers and groupers contributed to the bulk of the catch in low r fisheries. A summary of the top 5 fish families making up the catch for each case is provided in Appendix F.  Temporal level r The mean r value for studies that were conducted pre-2000 was 0.82±0.05, which was lower than the mean r of 0.90± 0.03 for studies done in year 2000 and later; this difference was not statistically significant. The countries for which there was inter-temporal catch composition data did not provide conclusive evidence for this trend. The fishery intrinsic growth rates for Bermuda and Barbados were lower in an earlier study, whereas the fishery r values for Fiji remained almost constant at 0.66 and 0.64 in 2002 and 1993, respectively. Studies done in Jamaica in 1990/1991 and in 2001/2002 also had similar fishery r values of 1.2. Another study done in 2000/2001 (same location as the 1990/1991 study) actually had a slightly lower fishery r value of 1.1 compared to the pre-2000 study. Likewise, the fishery level r for the same fishery recorded at the same location in Mauritius was slightly lower in 2007 compared to 2002 (1.24 vs. 1.51). In contrast, fishery r values in Apo Island (Philippines), which is adjacent to a marine reserve, decreased temporally, from 0.96 in 1981 to 0.67 in 2001. A similar pattern was also observed for non-protected sites, with a study done in 1987 having a higher fishery r value of 1.79 compared to a 1991 study with a fishery r value of 1.08. Two studies covering predominantly similar areas in Kenya overlapped in time. The first study done in 1995-1999 had a fishery level r of 1.48, while the second study from 1998-2006    77 presented fishery level r values of 1.71 and 0.53 for the trap and handline fisheries, respectively.  Regional level r There was a significant difference in fishery r values between regions (ANOVA F=4.69, df=3, p<0.01), with the Caribbean and Southeast Asia having the lowest and highest mean fishery r values of 0.73±0.05 and 0.99± 0.07, respectively (Table 5.4).  Fishery status level r The reef fisheries assessed to be overexploited by Newton et al. (2007) had, on average, the highest r (0.92±0.03). Underexploited fisheries had a slightly lower mean fishery r value of 0.88±0.05, whereas those assessed as fully exploited had the lowest mean r of 0.77±0.05. There was no significant difference in fishery r values between exploitation status. Sustainable reef fisheries had a mean r value of 0.80±0.03, which was significantly lower than the mean of 0.96±0.04 for unsustainable fisheries (F=11.49, p<0.01).  5.3.2 Discount rates and reef fishery status The official discount rates ( for all countries or territories were considerably lower than the fishery r values (Table 5.5). According to Clark’ s formula, this would mean that none of the reef fisheries would be at threat of being overfished or depleted. In comparison, half the countries or territories were considered to be overexploited in Newton’ s study. In most cases, the mean family level intrinsic growth rates for each case were also higher than official discount rates, suggesting there was minimal threat of overfishing most types of fish. There were 13 families for which r values were lower than official discount rates (Table 5.6). These families were caught in the reef fisheries of 13 countries, and mainly consisted of the slow growing sharks and rays (Carcharhinidae, Ginglymostomatidae, Squalidae, Rhinobatidae, and Dasyatidae), or of fish that were not important food fishes (e.g., Tetraodontidae, Monacanthidae). There was one important food fish family - barracudas (Sphyraenidae), because the catch consisted primarily or entirely of Sphyraena barracuda, which is a slow growing species within this family.  5.3.3 Private discount rates The mean annual private discount rate inferred across all fisheries cases was 107±4%. The estimated private discount rate for each country or territory is provided in Table 5.5 (these are    78 averaged values if there were multiple cases per country). Overall, the mean annual private discount rate of fishers in overexploited fisheries was 137%, while those for under and fully exploited fisheries were 58% and 79%, respectively. Fishers in Southeast Asia had the highest mean annual private discount rate (145%), while those in Oceania had the lowest (89%) (Table 5.7).  5.3.4 Sensitivity Analysis Using h=0.9 increased the estimated intrinsic growth rate of all species by an average of 36%, such that the overall average r across all species increased from 0.88 to 1.18.  Changes in species specific r values ranged from no change, to a maximum increase of 89%. The maximum change applied only to 3 species of sharks, which made up insignificant proportions of the fisheries in which they were caught. Using a h of 0.55 decreased the average species r to 0.49, a declie of 58%. The subsequent effect on fishery intrinsic growth rate for the top 3 cases with highest and lowest fishery r levels are shown in Table 5.8. Individual cases within each of the high and low fishery level r groups responded similarly to changes in h values. A h of 0.9 caused the fishery level r values of the high fishery r cases to increase by 7 to 12%, while those in the low fishery r cases increased by 84 to 100%. On the other hand, a h of 0.55 decreased the fishery level r of the high fishery r cases by around 56%, whereas the low fishery r cases decreased by 18 to 26%.  5.4 Discussion The purpose of this study was to apply Clark’ s (1973) ‘economics of overexploitation’  theory to investigate two aspects of discounting in small-scale reef fisheries worldwide: first, whether discount rates are indicative of the exploitation status of reef fisheries, and second, to find out what the private discount rates of small-scale reef fishers are.  As Clark’ s theory did not specify whether to use official or private discount rates, I first applied it by comparing official discount rates to the implicit growth rate of each fishery case. This comparison indicated no threat of overfishing or depletion because for each case, the official rate was lower than the implicit growth rate of the fishery. However, only 26% of the cases corresponded to an underexploited assessment as defined by Newton et al. (2007), whereas 57% were assessed as overexploited. This suggests that official discount rates may not appropriately capture the short-term outlook fishers place on exploiting coral reef resources. Indeed, small-scale fishers rarely have access to regular bank lending rates, and    79 fishing communities in parts of Asia normally face annual interest rates of 96-144% from informal money lenders (Béné et al., 2007). Thus, using official discount rates as the basis for fisheries policy analysis may result in overly optimistic decisions that are at odds with fishers’  preferences. In contrast, all open access fisheries are, theoretically, considered overexploited, since the open access discount rate is infinite, leading to  	r for all cases. As many developing country small-scale fisheries remain open access, this suggests the need for managers to use high, rather than low, discount rates for evaluating policy options.  Comparison of official discount rates to species’  intrinsic growth rates showed that in most cases, those families likely to be overfished were elasmobranchs (sharks and rays). This raises concerns for the conservation of elasmobranchs, given the general lack of protective measures afforded to sharks, despite the decline in their populations globally (Baum et al., 2003; Robbins et al., 2006). On a more positive note, the recent creation of shark sanctuaries and bans on shark fishing in Chile, Honduras, and the Bahamas in 2011 (Pew Environment Group, 2011) indicates that shark protection and conservation is starting to take a turn for the better.  Clark (1973) showed that even a sole owner who is the only person fishing a resource has the incentive to deplete the stock if his/her discount rate is sufficiently high. It is important to note that the private discount rates inferred in this chapter relate to a non open access fishery, and are therefore conservative estimates. In reality, the private discount rates of small-scale fishers may be a lot higher due to the open access nature of many developing country small- scale fisheries. Moreover, the incentive for high discounting exists even in non open access fisheries if the regulations are not well enforced. The average annual non open access discount rate inferred from Clark’ s theory -107%- is fairly consistent with the few empirical studies of fishers’  discount rates from developing countries. For instance, Akpalu (2008) found average rates of 130% in Ghana, although fishers’  discount rates in Sabah and Fiji were higher, at 265% and 208%, respectively (Chapter 3).  Nevertheless, the overall average of 107% is high, and does not bode too well for the future sustainability of reef fisheries. The potential for unsustainable fishing is amplified by the open access nature of many reef fisheries, where fishers are forced to entirely discount the future (Clark, 1990). Given private discount rates of this magnitude, it is not surprising that slow-growing but valuable reef species such as those targeted for the live reef food fish trade    80 have been progressively fished out throughout the Indo-Pacific (Sadovy et al., 2003a; b; Scales et al., 2007). The ‘roaming bandit’  style (Warren-Rhodes et al., 2003) of fishing typical of this trade further illustrates the short-term horizon adopted by trade participants, and underlines the crucial role discounting plays in determining the future sustainability of reef fisheries.  The future sustainability of coral reef fisheries depends on fishers’  willingness to participate in conservation. However, the high private discount rates estimated here pose a challenge for achieving this goal, as they imply that fishers would rather front load and obtain benefits from the fishery now, rather than later (Sumaila, 2004). Given the poor socio-economic backdrop of many coral reef fishing communities, this is not surprising because poverty is commonly thought to contribute to high discount rates (Mink, 1993). This is demonstrated by the widespread occurrence of Malthusian overfishing, in which poverty, population pressures, and lack of economic opportunities leads to destructive fishing behaviour (Pauly et al., 1989).  The highest private discount rates were found in Southeast Asia, and this warrants particular conservation concern because the global centre of coral reef biodiversity, the Coral Triangle, is situated in this region. Oceania had the lowest private discount rates. This may reflect the presence of a traditional conservation ethic among many in the Pacific Islands (Johannes, 1978). On the other hand, in Chapter 3, I found that private discount rates of Fijian fishers were similarly high to those in Sabah, Malaysia. Moreover, overfishing is prevalent in many Pacific Island countries and territories (Kronen et al., 2010b), suggesting that private discount rates may not be as low as indicated in this analysis. The comparatively low private discount rates inferred for Oceania may be due to the fact that many of this region’ s reef fisheries were assessed as being underexploited by Newton et al. (2007), thus leading to a low inferred private discount rate. As Newton et al.’ s study covered the time period from 1997-2001, the status of these reef fisheries may have declined since then.  The discrepancy in official discount rates and fishers’  private discount rates imply that fisheries managers do not appropriately capture the perspective of fishers. Differing perceptions among marine resource stakeholders may lead to management failure (Ruddle and Hickey, 2008). For instance, the substantially lower official discount rates used by managers may result in long-term oriented policies that protect coral reef resources, e.g., marine protected areas. However, this is in conflict with the more short-term oriented outlook    81 of fishers, which in many instances leads to non-compliant behaviour such as poaching within MPAs. Importantly, this highlights the need for understanding and integrating the perspectives of different resource users during the decision making process.  On average, the reef fisheries in this study had a productivity level equivalent to a surgeonfish, which are mainly herbivorous fish that feed on benthic algae. This implies that reef fisheries are overexploited, as catches that are dominated by highly productive, herbivorous species are characteristic of overfished fisheries (Koslow et al., 1988; Dalzell et al., 1996; Friedlander and DeMartini, 2002). Cases with above average fishery level r (i.e., r > 0.88) were mostly also assessed as overexploited by Newton et al. (2007). Thus, fishery level r was fairly representative of exploitation status for this group of cases. However, a larger proportion of cases that had below average fishery level r were considered to be over, rather than underexploited, by Newton et al. (2007).  A possible reason for the weak correspondence could be because Newton’ s assessment concentrated on catch rates and sustainable yield, which was not incorporated in this study. Their assessment also covered invertebrates, the status of which may have differed from those of finfish, even within a country.  I also recognise that the fishery level r estimates represented a snapshot of reef catch compositions for a specific location and time, and this local heterogeneity may not have been captured in the country level parameters used in Newton et al.’ s assessment of exploitation status and ecological footprints.  The assumed steepness parameter used in estimating the intrinsic growth rates of reef species, and hence private discount rates, could have affected my results. I chose to assign h values based on species’  resilience because I considered this method more realistic than using the common assumption of h=0.9. Sensitivity analysis showed that using uniform assumed steepness v alues of either 0.9 or 0.55 caused the estimated r values for all species to increase or decrease by an average of 36% and 58%, respectively. Although different h assumptions result in considerable changes in estimated species r values, the overall findings of this study do not appear to be affected. This is because fishery cases with similar catch compositions respond similarly to changes in r values; thus, the general patterns about fishery level intrinsic growth rates and private discount rates remain robust to these changes. Further, another method that could have been used under similar data limited circumstances is the empirical relationship r=2M, where M is the natural mortality rate (Norse et al., 2011). The r    82 estimates generated using this technique were, in general, higher than the estimates I obtained. Therefore, I can consider my results to be conservative.  It is acknowledged that the perpetual lack of reef fisheries data should not hinder research (Johannes, 1998). Therefore, it was necessary to take the ad hoc approach of using case studies to obtain global catch composition data, even though this resulted in cases that were not uniform in time and gear type.  It also necessitated the assumption that the case studies were representative of reef fisheries at the country or territory level, although I acknowledge that seasonal and habitat differences within countries and territories could lead to location specific differences in catch composition. Nevertheless, the potential for location bias was minimized to an extent because reef fish families usually remain the same between countries, and show similar responses to fishing, even though dominant species may change between places (Clua et al., 2005).  The use of different gears could have biased the fishery r estimates; however, removing each gear sequentially did not change the overall mean fishery level r significantly. The removal of nets did lower the r for overexploited fisheries, but this decrease was not statistically significant. The finding that the earlier time period (pre-2000) fisheries had, on average, lower intrinsic growth rates suggests that the results of this study are conservative. This is because the general pattern of temporal decline in reef fisheries worldwide (Cheung et al., 2007) means it is likely that the same fisheries would be in a worse shape now (i.e., higher r), except in cases where effective management restrictions have been implemented (e.g., McClanahan and Hicks, 2011). Excluding all pre-2000 cases lowered the fishery r value for Southeast Asia, and also increased the mean fishery r value for overexploited fisheries. However, the overall exploitation status and regional patterns of fishery intrinsic growth rates was not affected, suggesting that the study results are fairly robust.  Not all countries with temporal catch composition data indicated an increase in fishery r values through time. This was surprising, particularly for the cases of the Philippines and Jamaica, as reef fisheries in these two countries have been extensively overexploited for the past two decades (Koslow et al., 1988; Lavides et al., 2010). I acknowledge that this was possibly because I did not have studies that were matched in space and time. However, a reason for the temporal decrease in fishery level r for Apo Island (Philippines) may be due to    83 its location adjacent to a marine reserve, and the subsequent benefit from spillover effects (Russ et al., 2004).  Similarly, the implementation of a fisheries management policy may have contributed to the lower fishery r estimated for the heavily fished Discovery Bay area in Jamaica in 2000 compared to 1990/1991. A programme which exchanged large for small mesh sized traps was started in the early 1990s, and a shift in the catch composition to larger and more valuable species was subsequently documented (Sary et al., 1997). These cases demonstrate that the fishery level r appears to be a consistent measure of catch composition, and may be a useful measure to use in conjunction with other indicators such as fishing effort and fish size (Clua et al., 2005) to assess the status of a fishery.  From a behavioural perspective, the cases of Apo Island and Jamaica imply that it may be possible for fishers’  conservation orientation to change (lowering of discount rates) if the proper incentives are put in place. Sumaila (2005) proposed ‘stick and carrot’  incentives as a mean of compelling those with high discount rate to apply lower discount rates to marine resources. Asche (2001) showed that fishers’  discount rates decreased following the implementation of individual transferable quotas in Iceland and New Zealand. This shows that for open access fisheries, the use of properly enforced management initiatives may be able to alter the private discount rates of fishers from infinity to the private non open access rate. This may be possible if fishers’  self-interest in the sustainability of the fishery drive compliance and cooperation with monitoring and management. The use of territorial use rights (TURFS) in the formerly open access Chilean artisanal shellfish fishery is an example of such a co-management system (Parma et al., 2003). Overall, there is little research on what motivates fishers’  willingness to delay gratification, making this an important topic for further research and understanding. In Chapter 4, I found that site and fishery specific conditions were associated with low discounting behaviour among small-scale fishers. This supports other research which emphasizes that effective management interventions first need to understand the local socio-economic context under which fishers operate (Cinner et al., 2009a).  I apply Clark’ s theory to infer private discount rates, but acknowledge that several factors may confound the direct application of this theory. A recent debate has centred on the assertion that, contrary to Clark’ s theory, it may not be economically optimal to exploit a fish    84 population to extinction, even if it has a very low intrinsic growth rate (Grafton et al., 2010). However, Clark et al. (2010b) show that the Grafton result holds only if the authors’  assumed parameter combination of discount rate, price, fishing costs, and technology are held constant through time. In reality, this is not probable, as fish prices, fishing effort costs, and technology are prone to temporal change (Tietze et al., 2005).  In summary, this study is one of the first to examine Clark’ s classic theory about the ‘economics of overexploitation’  in the context of reef fisheries worldwide.  It makes two key contributions to the small-scale reef fisheries literature. First, it estimates a fishery level intrinsic growth rate which may serve as an indicator for the status of reef fisheries. Second, it provides an estimate of reef fishers’  private discount rates worldwide, which is, as far as I am aware, the first estimate of its kind. Importantly, reef fishers’  private discount rates may in fact be a lot higher than inferred in this study, as theoretically, the open access nature of many reef fisheries implies that fishers are compelled to discount their future at an infinite rate.  Although I have focused on coral reef fisheries, the approaches developed in this study can also be readily applied to other marine fisheries. In terms of policy implications, my results show that high private discount rates of fishers, and discrepancy between these discount rates and those utilized by fishery managers, are issues which will potentially affect the status of reef and other fisheries in the present and future. This suggests that in order to improve the chances of success, fisheries policies which affect local fishers may need to be evaluated using private discount rates. Addressing the divergent future outlooks of those who manage and those who use fisheries resources is a starting point for sustainable, conservation oriented fisheries management.      85  Table 5.1. List of families grouped under periodic, equilibrium, or opportunistic life history strategies, and corresponding range of steepness parameters (h values). Life history strategy Families Steepness (h value)   High Medium Low Periodic Acanthuridae, Albulidae, Aulostomidae, Belonidae, Bothidae, Bramidae, Caesionidae, Callionymidae, Carangidae, Centropomidae, Chaetodontidae, Chirocentridae, Coryphaenidae, Dactylopteridae, Diodontidae Echeneidae, Elopidae, Ephippidae, Exocoetidae, Fistulariidae, Gempylidae, Gerreidae, Haemulidae, Holocentridae, Kuhlidae, Kyphosidae, Labridae, Lethrinidae, Lutjanidae, Malacanthidae, Monodactylidae, Mugilidae, Mullidae, Muraenidae, Nemipteridae, Ostraciidae, Paralichthyidae, Pempheridae, Pinguipedidae, Platycephalidae, Pomacanthidae, Pomacentridae, Priacanthidae, Scaridae, Sciaenidae, Scombridae, Scorpaenidae, Serranidae, Siganidae, Sillaginidae, Sparidae, Sphyraenidae, Synodontidae, Triakidae, Trichiurinae, Xiphiidae, Zanclidae 0.87 0.7 0.55  Equilibrium Balistidae, Blenniidae, Carcharhinidae, Dasyatidae, Eleotridae, Ginglymostomatidae, Monacanthidae, Plotosidae, Pomacentridae, Rajidae, Rhinobatidae, Sphyrnidae, Terapontidae,Tetraodontidae,Triacanthidae, Torpedinidae 0.85 0.57 0.3  Opportunistic Atherinidae, Carangidae1, Clupeidae, Engraulidae, Exocoetidae, Hemiramphidae, Leiognathidae, Scombridae2 0.7 0.56 0.56 1  Atule mate 2  Rastrelliger brachysoma         86 Table 5.2.Summary of catch composition case studies by country or territory, with corresponding exploitation and sustainability status as assessed by Newton et al (2007). Under=underexploited, Over=overexploited, Full=fully exploited, S=sustainable, U=unsustainable. Country No. of cases Source Study year Gear (“-” =no specified gear) Status1 Sustainable1 Anguilla 1 Anguilla Fisheries Division, pers. comm. 2010 - Under S Antigua & Barbuda 1 Horsford 2004 2004 - Over U Bahamas 1 Cushion and Sullivan Sealey 2007 2007 - Full S Barbados  2 a. Selliah et al. 2001 b. Maraj et al. 2011 1999 2010 - Net Over U Bermuda 2 a. Butler et al. 1993 b. Luckhurst and Ward 1987 1993 1975-1981 - Trap Over S British Virgin Islands 1 Boulon and Clavijo 19862  1984 Trap Over S Comoros 1 C3 Madagascar and Indian Ocean Islands Programme 2010 2010 - Over S Cook Islands 3 Pinca et al. 2009a 2007 - Under S Cuba 1 Bustamante et al. 2000 1996 - Full S Dominica 2 Gobert 2000 1990-1992 - Over U East Timor 1 Barbosa  and Booth 2009 2000 - Under3 S Federated States of Micronesia 9 a. Rhodes and Tupper 2007 b.Houk et al. 2010 c. Kronen et al. 2009c 2003 2008-2009 2006 - - - Full S Fiji 11 a. Jennings and Polunin 1995 b. Friedman et al. 2010 1992-1993 2002-2009 - - Over S French Polynesia 3 Kronen et al. 2008 2003-2006 - Under S Grenada 1 Jeffrey 2000  1986-1993 - Over S Guadeloupe 2 Gobert 2000 1990-1992 - Over U Guam 1 Myers 1993 1981-1990 Spear/ speargun Over S Indonesia 8 a. Pet-Soede and Erdmann 1998 b. May 2004 1998-1999 2002-2003 Dynamite Handline, net, trap, spear/ speargun Over3 U Jamaica 8 a. Picou-Gill et al. 1991 b. Sary et al. 2003    c. Murray and Aiken 2005 1990-1991 2000-2001    2001-2002 - Handline, net, trap, spear/ speargun - Over U Kenya 3 a.McClanahan and Mangi 2001 b.McClanahan et al. 2010 1995-1999 1998-2006 - Handline, trap Over3 U Kiribati 5 Awira et al. 2008  2004 - Under U Madagascar 8 a. Laroche and Ramananarivo 1995 b. Laroche et al. 1997 c.  Davies et al. 2009 1990  1991 2008 Handline, net - Handline, net, spear/ speargun Over U Malaysia 2 a. Teh et al. 2005 b. Teh et al. 2007 2004 2005 Handline - Over3 U Maldives 1 Sattar 2008  2006-2007 - Under S Marshall Islands 4 Pinca et al. 2009b 2007 - Under S Martinique 2 Gobert 2000 1987-1992 Trap Over U Mauritius 4 a. Lynch et al. 2003  b. Hardman et al. 2008 2002  2007 Handline, net, trap Net Over U Mayotte 1 Minet and Weber 1992 1989 - Under U Nauru 1 Sauni et al. 2007  2005 - Full S Netherlands Antilles 2 a. Toller and Lundvall 2008 b. Johnson 2010 2007 2008 - Trap Under S New Caledonia 5 Kronen et al. 2009a 2003-2007 - Under S    87 Country No. of cases Source Study year Gear (“-” =no specified gear) Status1 Sustainable1 Niue 1 Kronen et al. 2009b 2005 - Under S Palau 3 Friedman et al. 2009a 2007 - Full S Papua New Guinea 3 McClanahan and Cinner 2008  2001-2002 Handline, net, spear/ speargun Under U Philippines  9 a. Amar et al. 1996 b. Campos et al. 1994 b. Maypa et al. 2002 1991-1992 1987-1988 1980-2001 - - - Over U Puerto Rico 1 Jimenez and Sadovy 1997 1990 Trap Over3 U Reunion 1 Biais and Taquet 1992 1988 - Over U Samoa 9 a. Craig et al. 2008 b. Vunisea et al. 2008 2002 2005 - - Over U Seychelles 1 Pistorius and Taylor 2009 1995-2007 - Full S Solomon Islands 5 Pinca et al. 2009c 2006 - Full S Sri Lanka 2 Anandappa and Simon 2008  2006 - Over U Tonga 5 Friedman et al. 2009b  2001-2008 - Over S Trinidad and Tobago 1 Cesar and Oxenford 2005  2001 - Over U Turks and Caicos 2 a. Keegan 1986 b. School of field studies 1981 2008 Trap - Full U Tuvalu 4 Sauni et al. 2008 2004-2005 - Under S U.S. Virgin Islands 1 Garrison et al. 2004  1992-1994 Trap Over S Vanuatu 3 Friedman et al. 2008  2003 - Under S Wallis and Futuna 3 Kronen et al. 2009d  2005 - Under S 1Source: Newton et al. 2007. 2  Study stated that the catch composition from British Virgin Islands  was similar to that recorded for this study of the US Virgin Islands. 3 See text for source of exploitation status.   Table 5.3. Top and bottom 3 records with highest and lowest fishery level r values, respectively. Country/territory Fishery r  Major families Source Highest fishery r Mauritius 1.82  Siganidae (52%), Acanthuridae (17%), Scaridae (10%), Mullidae (8%), Serranidae (4%), others (10%) Lynch et al. 2003 Philippines 1.79 Siganidae (48%), Scaridae (19%), Labridae (11%), Gobiidae (5%), others (17%) Campos et al. 1994 Kenya  1.71  Siganidae (36%), Lethrinidae (27%), Scaridae (24%), Mullidae (6%), others (8%) McClanahan et al. 2010 Lowest fishery r Turks and Caicos  0.30  Serranidae (33%), Lutjanidae (26%), Haemulidae (15%), Labridae (9%), other (17%) School of Field Studies (pers. comm.) 2011 Netherlands Antilles  0.31  Lutjanidae (46%), Haemulidae (15%), Serranidae (15%), Balistidae(10%), other (14%) Toller & Lundvall 2008 Seychelles  0.36 Serranidae (56%), Lutjanidae (24%), Lethrinidae (20%) Pistorius & Taylor 2009          88 Table 5.4. Mean fishery r value by region Region No.  of cases Mean ± std. error Caribbean 31 0.73±0.05 Indian Ocean1 22 0.98±0.09 Oceania 78 0.88±0.03 Southeast Asia1 20 0.99±0.07 1  Significant mean difference with Caribbean (Tukey test, p<0.05).     89   Table 5.5 Comparison of official discount rate (with fishery r (r) and corresponding exploitation status. Inferred private discount rates ( are provided in the last column. Country  r >r? >2r? Status1  Anguilla 0.10 0.49 no no Under 0.39 Antigua and Barbuda 0.11 0.61 no no Over 0.92 Bahamas 0.05 0.28 no no Full 0.28 Barbados 0.10 0.96 no no Over 1.43 Bermuda 0.07 0.51 no no Over 0.77 British Virgin Islands 0.10 0.58 no no Over 0.87 Comoros 0.11 0.76 no no Over 1.15 Cook Islands 0.10 0.91 no no Under 0.60 Cuba 0.10 0.48 no no Full 0.48 Dominica 0.10 0.68 no no Over 1.02 East Timor 0.10 0.83 no no Under2 0.56 Federated States of Micronesia 0.16 0.96 no no Full 0.89 Fiji 0.06 0.69 no no Over 1.04 French Polynesia 0.10 1.09 no no Under 0.69 Grenada 0.10 0.30 no no Over 0.45 Guadeloupe 0.10 0.84 no no Over 1.26 Guam 0.07 0.74 no no Over 1.11 Indonesia 0.16 1.13 no no Over2 1.69 Jamaica 0.18 1.00 no no Over 1.50 Kenya 0.13 1.12 no no Over2 1.86 Kiribati 0.10 0.60 no no Under 0.44 Madagascar 0.28 1.10 no no Over 1.49 Malaysia 0.13 0.73 no no Over2 1.10 Maldives 0.14 0.65 no no Under 0.47 Marshall Islands 0.10 0.88 no no Under 0.58 Martinique 0.07 1.14 no no Over 1.50 Mauritius 0.21 1.38 no no Over 2.06 Mayotte 0.07 0.63 no no Under 0.46 Nauru 0.10 1.15 no no Full 1.15 Netherlands Antilles 0.09 0.63 no no Under 0.46 New Caledonia 0.07 0.90 no no Under 0.60 Niue 0.07 1.15 no no Under 0.72 Palau 0.10 0.88 no no Full 0.88 Papua New Guinea 0.11 1.18 no no Under 0.74 Philippines 0.15 0.88 no no Over 1.41 Puerto Rico 0.10 0.58 no no Over2 0.86 Reunion 0.07 0.57 no no Over 0.85 Samoa 0.10 1.08 no no Over 1.62 Seychelles 0.10 0.36 no no Full 0.36 Solomon Islands 0.10 0.76 no no Full 0.76 Sri Lanka 0.07 0.74 no no Over 1.10 Tonga 0.12 0.84 no no Over 1.26 Trinidad and Tobago 0.10 0.37 no no Over 0.71 Turks and Caicos 0.07 0.37 no no Full 0.50 Tuvalu 0.10 0.76 no no Under 0.53 U.S. Virgin Islands 0.07 0.54 no no Over 0.80 Vanuatu 0.08 1.29 no no Under 0.79 Wallis and Futuna 0.07 0.81 no no Under 0.55 1Source: Newton et al. (2007) 2 See text for source of exploitation status    90   Table 5.6. Summary of families which may potentially be overfished (r< for each country or territory. Country/territory  Families with r< Anguilla 0.10 Ginglymostomatidae Antigua and Barbuda 0.11 - Bahamas 0.05 - Barbados 0.10 - Bermuda 0.07 - British Virgin Islands 0.10 - Comoros 0.11 - Cook Islands 0.10 - Cuba 0.10 Carcharhinidae, Dasyatidae Dominica 0.10 - East Timor 0.10 Carcharhinidae, Rajidae Federated States of Micronesia 0.16 Bolbometopon muricatum Fiji 0.06 - French Polynesia 0.10 - Grenada 0.10 - Guadeloupe 0.10 - Guam 0.07 - Indonesia 0.16 Sphyraenidae Jamaica 0.18 Muraenidae Kenya 0.13 Sphyraenidae Kiribati 0.10 - Madagascar 0.28 Congridae, Dasyatidae, Fistularidae, Muraenidae,Rhinobatidae, Tetraodontidae, Torpedinidae, Carcharhinidae, Sphyraenidae, Malaysia 0.13 Dasyatidae, Monacanthidae, Rhinobatidae, Carcharhinidae Maldives 0.14 - Marshall Islands 0.10 - Martinique 0.07 - Mauritius 0.21 Monacanthidae, Tetraodontidae, Fistularidae Mayotte 0.07 - Nauru 0.10 - Netherlands Antilles 0.09 Carcharhinidae New Caledonia 0.07 - Niue 0.07 - Palau 0.10 - Papua New Guinea 0.11 - Philippines 0.15 - Puerto Rico 0.10 Dasyatidae, Ginglymostomatidae Reunion 0.07 - Samoa 0.10 Carcharhinidae, Dasyatidae Seychelles 0.10 - Solomon Islands 0.10 - Sri Lanka 0.07 - Tonga 0.12 - Trinidad and Tobago 0.10 - Turks and Caicos 0.07 - Tuvalu 0.10 - U.S. Virgin Islands 0.07 - Vanuatu 0.08 - Wallis and Futuna 0.07 -    91 Table 5.7. Private discount rates (annual rate per year) by region and exploitation status.  Mean ± std. error Region Caribbean 1.02±0.09 Indian Ocean  1.42±0.14 Oceania 0.89±0.04 Southeast Asia 1.45±0.11 Status Overexploited 1.37±0.05 Underexploited 0.58±0.02 Fully exploited 0.79±0.05  Table 5.8. Results of sensitivity analysis of different h assumptions on fishery level intrinsic growth rates for the 3 cases with highest and lowest fishery level r. Fishery case Fishery level r  Current h h=0.9 h=0.55 Top 3 highest fishery level r Mauritius 1.82 2.00 0.79 Philippines 1.79 1.92 0.78 Kenya 1.71 1.91 0.76 Top 3 lowest fishery level r Bahamas 0.28 0.56 0.23 Turks and Caicos 0.30 0.57 0.23 Netherlands Antilles 0.31 0.57 0.23    92   Figure 5.1. Map of global coral reef distribution. Source: coral reef distribution data from UNEP-WCMC (   Figure 5.2. Distribution of fishery r values for all cases. The horizontal lines show the mean family intrinsic growth rate for Serranidae (groupers), Lutjanidae (snappers), Acanthuridae (surgeonfish), and Scaridae (parrotfish).     93  Chapter  6: Conclusion  The linking thread through the chapters in this thesis is an attempt to better understand the dynamics of discounting among small-scale fishers. I have examined this under different contexts: contrasting fisheries management systems, socio-economic conditions, and stages of fisheries exploitation. I started the thesis by using a case study to illustrate why small-scale fisheries warrant our attention. I then addressed my first and second research questions (what is the discount rate of fishers, and under what conditions fishers have low discount rates) in Chapters 3 and 4. Finally, I expanded my case studies to a global study of reef fisheries in order to further investigate the magnitude of private discount rates, and the relevance of discount rates to fisheries exploitation status. In the next section I discuss how my results have addressed the research questions I set out with, and the contribution and significance of my results given the current state of knowledge about fishers’  private discount rates. I then outline some limitations of this research, and identify directions for future research.  6.1 What are the private discount rates of fishers? Small-scale fishers have commonly been characterised as having high discount rates (Pollnac, 1985), although there is sparse empirical evidence to support this. Using two different approaches, I found that the absolute values of reef fishers’  private discount rates were indeed high (Chapter 3 and 5). In Sabah and Fiji, average annual discount rates for receiving one month delayed payments were 265% and 208%, respectively (Chapter 3), while an average of 107% was inferred for reef fishers worldwide (Chapter 5). As noted in the respective chapters, these represent fishers’  private discount rates in non open access situations, and are consistent in magnitude with the average of 130% Akpalu (2008) estimated for fishers in Ghana. On the other hand, they are substantially higher than the median of 30-40% estimated for fishers in Ireland (Curtis, 2002). While there are two other studies on fishers’  discount rates in developing countries (Fehr and Leibbrandt, 2008; Nguyen, 2009), the authors did not document the absolute rates. Thus, given the present limited studies, it appears that, at least in developing countries, a probable range for fishers’  annual discount rates is from 100% to 300%. This suggests that overall, fishers discount the future heavily, i.e., they are impatient and want to obtain benefits from the fisheries resources immediately.      94 6.2 Under what conditions do fishers exhibit low discounting? There are numerous studies in other fields examining the demographic and socio-economic correlates of discount rates (e.g., Pender, 1996; Becker and Mulligan, 1997; Godoy and Jacobson, 1999; Poulos and Whittington, 2000; Kirby et al., 2002). Although there are contradictory findings about the direction of relationship between explanatory variables and discount rates, it seems that variables such as age, education, and income are common correlates of discount rates. In contrast, I found that demographic and socio-economic variables were not useful predictors of low discounting behaviour among fishers (Chapter 4). Instead, fishery (boat ownership and relative fish catch) and site (protection initiative and study site) level variables predicted low discount rates among fishers in Sabah and Fiji. Some of these results are consistent with the literature. For instance, Pollnac and Poggie (1978) also found boat ownership to be correlated with a delayed economic gratification orientation, which is analogous to a low discount rate, among fishers.  The significance of the protection initiative variable supports my result from Chapter 3, wherein fishers from villages where there was some form of fishing ground protection had lower discount rates. As discussed in Chapter 4, this brings up the possibility that social capital may contribute to a conservation orientation, but I was not able to further address this with the data I had. There are additional questions arising from this chapter’ s results that I could not resolve. First, I could not single out what local socio-cultural, economic, or environmental factors made fishers in Sabah more likely to choose low discount rates compared to Fiji. A related question is why the same variables had contradictory effects on low discounting in Sabah and Fiji. Overall, my results did not provide a conclusive answer to my research question. Rather, the best answer I can provide is that low discounting behaviour appears to depend on local socio-cultural, economic, or ecological factors that I have not managed to fully capture in my model. This reiterates other research which point to the importance of understanding local conditions in order to understand why fishers make the fishing allocation decisions that they do (Cinner et al., 2009a; McClanahan et al., 2009a). Thus, although inconclusive, this outcome provides a starting point for pursuing further investigations.  6.3 Are discount rates reflective of the exploitation status of fisheries? Chapter 5 presented evidence that official discount rates, which are the rates that fisheries managers most likely use for evaluating projects, do not fully reflect the short term    95 perspective fishers employ when making fishing decisions, and subsequently offer too optimistic a perception about the state of fisheries. Instead, fishers’  private discount rates are more likely to capture the exploitation status. This provides two unique insights for management: First, there is a clear discrepancy in the future outlooks of managers and fishers which has to be rectified, or at least articulated in the decision making process. The results in Chapter 5 support an earlier observation by Clark (1985), who stated that disagreement over fisheries policies stems from a difference in discount rates. Second, to address this discrepancy, managers should consider using private discount rates to evaluate policies which affect local fishers, as this will likely increase the chances of acceptance and compliance by the fishing community. A more comprehensive comparison of discount rates and exploitation status will require a more uniform set of data on reef fish catch composition across space, time, and gears; this in turn points to the need for improved monitoring of reef fisheries catches, a long standing challenge in reef fisheries management (Johannes, 1998; Zeller et al., 2006a; 2007).  6.4 Significance and limitations of thesis research The overall significance of this research is that it has shown that small-scale fishers’  discount rates are generally high, a theme that has been repeated in the fisheries literature, but has limited empirical support. In addition, my study has contributed several unique insights about fishers’  private discount rates, an area of research that has not received much attention in the fisheries literature to date. The first is that discount rates in both open access and traditionally managed fisheries are similarly high, which is counter-intuitive to theoretical expectations. It is further intriguing that a greater proportion of fishers in the open access system chose low discount rates. Second, my research is perhaps one of the few to examine the socio-economic predictors of discounting behaviour among fishers. This research adds a new perspective to the growing body of literature on fishers’  behaviour, which is increasingly recognised as being necessary for more successful fisheries management (Salas and Gaertner, 2004). Lastly, Chapter 5 provides a comparative analysis between discount rates and fisheries exploitation status on a worldwide basis that has not been examined to date.  An important application of this study is that it provides fishery managers with an approximate range for fishers’  discount rates, so that managers can now take steps to address, and close the gap between official and fishers’  private discount rates. For instance, managers can consider using private discount rates instead of official rates to assess regulations that    96 will affect fishers. Another application of this study is that the estimation approach I took in Chapter 2 can readily be applied by fishery managers or other researchers to quantify the socio-economic contribution of small-scale as well as commercial fisheries.  The strength of this research is the integration of economic and ecological concepts to examine fishers’  discount rates. It is, as far as I am aware, the first to investigate fisher’ s private discount rates on a case study basis as well as globally through the use of experimental and inferential techniques. Using a combination of social, experimental economics, quantitative, and theoretical approaches, I have obtained similar estimates for the magnitude of fishers’  private discount rates, thus providing some assurance about the validity of my results.  There are, however, limitations to this study. One limitation is that I cannot identify the drivers that lead to the observed discount rates, as the comparative study in Chapter 3 and logistic regression model in Chapter 4 were not intended to examine causative factors of discount rates. Instead, the results raise interesting suggestions for correlates or predictors of discount rates that warrant further investigation. Further, a limitation of using case studies in Chapter 5 is the possibility of confounding factors on the composition of reef fish catches, due to the lack of uniformity in time and gear. However, I showed that these factors had minimal influence on the results.  I acknowledge that due to the lack of data, parts of my analysis necessitated making assumptions which may have been subjective. However, as indicated in Chapters 2 and 5, the general picture of my findings remains consistent, even though the magnitude of the outcomes may change under different assumptions of certain variables.  6.5 Future directions This thesis has investigated a relatively new research topic on which there is sparse literature for comparative purposes. I see two main research topics on fishers’  discount rates that would benefit from further research. First, there is a need for more empirical studies that elicit the private discount rates of fishers in order to establish a more representative range of fishers’ discount rates. The diverse discount rate elicitation methods currently in use in experimental economics makes it difficult to compare discount rates across studies (Cardenas and Carpenter, 2008). Thus, further empirical studies of fishers’  discount rates should follow a consistent elicitation methodology.    97  Second, there is a lot of scope for research on the correlates or drivers of fishers’  discounting behaviour. The results from the logistic model in Chapter 4 indicate that we need a better understanding of the interaction between local conditions and fishers’  discounting behaviour. Further research that aims to find a clearer picture of what factors predict or are correlated to fishers’  discount rates will need to identify explanatory variables that are more representative, or are better able to capture local differences in socio-cultural, environmental, or economic conditions. In Chapters 3 and 4, I suggested that social capital attributes may be correlated to lower discount rates, and this relationship warrants further research. As such, existing frameworks for assessing social capital (e.g., Rudd, 2004; Grafton, 2005) can be incorporated into surveys to elicit fishers’  discount rates. Once the correlates of fishers’  discount rates are established, future research can also focus on examining the causal relationship between socio-economic factors and discount rates. The developmental of such a model will enable managers to identify appropriate intervention policies which directly address the drivers of high or low discount rates.  One aspect of discounting I have not addressed in my thesis is the psychological and ethical aspects of time preference. Such studies may also provide a better insight into why fishers may make the fishing effort decisions that they do. Bearing in mind the complexity of human behaviour, and the multiple factors that affect our perceptions and future outlooks, it may be some time before we can obtain a comprehensive conceptual understanding about fishers’ discount rates. However, the studies I mentioned above may be a starting point towards this goal.     98 Bibliography  Abdul Mannan, Q., 1982. Sabah fishermen and their economy. Ko-Nelayan, Kota Kinabalu, 90 p.  Adams, T., 1992. Resource aspects of the Fiji bêche-de-mer industry. SPC Bêche-de-mer Information Bulletin 4, 13-16.  Adeogun, O., Abohweyere, P., Ogunbadejo, H., Tanko, A., Jim-Saiki, L., 2009. Economic viability of small-scale marine capture fisheries in the Bonny Area, Rivers State, Nigeria. Marine Resource Economics 24, 195-203.  Ainsworth, C., Sumaila, U., 2005. Intergenerational valuation of fisheries resources can justify long-term conservation: a case study in Atlantic cod (Gadus morhua). Canadian Journal of Fisheries and Aquatic Science 62, 1104-1110.  Ainsworth, C., Pitcher, T., Rotinsulu, C., 2008. Evidence of fisher depletions and shifting cognitive baselines in Eastern Indonesia. Biological Conservation 141, 848-859.  Akpalu, W., 2008. Fishing regulations, individual discount rate, and fisherman behaviour in a developing country fishery. Environment and Development Economics 13, 591-606.  Allison, E., Ellis, F., 2001. The livelihoods approach and management of small-scale fisheries. Marine Policy 25, 377-388.  Almada-Villela, P.C., 1997. Pilot fisheries socio-economic survey of two coastal areas in eastern Sabah, in: Fowler, S.L., Reed, T.M., Dipper, F.A. (Eds.), Elasmobranch biodiversity, conservation and management: Proceedings of the international seminar and workshop, Sabah, Malaysia, July 1997. IUCN, Gland and Cambridge, pp. 33-44.  Amadja, S., 2008. Discount rate estimation and the role of time preference in rural household behavior: Disease prevention in India and forest management in the US. PhD thesis, North Carolina State University. 300 p.     99 Amar, E., Cheong, R., Cheong, M., 1996. Small-scale fisheries of coral reefs and the need for community-based resource management in Malalison Island, Philippines. Fisheries Research 25, 265-277.  Anandappa, S., Simon, D., 2008. Poverty, livelihoods and reef fishing practices at Hikkaduwa, Sri Lanka. CEDAR Research Paper No.45. Department of Geography University of London, Egham, Surrey, 45 p.  Andrew, N., Béné, C., Hall, S., Allison, E., Heck, S., Ratner, B., 2007. Diagnosis and management of small-scale fisheries in developing countries. Fish and Fisheries 8, 227-240.  Asche, F., 2001. Fishermen's discount rates in ITQ systems. Environmental and Resource Economics 19, 403-410.  Awira, R., Friedman, K., Sauni, S., Kronen, M., Pinca, S., Chapman, L., Magron, F., 2008. Kiribati country report: Profiles and results from survey work at Abaiang, Abemama, Kuria and Kirimati: (May to November 2004). Pacific Regional Oceanic and Coastal Fisheries Development Programme (PROCFish/C/CoFish) / Secretariat of the Pacific Community, Noumea, 313 p.  Barbier, E., Burgess, J., Folke, C., 1994. Paradise Lost? The Ecological Economics of Biodiversity. Earthscan Publications, London.  Barbosa, M., Booth, S., 2009. Timor-Leste's fisheries catches (1950-2009): Fisheries under different regimes, in: Zeller, D., Harper, S. (Eds.), Fisheries catch reconstructions: Islands, Part 1. University of British Columbia Fisheries Cetnre, Vancouver, pp. 39-50.  Baum, J., Myers, R., Kehler, D., Worm, B., Harley, S., Doherty, P., 2003. Collapse and conservation of shark populations in the Northwest Atlantic. Science 299, 389-392.  Becker, G., Mulligan, C., 1997. The endogenous determination of time preference. Journal of Economic Psychology 23, 291-316.     100 Bell, J., Ratner, B., Stobutzki, I., Oliver, J., 2006. Addressing the coral reef crisis in developing countries. Ocean and Coastal Management 49, 976-985.  Belli, P., Anderson, J., Barnum, H., Dixon, J., Tan, J., 1998. World Bank Handbook on economic analysis of investment operations 1998. World Bank, Washington D.C., 159 p.  Bellwood, D., Hughes, T., Folke, C., Nystrom, M., 2004. Confronting the coral reef crisis. Nature 429, 827-833.  Béné, C., 2006. Small-scale fisheries: assessing their contribution to rural livelihoods in developing countries. FAO Fisheries Circular No. 1008. FAO, Rome, 46 p.  Béné, C., Macfadyen, G., Allison, E., 2007. Increasing the contribution of small-scale fisheries to poverty alleviation and food security. FAO Fisheries Technical Paper 481. FAO, Rome, 125 p.  Béné, C., Hersoug, B., Allison, E., 2010. Not by rent alone: analysing the pro-poor functions of small-scale fisheries in developing countries. Development Policy Review 28, 325-358.  Berkes, F., Mahon, R., McConney, P., Pollnac, R., Pomeroy, R., 2001. Managing small-scale fisheries: alternative directions and methods. IDRC, Ottawa.  Biais, G., Taquet, M., 1992. La peche locale aux abords de La Reunion. Reperes Ocean No. 2-199. Institut francais de recherche pour l'exploitation de la mer (IFREMER), Centre de Brest, 77 p.  Biusing, R.E., 2001. Assessment of coastal fisheries in the Malaysian Sabah portion of the Sulu-Sulawesi Marine Ecoregion (SSME). WWF Malaysia, Sabah, 719 p.  Bonfil, R., 1994. Overview of world elasmobranch fisheries. FAO Fisheries Technical Paper 341. FAO, Rome, 119 p.     101 Boulon, R., Clavijo, I., 1986. Utilization of the Virgin Islands Biosphere Reserve by artisanal fishermen. Biosphere Reserve Research Report No. 11. US National Park Service/VIRMC, St. Thomas, USVI, 37 p.  Bustamante, G., Chiappone, M., Kelly, J., Lowe, A., Sullivan Sealey, K., 2000. Fish and fisheries in Guantanamo Bay, Cuba: recommendations for their protection. Proceedings of the 51st Gulf and Fisheries Institute, 51, 242-257.  Butler, J., Burnett-Herkes, J., Barnes, J., Ward, J., 1993. The Bermuda fisheries: a tragedy of the commons averted? Environment 35, 6-15, 25-33.  C3 Madagascar and Indian Ocean Islands Programme, 2010. Socio-economic assessment and identification of potential sites for community-based coral reef management in the Comoros. A report submitted to the NOAA coral reef conservation program, 22 p.  Cabanban, A.S., Biusing, R.E., 1999. Coral reef fisheries and their contribution to marine fish production in Sabah, Malaysia, in: Cabanban, A.S., Philips, M. (Eds.), Proceedings of the workshop on aquaculture of coral reef fishes and sustainable reef fisheries. Institute of Development Studies, Kota Kinabalu.  Campos, W., Del-Norte Campos, A., McManus, J., 1994. Yield estimates, catch, effort and fishery potential of the reef flat in Cape Bolinao, Philippines. Journal of Applied Ichthyology 10, 82-95.  Cardenas, J., Carpenter, J., 2008. Behavioural development economics: Lessons from field labs in the developing world. Journal of Development Studies 44, 311-338.  Casse, T., Neilsen, U., Ranaivoson, S., Randrianamarivo, J.R., 2005. Farmer strategies and forest conservation: a case study from south-western Madagascar. International Journal of Social Economics 32, 704-716.  Cesar, E., Oxenford, H., 2005. Testing a collapsible trap design in the deep demersal trap fishery of Tobago, Eastern Caribbean. Proceedings of the 56th Gulf and Caribbean Fisheries Institute, 56, 117-132.    102  Chabris, C., Laibson, D., Morris, C., Schuldt, J., Taubinsky, D., 2008. Individual laboratory- measured discount rates predict field behavior. Journal of Risk and Uncertainty 37, 237-269.  Charles, A.T., 1998. Living with uncertainty in fisheries: analytical methods, management priorities and the Canadian groundfishery experience. Fisheries Research, 37, 37-50.  Charles, A., 2011. Good practices for governance of small-scale fisheries, in: Chuenpagdee (Ed.), Contemporary visions for world small-scale fisheries, Eburon, pp. 285-298.  Cheung, W., Watson, R., Morato, T., Pitcher, T., Pauly, D., 2007. Intrinsic vulnerability in the global fish catch. Marine Ecology Progress Series 333, 1-12.  Cinner, J., Marnane, M., McClanahan, T., 2005. Conservation and community benefits from traditional coral reef management at Ahus Island, Papua New Guinea. Conservation Biology 19, 1714-1723.  Cinner, J., Daw, T., McClanahan, T., 2009a. Socioeconomic factors that affect artisanal fishers' readiness to exit a declining fishery. Conservation Biology 23, 124-130.  Cinner, J., McClanahan, T., Daw, T., Graham, N., Maina, J., Wilson, S., Hughes, T., 2009b. Linking social and ecological systems to sustain coral reef fisheries. Current Biology 19, 206- 212.  Clark, C., 1973. The economics of overexploitation. Science 181, 630-634.  Clark, C., 1985. Bioeconomic modelling and fisheries management. Wiley, New York.  Clark, C., 1990. Mathematical bioeconomics: the optimal management of renewable resources. J. Wiley, New York.  Clark, C., 2006. The worldwide crisis in fisheries economic models and human behavior. Cambridge University Press, Cambridge, UK.     103 Clark, C.W., Munro, G.R., 1975. The economics of fishing and modern capital theory: a simplified approach. Journal of Environmental Economics and Management, 2, 92-106.  Clark, C.W., Munro, G.R., Sumaila, U.R., 2005. Subsidies, buybacks and sustainable fisheries. Journal of Environmental Economics and Management, 50, 47-58.  Clark, C., Munro, G., Sumaila, U., 2010a. Limits to the privatization of fishery resources. Land Economics 86, 209-218.  Clark, C., Munro, G., Sumaila, U., 2010b. Limits to the privatization of fishery resources: Reply. Land Economics 86, 614-618.  Clua, E., Beliaeff, B., Chauvet, C., David, G., Ferraris, J., Kronen, M., Kulbicki, M., Labrosse, P., Letourneur, Y., Pelletier, D., Thebaud, O., Leopold, M., 2005. Towards multidisciplinary indicator dashboards for coral reef fisheries management. Aquatic Living Resources 18, 199-213.  Cooke, A., Polunin, N., Moce, K., 2000. Comparative assessment of stakeholder management in traditional Fijian fishing grounds. Environmental Conservation 27, 291-299.  Cooke, F.M., 2008. Land development and livelihood vulnerability in Sabah. Paper presented at the 17th Biennial conference of the Asian Studies Association of Australia, Melbourne 1-3 July 2008, Monash University.  Costanza, R.E., 1991. Ecological Economics: The Science and Management of Sustainability. Columbia University Press, New York.  Costello, C., Gaines, S., Lynham, J., 2008. Can catch shares prevent fisheries collapse? Science 321, 1678-1681.  Craig, P., Green, A., Tuilagi, F., 2008. Subsistence harvest of coral reef resources in the outer islands of American Samoa: modern, historic and prehistoric catches. Fisheries Research 89, 230-240.     104 Cunningham, S., Neiland, A., Arbuckle, M., Bostock, T., 2009. Wealth-based fisheries management: using fisheries wealth to orchestrate sound fisheries policy in practice. Marine Resource Economics 24, 271-287.  Curtis, J., 2002. Estimates of fshermen’ s personal discount rate. Applied Economic Letters 9, 775-778.  Cushion, N., Sullivan Sealey, K., 2007. Landings, effort and socio-economics of a small- scale commercial fishery in the Bahamas. Proceedings of the 60th Gulf and Caribbean Fisheries Institute, 60, 162-166.  Dalzell, P., 1996. Catch rates, selectivity and yields of reef fishing, in: Polunin, N., Roberts, C. (Eds.), Reef Fisheries. Chapman and Hall, London, pp. 161-192.  Dalzell, P., Adams, T., Polunin, N., 1996. Coastal fisheries in the Pacific islands. Oceanography and Marine Biology 34, 395-531.  Davies, T., Beajara, N., Tregenza, T., 2009. A socio-economic perspective on gear-based management in an artisanal fishery in southwest Madagascar. Fisheries Management and Ecology 16, 279-289.  Department of Statistics Malaysia, 1995. Population and Housing Census of Malaysia 1991, Putrajaya, Malaysia. Available at Accessed 10 April 2011.  Department of Statistics Malaysia, 2007. Yearbook of Statistics Sabah 2007. Department of Statistics, Kota Kinabalu.  Department of Statistics Malaysia, 2009a. Population, Household and Living Quarters Malaysia 2009. Department of Statistics Malaysia, Putrajaya.  Department of Statistics Malaysia, 2009b. Report on the census of agricultural establishments: fisheries. Department of Statistics Malaysia, Putrajaya. Available at Accessed 10 April 2011.     105 Department of Statistics Malaysia, 2011. GDP by state and kind of economic activity for the year 2009. Available at Accessed 10 April 2011.  Deutsch, C., 2003. Coral Reefs in East Timor: An investigation of the impacts of fishing on reef fish abundance. EM 531 unpublished project report. Available at Accessed 4 October 2011.  Dietz, T., Ostrom, E., Stern, P., 2003. The struggle to govern the commons. Science 302, 1907-1912.  Dulvy, N., Freckleton, R., Polunin, N., 2004a. Coral reef cascades and the indirect effects of predator removal by exploitation. Ecological Letters 7, 410-416.  Dulvy, N., Ellis, J., Goodwin, N., Grant, A., Reynolds, J., Jennings, S., 2004b. Methods of assessing extinction risk in marine fishes. Fish and Fisheries 5, 255-276.  Dulvy, N., Polunin, N., 2004. Using informal knowledge to infer human-induced rarity of a conspicious reef fish. Animal Conservation 7, 365-374.  Dyck, A., Sumaila, U., 2010. Economic impact of ocean fish populations in the global fishery. Journal of Bioeconomics 12, 227-243.  FAO, 2004. The state of the world fisheries and aquaculture 2004. FAO, Rome, 153 p.  FAO, 2009. National Fishery Sector Overview Timor Leste Fishery and Aquaculture Country Profile. Available at Accessed 8 August 2011.  Fearnside, P.M., 1989. Forest Management in Amazonia: The need for new criteria in evaluating development options. Forest Ecology and Management 27, 61-79.     106 Fehr, E., Leibbrandt, A., 2008. Cooperativeness and impatience in the tragedy of the commons. Institute for Empirical Research in Economics Working Paper No. 378. University of Zurich, 23 p.  Fiji Islands Bureau of Statistics, 2007. Fiji national census of population 2007. Available at Accessed 10 February 2009.  Fiji Islands Bureau of Statistics, 2009. Fiji Facts and Figures as at 1st July 2009. Available at Accessed 10 February 2009.  Fisher, H., 2000. A socio-economic assessment of coastal communities of Pulau Banggi Sabah, East Malaysia. Master of Science thesis, University of Minnesota.  Foo, L.H., Khor, G.L., Tee, E.S., Prabakaran, D., 2006. Dietary intake of adolescents in a rural fishing community in Tuaran district, Sabah. Malaysian Journal of Nutrition 12, 11-21.  Frederick, S., 1999. Discounting, time preference, and identity. Ph.D. dissertation thesis, Carnegie Mellon University.  Frederick, S., Loewenstein, G., O'Donoghue, T., 2003. Time discounting and time preference: a critical review, in: Loewenstein, G., Read, D., Baumeister, R. (Eds.), Time and Decision. Russell Sage Foundation, New York, pp. 13-86.  Friedlander, A., DeMartini, E., 2002. Contrasts in density, size, and biomass of reef fishes between the northwestern and the main Hawaiian island: the effects of fishing down apex predators. Marine Ecology Progress Series 230, 253-264.  Friedman, K., Pakoa, K., Kronen, M., Chapman, L., Sauni, S., Vigliola, L., Boblin, P., Magron, F., 2008. Vanuatu country report: Profile and results from survey work at Paunangisu village, Moso Island, Uri and Uripiv islands and the Maskelyne Archipelago (July to December 2003). Pacific Regional Oceanic and Coastal Fisheries Development Programme (PROCFish/C/CoFish) / Secretariat of the Pacific Community, Noumea, 357 p.     107 Friedman, K., Kronen, M., Pinca, S., Lasi, F., Pakoa, K., Awira, R., Boblin, P., Tardy, E., Chapman, L., Magron, F., 2009a. Palau country report: Profiles and results from survey work at Ngarchelong, Ngatpang, Airai and Koror (April to June 2007). Pacific Regional Oceanic and Coastal Fisheries Development Programme (PROCFish/C/CoFish) / Secretariat of the Pacific Community, Noumea, 411 p.  Friedman, K., Pinca, S., Kronen, M., Boblin, P., Chapman, L., Magron, F., Vunisea, A., Labrosse, P., 2009b. Tonga country report: profiles and results from survey work at Ha'atafu, Manuka, Koulo and Lofanga (November and December 2001; March to June 2002; April to June, September and October 2008). Pacific Regional Oceanic and Coastal Fisheries Development Programme (PROCFish/C/CoFish) / Secretariat of the Pacific Community, Noumea, 369 p.  Friedman, K., Kronen, M., Vunisea, A., Pinca, S., Pakoa, K., Magron, F., Chapman, L., Sauni, S., Vigliola, L., Tardy, E., Labrosse, P., 2010. Fiji Islands country report: profiles and results from survey work at Dromuna, Muaivuso, Mali and Lakeba (September to November 2002, April to June 2003, June and July 2007, and February 2009). Pacific Regional Oceanic and Coastal Fisheries Development Programme (PROCFish/C/CoFish) / Secretariat of the Pacific Community, Noumea, 467 p.  Froese, R., Binohlan, C., 2000. Empirical relationships to estimate asymptotic length, length at first maturity and length at maximum yield per recruit in fishes, with a simple method to evaluate length frequency data. Journal of Fish Biology 56, 758-773.  Froese, R., Pauly, D., 2011. FishBase. Available at Accessed 6 September 2011.  Garrison, V., Rogers, C., Beets, J., Friedlander, A., 2004. The habitats exploited and the species trapped in a Caribbean island trap fishery. Environmental Biology of Fishes 71, 247- 260.  Gillett, R., Lightfoot, C., 2002. The contribution of fisheries to the economies of Pacific island countries. Asian Development Bank, Forum Fisheries Agency and World Bank, Manila, Philippines, 217 p.    108  Gobert, B., 2000. Comparative assessment of multispecies reef fish resources in the Lesser Antilles. Fisheries Research 44, 247-260.  Godoy, R., Jacobson, M., 1999. Covariates of private time preference: a pilot study among the Tsimane' Indians of the Bolivian rain forest. Evolution and Human Behaviour 20, 249- 256.  Godoy, R., Kirby, K., Wilkie, D., 2001. Tenure security, private time preferences, and use of natural resources among lowland Bolivian Amerindians. Ecological Economics 38, 105-118.  Gordon, H., 1954. The economic theory of a common property resource: The fishery. Journal of Political Economy 62, 124-142.  Grafton, R., Arnason, R., Bjorndal, T., Campbell, D., Campbell, H., Clark, C., Connor, R., Dupont, D., Hannesson, R., Hilbron, R., Kirkley, J., Kompas, T., Lane, D., Munro, G., Pascoe, S., Squires, D., Steinshamn, S., Turris, B., Weninger, Q., 2006. Incentive-based approaches to sustainable fisheries. Canadian Journal of Fisheries and Aquatic Sciences 63, 699-710.  Grafton, R., Kompas, T., Hilborn, R., 2010. Limits to the privatization of fishery resources: Comment. Land Economics 86, 609-613.  Grafton, R.Q., 2005. Social capital and fisheries governance. Ocean and Coastal Management 48, 753-766.  Graham, N., Dulvy, N., Jennings, S., Polunin, N., 2005. Size-spectra as indicators of the effects of fishing on coral reef fish assemblages. Coral Reefs 24, 118-124.  Greboval, D., Munro, G., 1999. Overcapitalization and excess capacity in world fisheries: underlying economics and methods of control, in: Greboval, D. (Ed.), Managing fishing capacity: selected papers on underlying concepts and issues. FAO Fisheries Technical Paper 386. FAO, Rome, pp. 1-48.     109 Gunatilake, H.M., Wickramasinghe, W.A.R., Abeygunawardena, P., 2007. Time preference and natural resource use by local communities: the case of Sinharaja forest in Sri Lanka. Asian Development Bank, Manila, Philippines, 36 p.  Guyader, O., Berthou, P., Koustikopoulos, C., Alban, F., Demaneche, S., Gaspar, M., Eschbaum, R., Fahy, E., Tully, O., Reynal, L., Albert, A., 2007. Small-scale coastal fisheries in Europe. IFREMER Final Report No. FISH/2005/10. IFREMER, 447 p.  Hardin, G., 1968. The tragedy of the commons. Science 162, 1243-1248.  Hardisty, D., Weber, W., 2009. Dicounting future green: money vs. the environment. Journal of Experimental Psychology: General 138, 329-340.  Hardman, E., Blais, F., Desire, M., Raffin, J., Perrine, S., Meunier, S., 2008. Annual report on the status of the artisanal seine net fishery of Rodrigues 2007. Shoals Rodrigues, Point Monier, Rodrigues, 53 p.  Harrison, G., Lau, M., Williams, M., 2002. Estimating individual discount rates in Denmark: A field experiment. The American Economic Review 92, 1606-1617.  Holden, S., Shiferaw, B., Wik, M., 1998. Poverty, market imperfections and time preferences: of relevance for environmental policy? Environment and Development Economics 3, 105-130.  Horsford, I., 2004. Status and trends in the fisheries sector of Antigua-Barbuda. Ministry of Agriculture, Lands, Marine Resources and Agro-Industries. Available at Accessed 29 June 2011.  Houk, P., Rhodes, K., Mcilwain, J., Cuetos-Bueno, J., 2010. Comparative assessment of commercial coral reef fisheries across Micronesia: the need for improved management. A report for the Nature Conservancy. The Nature Conservancy Micronesia Program, 42 p.  Hughes, T.P., 1994. Catastrophes, phase shifts, and large-scale degradation of a Caribbean coral reef. Science 265, 1547-1551.    110  Hutchings, J.A., Reynolds, J.D., 2004. Marine fish population collapses: consequences for recovery and extinction risk. BioScience, 54, 297-309.  ICTSD, 2006. Fisheries, international trade and sustainable development: policy discussion paper. ICTSD Natural Resources, International Trade and Sustainable Development Series. International Centre for Trade and Sustainable Development, Geneva, Switzerland. 114 p.  Institut Penyelidikan Marin Borneo, 2003. Laporan kajian mengenai pengetahuan dan pandangan masyarakt Pulau Banggi terhadap penubuhan Taman Tun Mustapha. Institut Penyelidikan Marin Borneo, Universiti Malaysia Sabah, Kota Kinabalu, 36 p.  Jeffrey, C., 2000. Annual, coastal and seasonal variation in Grenadian demersal fisheries (1986-1993) and implications for management. Bulletin of Marine Science 66, 305-319.  Jennings, S., Polunin, N., 1995. Comparative size and composition of yield from six Fijian reef fisheries. Journal of Fish Biology 46, 28-46.  Jennings, S., Lock, J., 1996. Population and ecosystem effects of reef fishing, in: Polunin, N., Roberts, C. (Eds.), Reef Fisheries. Chapman and Hall, London, pp. 193-218.  Jennings, S., Polunin, N., 1996. Impacts of fishing on tropical reef ecosystems. Ambio 25, 44-49. Jentoft, S., McCay, B., Wilson, D., 1998. Social theory and fisheries co-management. Marine Policy 22, 423-436.  Jentoft, S., 2000. The community: a missing link of fisheries management. Marine Policy 24, 53-60.  Jimenez, A., Sadovy, Y., 1997. The effect of fish trap mesh size on species composition and catch value in western Puerto Rico. Proceedings of the 44th Gulf and Caribbean Fisheries Institute, 44, 5-30.     111 Johannes, R., 1978. Traditional marine conservation methods in Oceania and their demise. Annual Review of Ecology and Systematics 9, 349-364.  Johannes, R., 1998. The case for data-less marine resource management: examples from tropical nearshore finfisheries. Trends in Ecology and Evolution 13, 243-245.  Johannes, R., 2002. The renaissance of community-based marine resource management in Oceania. Annual Review of Ecology and Systematics 33, 317-340.  Johnson, A., 2010. Reducing bycatch in coral reef trap fisheries: escape gaps as a step towards sustainability. Marine Ecology Progress Series 415, 201-209.  Jul Larsen, E., 2003. Analysis of effort dynamics in the Zambian ishore fisheries of Lake Kariba, in: Jul Larsen, E., Kolding, J., Overa, R., Nielsen, J., van Zwieten, P. (Eds.), Management, co-management or no management? Major dilemmas in southern African freshwater fisheries. FAO Fisheries Technical Paper 426/1. FAO, Rome, pp. 233-252.  Kamaruddin, R., Abdul Rashid, Z., 2007. Structural change, linkages vs. leakages in the Malaysian economy: 1991-2000. Paper presented at the 16th International Conference on Input-Output Techniques, Istanbul, 2-6 July 2007. Available at Accessed 20 December 2010., Istanbul.  Keegan, W., 1986. The ecology of Lucayan Arawak fishing practices. American Antiquity 51, 816-825.  Kennelly, S.J., Broadhurst, M.K., 2002. By-catch begone: changes in the philosophy of fishing technology. Fish and Fisheries, 3, 340-355.  Kirby, K., Petryb, N., Bickelb, W., 1999. Heroin addicts have higher discount rates for delayed rewards than non-drug-using controls. Journal of Experimental Psychology 128, 78- 87.     112 Kirby, K., Godoy, R., Reyes-Garcia, V., Byron, E., Apaza, L., Leonard, W., Perez, E., Vadez, V., Wilkie, D., 2002. Correlates of delay-discount rates: Evidence from Tsimane' Amerindians of the Bolivian rain forest. Journal of Economic Psychology 23, 291-316.  Kolb, J.A., Scheraga, J.D., 1990. Discounting the benefits and costs of environmental regulations. Journal of Policy Analysis and Management 9, 381-390.  Koslow, J., Hanley, F., Wicklund, R., 1988. Effects of fishing on reef fish communities at Pedro Bank and Port Royal Cays, Jamaica. Marine Ecology Progress Series 43, 201-212.  Kramer, R., Simanjuntak, S., Liese, C., 2002. Migration and fishing in Indonesian coastal villages. Ambio 31, 367-372.  Kronen, M., Boblin, P., Friedman, K., Pinca, S., Magron, F., Awira, R., Pakoa, K., Lasi, F., Tardy, E., Vigliola, L., Chapman, L., 2009a. New Caledonia country report: Profile and results from survey work at Ouassé, Thio, Luengoni, Oundjo and Moindou (March, April and November 2003; January, February, April, June, August and November 2004; April and May 2005; January to March 2006; and January and February 2007). Pacific Regional Oceanic and Coastal Fisheries Development Programme (PROCFish/C/CoFish) / Secretariat of the Pacific Community, Noumea, 409 p.  Kronen, M., Fisk, D., Pinca, S., Magron, F., Friedman, K., Boblin, P., Awira, R., Chapman, L., 2009b. Niue country report: Profile and results from in-country survey work (May to June 2005). Pacific Regional Oceanic and Coastal Fisheries Development Programme (PROCFish/C/CoFish) / Secretariat of the Pacific Community, Noumea, 173 p.  Kronen, M., Friedman, K., Boblin, P., Chapman, L., Vunisea, A., Lassi, F., Awira, R., Pakoa, K., Magron, F., Tardy, E., Pinca, S., 2009c. Federated States of Micronesia country report: Profiles and results from survey work at Yyin and Riiken (Yap) and Piis-Panewu and Romanum (Chuuk) (April - May 2006). Pacific Regional Oceanic and Coastal Fisheries Development Programme (PROCFish/C/CoFish) / Secretariat of the Pacific Community, Noumea, 367 p.     113 Kronen, M., Tardy, E., Boblin, P., Chapman, L., Lasi, F., Pakoa, K., Vigliola, L., Friedman, K., Magron, F., Pinca, S., 2009d. Wallis et Futuna - Rapport de Pays : profils et résultats des enquêtes réalisées à Vailala, Halalo, Leava et Vele (août - décembre 2005 et mars 2006). Pacific Regional Oceanic and Coastal Fisheries Development Programme (PROCFish/C/CoFish) / Secretariat of the Pacific Community, Noumea, 346 p.  Kronen, M., Vunisea, A., Magron, F., McArdle, B., 2010a. Socio-economic drivers and indicators for artisanal coastal fisheries in Pacific island countries and territories and their use for fisheries management strategies. Marine Policy 34, 1135-1143.  Kronen, M., Magron, F., McArdle, B., Vunisea, A., 2010b. Reef finfishing pressure risk model for Pacific Island countries and territories. Fisheries Research 101, 1-10.  Laroche, J., Ramananarivo, N., 1995. A preliminary survey of the artisanal fishery on coral reefs of the Tulear region (southwest Madagascar). Coral Reefs 14, 193-200.  Laroche, J., Razanoelisoa, J., Fauroux, E., Rabenevanana, M., 1997. The reef fisheries surrounding the coastal citites of Madagascar. Fisheries Management and Ecology 4, 285- 299.  Lavides, M., Polunin, N., Stead, S., Tabaranza, D., Comeros, M., Dongallo, J., 2010. Finfish disappearance around Bohol, Philippines inferred from traditional ecological knowledge. Environmental Conservation 36, 235-244.  Leete, R., 2008. Sabah's human development progress and challenges. United Nations Development Programme, Kuala Lumpur.  Lobe, K., Berkes, F., 2004. The padu system of community-based fisheries management: change and local institutional innovation in south India. Marine Policy 28, 271-281.  Loewenstein, G., Read, D., Baumeister, R. (Eds.), 2003. Time and decision: economic and psychological perspectives on intertemporal choice. Russell Sage Foundation, New York.     114 Loper, C., Pomeroy, R., Hoon, V., McConney, P., Pena, M., Sanders, A., Sriskanthan, G., Vergara, S., Pido, M., Vave, R., Vieux, C., Wanyonyi, I., 2008. Conditions along the world's tropical coasts: 2008. National Oceanic and Atmospheric Administration, Global Coral Reef Monitoring Network, Conservation International.  Luckhurst, B., Ward, J., 1987. Behavioural dynamics of coral reef fishes in Antillian fish traps at Bermuda. Proceedings of the 38th Gulf and Caribbean Fisheries Institute, 38, 528- 546.  Lumley, S., 1997. The environment and the ethics of discounting: An empirical analysis. Ecological Economics 20, 71-82.  Lynch, T., Edwards, A., Gell, F., Hooper, T., Gaspard, R., Milazar, G., Blais, F., Raffin, J., Perrine, S., Pierre Louis, J., 2003. Preliminary report on the status of artisanal large net, basket trap and line fisheries of Rodrigues. Shoals Rodrigues, Pointe Monier, Rodrigues, 50p.  Mangel, M., Brodziak, J., DiNardo, G., 2010. Reproductive ecology and scientific inference of steepness: a fundamental metric of population dynamics and strategic fisheries management. Fish and Fisheries 11, 89-104.  Mangi, S.C., Roberts, C.M., Rodwell, L.D., 2007.Reef fisheries management in Kenya: Preliminary approach using the driver-pressure-state-impacts-response (DPSIR) scheme of indicators. Ocean and Coastal Management, 50, 463-480.  Mantjoro, E., 1996. Traditional management of communal-property resources: the practice of the Sasi system. Ocean & Coastal Management 32, 17-37.  Maraj, V., Cox, S., Oxenford, H., 2011. The small-scale seine fishery of Barbados: description and current contribution to the fishing industry. CERMES Technical Report No. 42. Centre for Resource Management and Environmental Studies. University of the West Indies, Cave Hill, Barbados, 26 p.  Markandya, A., Pearce, D., 1991. Development, the environment, and the social rate of discount. The World Bank Research Observer 6, 137-152.    115  Marshak, A., Hill, R., Sheridan, P., Scharer, M., Appeldoorn, R., 2008. In-situ observations of Antillean fish trap contents in southwest Puerto Rico: relating catch to habitat and damage potential. Proceedings of the 60th Gulf and Caribbean Fisheries Institute, 60, 447-453.  Matos-Caraballo, D., 2005. Comprehensive census of the marine commercial fishery of Puerto Rico 2002. Proceedings of the 56th Gulf and Caribbean Fisheries Institute, 56, 97-110.  Matthews, E., Veitayaki, J., Bidesi, V., 1998. Fijian villagers adapt to changes in local fisheries. Ocean and Coastal Management 38, 207-224.  May, D., 2004. Reef fishing activity in the Kaledupa stakeholder area. Operation Wallacea. Available at VITY%20IN%20THE%20KALEDUPA%20STAKEHOLDER%20AREA.pdf. Accessed 29 June 2011.  Maypa, A., Russ, G., Alcala, C., Calumpong, H., 2002. Long term trends in yield and catch rates of the coral reef fishery at Apo Island, central Philippines. Marine & Freshwater Research 53, 207-213.  McClanahan, T., Glaesel, H., Rubens, J., Kiambo, R., 1997. The effects of traditional fisheries management on fisheries yields and the coral reef ecosystems of southern Kenya. Environmental Conservation 24, 105-120.  McClanahan, T., Mangi, S., 2001. The effect of a closed area and beach seine exclusion on coral reef fish catches. Fisheries Management and Ecology 8, 107-121.  McClanahan, T., Mangi, S., 2004. Gear-based management of a tropical artisanal fishery based on species selectivity and capture size. Fisheries Management and Ecology 11, 51-60.  McClanahan, T., Cinner, J., 2008. A framework for adaptive gear and ecosystem-based management in the artisanal coral reef fishery of Papua New Guinea. Aquatic Conservation: Marine and Freshwater Ecosystems 18, 493-507.    116  McClanahan, T., Hicks, C., Darling, E., 2008. Malthusian overfishing and efforts to overcome it on Kenyan coral reefs. Ecological Applications 18, 1516-1529.  McClanahan, T., Castilla, J., White, A., Defeo, O., 2009a. Healing small-scale fisheries by facilitating complex socio-ecological systems. Reviews in Fish Biology and Fisheries 19, 33- 47.  McClanahan, T., Cinner, J., Graham, N., Daw, T., Maina, J., Stead, S., Wamukota, A., Brown, K., Venus, V., Polunin, N., 2009b. Identifying Reefs of Hope and Hopeful Actions:Contextualizing Environmental, Ecological, and Social Parameters to Respond Effectively to Climate Change. Conservation Biology 23, 662-671.  McClanahan, T., Kaunda-Arara, B., Omukoto, J., 2010. Composition and diversity of fish and fish catches in closures and open-access fisheries of Kenya. Fisheries Management and Ecology 17, 63-76.  McClanahan, T., Hicks, C., 2011. Changes in life history and ecological characteristics of coral reef fish catch composition with increasing fishery management. Fisheries Management and Ecology 18, 50-60.  McManus, J., Menez, L., Kesner-Reyes, K., Vergara, S., Ablan, M., 2000. Coral reef fishing and coral-algal phase shifts: implications for global reef status. ICES Journal of Marine Science 57, 572-578.  McManus, J.W., 1997. Tropical marine fisheries and the future of coral reefs: a brief review with emphasis on Southeast Asia. Coral Reefs 16, S121-S127.  McWilliams, A., 2003. Timorese Seascapes. The Asia Pacific Journal of Anthropology 3, 6- 32.  Menard, S., 1995. Applied logistic regression analysis. SAGE Publications, Thousand Oaks, California.     117 Minet, J., Weber, J., 1992. Perpectives de developpement du secteur des peches a Mayotte. IFREMER.  Mink, S., 1993. Poverty, population, and the environment. The World Bank, Washington D.C.  Mohd. Ariff, M.R., 1999. Dari pemungutan tripang ke penundaan udang: Sejarah perkembangan perusahaan perikanan di Borneo Utara, 1750-1990. Penerbit Universiti Malaya, Kuala Lumpur.  Moseley, W., 2001. African evidence on the relation of poverty, time preference and the environment. Ecological Economics 38, 317-326.  Murray, A., Aiken, K., 2006. Artisanal fishing in Jamaica today: a study of a large fishing site. Proceedings of the 57th Gulf and Caribbean Fisheries Institute, 57, 215-233.  Myers, R., 1993. Guam's small boat based fisheries. Marine Fisheries Review 55, 117-128.  Myers, R., Mertz, G., Fowlow, P., 1997. Maximum population growth rates and recovery times for Atlantic cod, Gadus morhua. Fishery Bulletin 95, 762-772.  Myers, R., Bowen, K., Barrowman, N., 1999. Maximum reproductive rate of fish at low population sizes. Canadian Journal of Fisheries and Aquatic Sciences 56, 2404-2419.  Newton, K., Cote, I., Pilling, G., Jennings, S., Dulvy, N., 2007. Current and future sustainability of island coral reef fisheries. Current Biology 17, 655-658.  Nguyen, Q., 2009. Do fishermen have different preferences?: Insights from an experimental study and household data MPRA Paper No. 16012. Munich Personal RePEc Archive. Available at Accessed 15 October 2010.  Noh, K.M., Tai, S.Y., 2001. Fisheries resource accounting for Sabah and Sarawak, in: Tai, S.Y., Radam, A. (Eds.), Selected readings on economic analysis of fisheries and natural resources. Universiti Putra Malaysia, Kuala Lumpur, pp. 106-138.    118 Norse, E., Brooke, S., Cheung, W., Clark, M., Ekeland, I., Froese, R., Gjerde, K., Haedrich, R., Heppell, S., Morato, T., Morgan, L., Pauly, D., Sumaila, R., Watson, R., 2011. Sustainability of deep-sea fisheries. Marine Policy 36, 307-320.  Nyström, M., Folke, C., Moberg, F., 2000. Coral reef disturbance and resilience in a human- dominated environment. Trends in Ecology and Evolution 15, 413-417.  Oakley, S., Pilcher, N., Atack, K., Digges, C., Enderby, S., Mackey, G., Clubb, R., Stapelton, K., Toh, S.M., Huet, C., Morton, T., 1999. Reefs under attack: the status of coral reefs of Sabah, East Malaysia. Paper presented at the 4th international conference on the marine biology of the South China Sea 20-22 October 1999, Quezon City.  OMB, 1992. Guidelines and discount rates for benefit-cost analysis of federal programs. OMB Circular A-94, Washington, D.C.  Ono, R., 2010. Ethno-archaeology and early Austronesian fishing strategies in near-shore environments. Journal of Polynesian Society 119, 269-314.  Ostrom, E., 1990. Governing the commons: the evoloution of institutions for collective action. Cambridge University Press, New York.  Ostrom, E., Burger, J., Field, C., Norgaard, R., Policansky, D., 1999. Revisiting the commons: local lessons, global challenges. Science 284, 278-282.  Ovasisi, L., 2006. The live reef fish trade in Fiji. SPC Live Reef Fish Information Bulletin 16, 29-32.  Padilla, E., 2002. Intergenerational equity and sustainability. Ecological Economics 41, 69- 83.         119 Parma, A., Orensanz, J., Elías, I., Jerez, G., 2003. Diving for shellfish and data: incentives for the participation of fishermen in the monitoring and management of artisanal fisheries around southern South America, in: Newman, S., Gaughan, D., Jackson, G., Mackie, M., Molony, B., St. John, J., Kaiola, P. (Eds.), Towards sustainability of data limited multi-sector fisheries. Australian Society for Fish Biology, Department of Fisheries, Perth, Western Australia, pp. 8-29.  Pauly, D., 1980. On the interrelationships between natural mortality, growth parameters, and mean environmental temperature in 175 fish stocks. Journal du Conseil international pour l'Exploration de la Mer 39, 175-192.  Pauly, D., Silvestre, G., Smith, I.R., 1989. On development, fisheries, and dynamite: a brief review of tropical fisheries management. Natural Resource Modelling 3, 307-329.  Pauly, D., 1997. Small-scale fisheries in the tropics: marginality, marginalization, and some implications for fisheries management, in: Pikitch, E., Huppert, D., Sissenwine, M. (Eds.), Global Trends: Fisheries Management. American Fisheries Society, Bethesda, pp. 40-49.  Pauly, D., Christensen, V., Guenette, S., Pitcher, T., Sumaila, U., Walters, C., Watson, R., Zeller, D., 2002. Towards sustainability in world fisheries. Nature 418, 689-695.  Pauly, D., Alder, J., Bennett, E., Christensen, V., Tyedmers, P., Watson, R., 2003. The future for fisheries. Science 302, 1359-1361.  Pender, J., 1996. Discount rates and credit markets: theory and evidence from rural India. Journal of Development Economics 50, 257-296.  Pet-Soede, L., Erdmann, M., 1998. Blast fishing in southwest Sulawesi, Indonesia. NAGA, the ICLARM Quarterly April-June 1998, 4-9.  Pew Environment Group, 2011. Global Shark Conservation. The Pew Charitable Trusts, Washington, D.C. Available at Accessed 18 August 2011.     120 Picou-Gill, M., Woodley, J., Miller, M., Sary, Z., van Barneveld, W., Vatcher, S., Brown, M., 1991. Catch analysis at Discovery Bay, Jamaica: the status of an artisanal reef fishery. Presented at the 44th Annual meeting of the Gulf and Caribbean Fisheries Institute. Nassau, Bahamas, November 1991. Non peer-reviewed section. 706-713.  Pilcher, N., Cabanban, A.S., 2000. The status of coral reefs in eastern Malaysia. Global Coral Reef Monitoring Network (GCRMN) Report. Australian Institute of Marine Science, Townsville.  Pinca, S., Ribanataake, A., Kronen, M., Chapman, L., Lasi, F., Pakoa, K., Boblin, P., Friedman, K., Magron, F., Tardy, E., 2009a. Cook Islands country report: Profiles and results from survey work at Aitutaki, Palmerston, Mangaia and Rarotonga (February and October 2007). Pacific Regional Oceanic and Coastal Fisheries Development Programme (PROCFish/C/CoFish) / Secretariat of the Pacific Community, Noumea, 339 p.  Pinca, S., Tardy, E., Ribanataake, A., Kronen, M., Pakoa, K., Boblin, P., Friedman, K., Vunisea, A., Chapman, L., Lasi, F., Magron, F., 2009b. Marshall Islands country report: profiles and results from survey work at Likiep, Ailuk, Arno and Laura (August and September 2007). Pacific Regional Oceanic and Coastal Fisheries Development Programme (PROCFish/C/CoFish) / Secretariat of the Pacific Community, Noumea, 337 p.  Pinca, S., Vunisea, A., Lasi, F., Friedman, K., Kronen, M., Awira, R., Boblin, P., Tardy, E., Chapman, L., Magron, F., 2009c. Solomon Islands country report: profiles and results from survey work at Nggela, Marau, Rarumana and Chubikopi (June to September 2006 and December 2006). Pacific Regional Oceanic and Coastal Fisheries Development Programme (PROCFish/C/CoFish) / Secretariat of the Pacific Community, Noumea, 409 p.  Piper, M., 1984. Settlements on eight Semporna Islands. Sabah Society Journal 7, 303-329.  Pistorius, P., Taylor, F., 2009. Declining catch rates of reef fish in Aldabra's marine protected area. Aquatic Conservation: Marine and Freshwater Ecosystems 19, S2-S9.  Poggie, J.J., 1978. Deferred gratification as an adaptive characteristic for small-scale fishermen. Ethos 6, 114-123.    121  Pollnac, R., Poggie, J.J., 1978. Economic gratification orientations among small-scale fishermen in Panama and Puerto Rico. Human Organization 37, 355-367.  Pollnac, R.B., 1985. Social and Cultural Characteristics in Small-scale Fishery Development, in: Cernea, M.M. (Ed.), Putting People First Sociological Variables in Rural Development. Oxford University Press, Washington D.C., pp. 189-223.  Pomeroy, R.S., Mascia, M.B., Pollnac, R.B., 2007. Marine Protected Areas: The social dimension, FAO Expert Workshop on Marine Protected Areas and Fisheries Management: Review of Issues and Considerations. FAO, Rome, pp. 149-275.  Poulos, C., Whittington, D., 2000. Time preferences for life-saving programs: Evidence from six less developed countries. Environmental Science and Technology 34, 1445-1455.  Pretty, J., 2003. Social capital and the collective management of resources. Science 302, 1912-1914.  Rabl, A., 1996. Discounting of long-term costs: what would future generations prefer us to do? Ecological Economics 17, 137-145.  Ramsey, F.P., 1928. A mathematical theory of saving. Economic Journal 38, 543-559.  Reynolds, J., Dulvy, N., Goodwin, N., Hutchings, J., 2005. Biology of extinction risk in marine fishes. Proceedings of the Royal Society B 272, 2337-2344.  Rhodes, K., Tupper, M., 2007. A preliminary market-based analysis of the Pohnpei, Micronesia, grouper (Serranidae: Epinephelinae) fishery reveals unsustainable fishing practices. Coral Reefs 26, 335-344.  Robbins, W., Hisano, M., Connolly, S., Choat, J., 2006. Ongoing collapse of coral-reef shark populations. Current Biology 16, 2314-2319.     122 Roberts, C., 1995. Effects of fishing on the ecosystem structure of coral reefs. Conservation Biology 9, 988-995.  Rose, K., Cowan, J., Winemiller, K., Myers, R., Hilborn, R., 2001. Compensatory density dependence in fish populations: importance, controversy, understanding and prognosis. Fish and Fisheries 2, 293-327.  Rosenberg, A.A., 2003. Managing to the margins: the overexploitation of fisheries. Frontiers in Ecology and the Environment, 1, 102-106.  Rudd, M., 2004. An institutional framework for designing and monitoring ecosystem-based fisheries management policy experiments. Ecological Economics 48, 109-124.  Ruddle, K., Hickey, F., 2008. Accounting for the mismanagement of tropical nearshore fisheries. Environmental Development Sustained 10, 565-589.  Russ, G., Alcala, A., Maypa, A., Calumpong, H., White, A., 2004. Marine reserve benefits local fisheries. Ecological Applications 14, 597-606.  Saat, G., 2003. The identity and social mobility of Sama-Bajau. Sari 21, 3-11.  Sadiq, K., 2009. Paper citizens: how illegal immigrants acquire citizenship in developing countries. Oxford University Press, New York.  Sadovy, Y., Donaldson, T., Graham, T., McGilvray, F., Muldoon, G., Phillips, M., Rimmer, M., Smith, A., Yeeting, B., 2003a. While stocks last: the live reef food fish trade. Asian Development Bank, 143 p.  Sadovy, Y., Kulbicki, M., Labrosse, P., Letourneur, Y., Lokani, P., Donaldson, T., 2003b. The humphead wrasse, Cheilinus undulatus: synopsis of a threatened and poorly known giant coral reef fish. Reviews in Fish Biology and Fisheries 13, 327-364.  Sadovy, Y., 2005. Trouble on the reef: the imperative for managing vulnerable and valuable fisheries. Fish and Fisheries 6, 167-185.    123  Sadovy, Y., Punt, A., Cheung, W., Vasconcellos, M., Suharti, S., Mapstone, B., 2007. Stock assessment approach for the Napoleon Fish, Cheilinus Undulatus, in Indonesia. FAO Fisheries Circular No. 1023. FAO, Rome.  Sáeza, C., Requenab, J., 2007. Reconciling sustainability and discounting in cost–benefit analysis: A methodological proposal. Ecological Economics 60, 712 – 725.  Salas, S., Gaertner, D., 2004. The behavioural dynamics of fishers: management implications. Fish and Fisheries 5, 153-167.  Salas, S., Chuenpagdee, R., Seijo, J.C., Charles, A., 2007. Challenges in the assessment and management of small-scale fisheries in Latin America and the Caribbean. Fisheries Research, 87, 5-16.  Sale, P., 1991. The ecology of fishes on coral reefs. Academic Press, San Diego.  Sale, P., 2008. Management of coral reefs: Where we have gone wrong and what we can do about it. Marine Pollution Bulletin 56, 805-809.  Samuelson, D., 1937. A note on the measurement of utility. Review of Economic Studies 4, 155-161.  Sary, Z., Oxenford, H., Woodley, J., 1997. Effects of an increase in trap mesh size on an overexploited coral reef fishery at Discovery Bay, Jamaica. Marine Ecology Progress Series 154, 107-120.  Sary, Z., Munro, J., Woodley, J., 2003. Status report on a Jamaican reef fishery: current value and the costs of non-management. Proceedings of the 54th Gulf and Caribbean Fisheries Institute, 54, 98-111.  Sather, C., 1985. Boat crews and fishing fleets; the social organisation of maritime labour among the Bajau Laut of southeastern Sabah. Contributions to Southeast Asian Ethnography 4, 165-214.    124  Sather, C., 1997. The Bajau Laut. Adaptation, history, and fate in a maritime fishing society of south-eastern Sabah. Oxford University Press, Kuala Lumpur.  Sattar, S., 2008. Review of the reef fishery of Maldives. Marine Research Centre, Ministry of Fisheries, Agriculture and Marine Resources, Male, Republic of Maldives.  Sauni, S., Vunisea, A., Kronen, M., Friedman, K., Magron, F., Pakoa, K., Awira, R., Lasi, F., Chapman, L., Pinca, S., Vigliola, L., Boblin, P., Tardy, E., 2007. Nauru country report: Profile and results from in-country survey work (October and November 2005). Pacific Regional Oceanic and Coastal Fisheries Development Programme (PROCFish/C/CoFish) / Secretariat of the Pacific Community, Noumea, 137 p.  Sauni, S., Kronen, M., Pinca, S., Sauni, L., Friedman, K., Chapman, L., Magron, F., 2008. Tuvalu country report: profile and reports from in-country survey work (Oct-Nov 2004 and March-April 2005). SPC Coastal Fisheries Programme PROCFish/C/CoFish. Secretariat of the Pacific Community, Noumea, 311 p.  Scales, H., Balmford, A., Manica, A., 2007. Impacts of the live reef fish trade on populations of coral reef fish off northern Borneo. Proceedings of the Royal Society B 274, 989-994.  Sekhar, N.U., 2007. Social capital and fisheries management: the case of Chilika Lake in India. Environmental Management 39, 497-505.  Selliah, N., Oxenford, H., Parker, C., 2001. Selecting biodegradable fasteners and testing the effects of escape panels on catch rates of fish traps. Proceedings of the 52nd Gulf and Caribbean Fisheries Institute, 52, 634-653.  Shiferaw, B., Holden, S., 1998. Resource degradation and adoption of land conservation technologies in the Ethiopian Highlands: A case study in Andit Tid, North Shewa. Agricultural Economics 18, 233-247.      125 Smith, M., Roheim, C., Crowder, L., Halpern, B., Turnipseed, M., Anderson, J., Asche, F., Bourillon, L., Guttormsen, A., Khan, A., Liguori, L., McNevin, A., O'Connor, M., Squires, D., Tyedmers, P., Brownstein, C., Carden, K., Klinger, D., Sagarin, R., Selkoe, K., 2010. Sustainability and global seafood. Science 327, 784-786.  Soman, D., Ainslie, G., Frederick, S., Li, X., Luynch, J., Moreau, P., Mitchell, A., Read, D., Sawyer, A., Trope, Y., Wertenbroch, K., Zauberman, G., 2005. The psychology of intertemporal discounting: why are distant events valued differently from proximal ones? Marketing Letters 16, 347-360.  Sumaila, U.R., 2003. An economic indicator for monitoring fishing pressure on marine ecosystems. Ecofish report - Sustainability Indicators. Available at Accessed 20 May 2010.  Sumaila, U.R., 2004. Intergenerational cost benefit analysis and marine ecosystem restoration. Fish and Fisheries 5, 329-343.  Sumaila, U.R., 2005. Differences in economic perspectives and implementation of ecosystem-based management of marine resources. Marine Ecology Progress Series 300, 241-296.  Sumaila, U.R., Walters, C., 2005. Intergenerational discounting: a new intuitive approach. Ecological Economics 52, 135-142.  Sumaila, U.R., Marsden, D., Watson, R., Pauly, D., 2007. A global ex-vessel fish price database: construction and applications. Journal of Bioeconomics 9, 39-51.  Sumaila, R., Teh, L., Watson, R., Tyedmers, P., Pauly, D., 2008. Fuel price increase, subsidies, overcapacity and resource sustainability. ICES Journal of Marine Science, 65, 832- 840.  Sumaila, R., Khan, A., Dyck, A., Watson, R., Munro, G.R., Tyedmers, P., Pauly, D., 2010. A bottom-up re-estimation of global fisheries subsidies. Journal of Bioeconomics 12, 201-225.     126 Tanaka, T., Camerer, C.F., Nguyen, Q., 2010. Risk and time preferences: linking experimental and household survey data from Vietnam. American Economic Review 100, 557-571.  Teh, L., Sumaila, U.R., Cabanban, A.S., 2005. The reef fisheries of Pulau Banggi, Sabah: A preliminary profile and assessment of ecological and socio-economic sustainability. Fisheries Research 76, 359-367.  Teh, L., Sumaila, U.R., 2007. Malthusian overfishing in Pulau Banggi? Marine Policy 31, 451-457.  Teh, L.S.L., Zeller, D., Cabanban, A., Teh, L.C.L., Sumaila, U.R., 2007. Seasonality and historic trends in the reef fisheries of Pulau Banggi, Sabah, Malaysia. Coral Reefs 26, 251- 263.  Teh, L., Teh, L., Zeller, D., Cabanban, A., 2009a. Historical perspective of Sabah's marine fisheries, in: Zeller, D., Harper, S. (Eds.), Fisheries catch reconstructions: Islands, Part 1. University of British Columbia Fisheries Centre, Vancouver, pp. 77-98.  Teh, L.C.L., Teh, L.S.L., Starkhouse, B., Sumaila, U.R., 2009b. An overview of socio- economic and ecological perspectives of Fiji's inshore reef fisheries. Marine Policy 33, 807- 817.  Tietze, U., Thiele, W., Lasch, R., Thomsen, B., Rihan, D., 2005. Economic performance and fishing efficiency of marine capture fisheries. FAO Fisheries Technical Paper 482. FAO, Rome, 68 p.  Toller, W., Lundvall, S., 2008. Assessment of the commercial fishery of Saba Bank. Report for the Department of Environment of the Netherlands Antilles. Available at final_LR.pdf. Accessed 29 June 2011.      127 Torres, W.I., 2005. Voyages and ethnicity across reordered frontiers: conflict resolution and leadership in the dynamics of ethnic identity formation among the Sama Dilaut of Semporna, in: Economic Prospects, Cultural Encounters and Political Decisions: Reconstructing Scenes In A Moving Asia (East And Southeast) The Work of the 2002/2003 API Fellows. Sasyaz Holdings, Kuala Lumpur, pp. 19-38.  Turner, R., Cakacaka, A., Graham, N., Polunin, N., Pratchett, M., Stead, S., Wilson, S., 2007. Declining reliance on marine resources in remote South Pacific societies: ecological versus socio-economic drivers. Coral Reefs 26, 997-1008.  UNDP, 2007. Malaysia measuring and monitoring poverty and inequality. United Nations Development Programme, Kuala Lumpur.  UNDP, 2011. Human Development Report 2010. Palgrave Macmillan, New York. Available at Accessed 26 July 2011.  Unsworth, R., Cullen, L., 2010. Recognising the necessity for Indo-Pacific seagrass conservation. Conservation Letters 3, 63-73.  Veitayaki, J., 1998. Traditional and community based marine resources management system in Fiji: an evolving integrated process. Coastal Management 26, 47-60.  Vunisea, A., Friedman, K., Awira, R., Kronen, M., Pinca, S., Chapman, L., Magron, F., Sauni, S., Pakoa, K., Lasi, F., 2008. Samoa country report: profiles and results from survey work at Manono-Uta, Salelavalu, Vailoa and Vaisala (June 2005 and August/September 2005). Pacific Regional Oceanic and Coastal Fisheries Development Programme (PROCFish/C/CoFish) / Secretariat of the Pacific Community, Noumea, 349 p.  Warren-Rhodes, K., Sadovy, Y., Cesar, H., 2003. Marine ecosystem appropriation in the Indo-Pacific: A case study of the live reef fish food trade. Ambio 32, 481-488.  Weitzman, M., 2001. Gamma discounting. American Economic Review 91, 260-271.     128 Whittingham, E., Campbell, J., Townsley, P., 2003. Poverty and Reefs. DFID–IMM– IOC/UNESCO, Paris, 260 p.  Wilen, J., Smith, M., Lockwood, D., Botsford, L., 2002. Avoiding surprises: incorporating fishermen behaviour into management models. Bulletin of Fisheries Science 70, 553-575.  Wilson, S., Graham, N., Pratchett, M., Jones, G., Polunin, N., 2006. Multiple disturbances and the global degradation of coral reefs: Are reef fishes at risk or resilient? Global Change Biology 12, 2220-2234.  Winemiller, K., Rose, K., 1992. Patterns of life-history diversification in North American fishes: implications for population regulation. Canadian Journal of Fisheries and Aquatic Sciences 49, 2916-2218.  Wood, E., 2001. Use of natural resources in the proposed Semporna Islands park. Marine Conservation Society, Sabah Parks, WWF Malaysia, Nature Link.  Worm, B., Hilborn, R., Baum, J., Branch, T., Collie, J., Costello, C., Fogarty, M., Fulton, E., Huchings, J., Jennings, S., Jensen, O., Lotze, H., Mace, P., McClanahan, T., Minto, C., Palumbi, S., Parma, A., Ricard, D., Rosenberg, A., Watson, R., Zeller, D., 2009. Rebuilding global fisheries. Science 325, 578-584.  Wright, A., 1993. Shallow water reef-associated finfish, in: Wright, A., Hill, L. (Eds.), Nearshore marine resources of the South Pacific. Forum Fisheries Agency, Honiara and Institute of Pacific Studies, Suva, pp. 203-284.  WWF, 2003. Fiji Islands marine ecoregion. WWF Fiji Programme, Suva.  Yesuf, M., Bluffstone, R., 2008. Wealth and time preference in rural Ethiopia. Environment for Development Discussion Paper Series EfD DP 08-16, 19 p.  Zeller, D., Booth, S., Craig, P., Pauly, D., 2006a. Reconstruction of coral reef fisheries catches in American Samoa, 1950-2002. Coral Reefs 25, 144-152.     129 Zeller, D., Booth, S., Pauly, D., 2006b. Fisheries contributions to GDP: Underestimating small-scale fisheries in the Pacific. Marine Resource Economics 21, 355-374.  Zeller, D., Booth, S., Davis, G., Pauly, D., 2007. Re-estimation of small-scale fisheries catches for U.S. flag island areas in the Western Pacific: The last 50 years. Fishery Bulletin 105, 266-277.     130 Appendices Appendix A  Reconstructed and reported traditional catch (tonnes year-1) and reconstructed and reported number of traditional fishers for 1950-2009.1 Year Reconstructed no. of traditional fishers Reported number of traditional fishers2 Reconstructed traditional catch (tyr-1) Reported traditional landings2(t yr-1) 1950 9531 - 25034 - 1951 9008 - 24729 - 1952 9088 - 26028 - 1953 9262 - 27623 - 1954 9310 - 28872 - 1955 9377 - 30193 - 1956 9491 - 31687 - 1957 9604 - 33204 - 1958 9780 - 34971 - 1959 9916 - 36635 - 1960 10264 - 39141 - 1961 10328 - 40607 - 1962 10364 - 41980 - 1963 10373 - 43248 - 1964 10451 - 44815 - 1965 10518 - 46347 - 1966 9894 - 44774 - 1967 9775 - 45394 - 1968 9717 - 46278 - 1969 9701 - 47354 - 1970 9714 - 48569 - 1971 9842 - 49212 - 1972 9702 - 48509 - 1973 9883 - 49415 - 1974 10535 - 52673 - 1975 10772 - 53859 - 1976 11161 - 55806 - 1977 11491 - 57457 - 1978 11846 - 59230 - 1979 12047 - 60237 - 1980 12524 - 62619 - 1981 12242 - 60535 8387 1982 12946 - 63306 10563 1983 13627 - 65886 19892 1984 14390 - 68783 20311 1985 15177 - 71713 16616 1986 16012 - 74777 16419 1987 16906 - 78020 16221 1988 17753 4921 80953 21568 1989 18879 5035 85051 13491 1990 20092 4738 89411 12380 1991 22123 4752 97231 20378 1992 23786 5672 103231 44440 1993 25545 6351 109460 43566 1994 28228 7331 119405 42901 1995 29514 7341 123220 52795 1996 31129 7851 128250 52795 1997 32362 7851 131550 51719 1998 33581 7708 134661 49605 1999 36136 7708 142919 48784 2000 34205 7708 133400 46652 2001 35649 7708 137070 40914    131 Year Reconstructed no. of traditional fishers Reported number of traditional fishers2 Reconstructed traditional catch (tyr-1) Reported traditional landings2(t yr-1) 2002 36257 7708 137413 44802 2003 37847 7708 141360 41001 2004 39600 7708 145730 48920 2005 41824 7708 153911 41269 2006 43034 7708 158365 43378 2007 45041 9102 165752 41022 2008 46843 8040 172381 36172 2009 48833 8544 179705 36922 1 Source: Teh et al. (2009a). 2  Note that this differs from Teh et al. (2009a) due to the categorisation of gillnets as a commercial gear in Chapter 2.    132 Appendix B  Interview questionnaire for Sabah (English version)            ID____ 1. Personal/Demographic  Name:        Date: Age:        Gender: Education: Primary/Secondary/None   Ethnicity: Kampung:       Full-time/ Part-time: I. DEMOGRAPHIC INFORMATION 1. Where are you originally from? 2. How long have you lived in this village?____________ 3. Why did you move to this village? Fishing Other work Family & friends Other  4. Marital status: single___ married__ 5. How many people live in your house?  _______ Adult male Adult female Male children Female children  II. FISHING ACTIVITY 1. How many years have you been fishing? 2. Do you fish by yourself or with someone else? FISH ALONE WITH SOMEONE  1 other              2 other                 3 other                > 3 others  3. Do you own your own boat? YES:            Pumbot / sangkut                       HP:                   Length:                       Age: NO:              Owner is:                 Friend                       Family                      Pay rent? Y/N          RM_____per day/week/month  Fill in table for no. 4-11 Gear, catch, effort 4. What fishing gears do you use? 5. After listing gears: Please rank the gears according to which is most important    133 6. How many days per week do you use each gear? (low and high) 7. How much fish do you catch with xxx gear a day? We know that the amount of fish you catch is different each day, but say on a good day, how much fish would you catch? On a bad day? On an average day? 8. How many hours do you fish each day during bad, good, average day? ** specify which months? 9. How many times do you go out each day? 10. For each gear you use, what % of your catch consists of ikan batu, tengirri, tulai/rumahan/termanung, sunnoh, ikan putih etc. 11. Do you fish from your village all year? Y/N Fishing location Months Gear Village Other __________________    134    Gear  Rank  Days per week (Low) Days per week (High) Bad catch (kg/day)/ season  Good catch (kg/day)/ season Average catch (kg/day) / season No. Hrs Good/ no. times per day No. Hrs Bad/no. times per day No. Hrs Average/ no. times per day No. Ppl in household who use  gear Pancing          /        /        /        /        /        / Hantuk           /        /        /        /        /        / Tunda           /        /        /        /        /        / Pukat (specify type)           /        /        /        /        /        / Bubu           /        /        /        /        /        / Panar            /        /        /        /        /        / Cast net           /        /        /        /        /        / Prong (sotong)           /        /        /        /        /        / Gleaning           /        /        /        /        /        / Other           /        /        /        /        /        /   135 Catch composition (%) Gear Ikan batu Sunnoh (hidup) Putih Tengirri Termanung/tulai/ rumahan Other Pancing Hantuk Tunda Pukat Bubu Panar Cast net Prong (sotong) Gleaning Other   136 III. ECONOMIC  Fishing Income 1. How much of the fish that you catch per day is kept for food, and how much is sold? Kept (% or kg) Sold (% or kg)   2a. How many years have you been catching fish to sell? 2b. Before that did you fish for food, or not fish at all? 3a. Which species do you sell? 3b. What is your daily/weekly income from selling fish and other marine resources?  Income/day/ week/month (Good) Net? Which months? Income/day/ week/month (Bad) Net? Which months? Ikan mati Ikan hidup Sotong Balat / lepas  Other  4. What is your fishing cost per month for fishing gear, fuel, and boat maintenance? Item RM Month/week/day Gear     Fuel RM per litre/gallon/bottle:  Boat maintenance    137 Occupations (Occupational multiplicity) 1. How many people in your household work? 2. What jobs do you and other members of your household do to earn money or bring food for the family? Fill in Table ACTIVITY Check if applicable No. of People Rank of Importance Notes/Detail Fishing Gleaning Seaweed Grocery/coffee shop Farming Food stall Salaried Employment _________________  Tourism Informal Economic Activities  Remittance from family Other  3. Occupational mobility: What other work have you done in the last 5 years? Occupation Main job Why stop? Could get similar now? (y/n) Prefer to current activity? (y/n)      Household income and expenditure 1. What is total household income from all family members per month? (net?)   138 2. What is household expenditure per week/ per month? (if cannot answer, then what is your expenditure in the past 2 weeks?) Income Time (Week/month) Expenditure Week/month   3. Do you have access to credit? (If you have to borrow money to buy fishing gear, food, etc., who do you borrow from?) No (don’ t borrow) ____ Fish middleman ____      Store owner ____ Family/friends _____     Bank/financial institution ____ Co-operative____ 4a. If have access to credit: Do you repay your money with interest? Y/N 4b. How much interest do you have to pay? 5a. Do you have any savings? Y/N 5b. On average, how much (% or RM) of your monthly fishing income can you save?  IV. CAPACITY TO ADAPT  Capacity to anticipate change and develop response 1a. If you were to get 20% less catch all year, what would you do? keep fishing at same amount Fish harder Fish less move locations change gear leave fishery- where to? Other If keep fishing, for how long? 1b. if you were to get 50% less catch all year what would you do? keep fishing at same amount fish harder Fish less move locations change gear leave fishery- where to? Other If keep fishing, for how long?  Causality & intervention – perception of link between humans and marine environment 1. Is there more or less fish on the reef now compared to 5 years ago? 2. How do you know there are more/less fish now?   139 3. What do you think caused there to be more/less fish? 4. What can be done around here (kampung/Banggi) to increase the number of fish on the reef? Possible responses (Tick if any are mentioned):  a) restrict gear ______ b) stop fish bombing ______ c) ban commercial vessels ______  d) control migrants ______  V. SOCIAL DIMENSIONS  Reciprocity and exchanges 1. Do you share your fish with friends/family? How often? 1x/week    >1x/week    1x/month  <1x/month 2. Do you ever share your boat with another fisherman/take another fisherman out to fish with you? If YES, do you charge rent for your boat? Get fish in return?  Common rules, norms, and sanctions 1. Are there any rules/regulations about marine resource use in this area? 2. Are there fishing grounds that you are not allowed to use? 2a. If YES – do you continue to use those fishing grounds? Never Sometimes All the time  Do other people in this kampung continue to use the closed off fishing grounds? Never Sometimes All the time  2b. If NO - Pretend that there is a new law that says you cannot fish in your fishing grounds. What would you do? Go to another fishing ground Get another job Continue fishing (ignore law)  3. Are you involved in making decisions about marine resource use / management? 4. Who would you trust to manage your fishing grounds?     140 Government (Fisheries, Sabah Parks) Ketua kampung Other kampung people All fishers who fish here, including those from adjoining kampungs NGO e.g., WWF Why? Pretend you could make decisions about the fishing grounds: 5. Which are the fishing areas you would close off? To whom? 6. Which are the fishing grounds that are essential for you, that you want to leave open (while all other areas are closed)? 7. Are there groups of fishers you want to exclude from your fishing grounds? Who? Area to close off Area to leave open Who to exclude     8. Of all the people who fish in Banggi water, what do you think is their inclination to cooperate and follow new regulations?  Not          A little         Willing           Very willing     willing                                 willing 1                  2                   3                    4 Fishers from this kampung Fishers from other kampung Commercial fishers (trawl/ purse seine) Migrants   9. What priorities are important to you if a managed area were to be created?  Not                A little bit                           Important         Very important      important      Indifferent                             important 1                      2                    3                      4                   5 Maximise catch Maximise revenue Generate ecotourism Control commercial intrusion Conserve species    141 Stop fish bombing Other  Connectedness, networks and groups 1a. Is there a fishermen’ s organization in this kampung? Y/N 1b. If YES Are you a member? 1c. Have you participated in any meetings? 1d. How many of these meetings have you attended?       None__   Few__  Most__   All__ 2. Have you been involved in any non-family related community activities in the past year (e.g. gotong royong, pesta, tamu, sporting events)?  VI. DISCOUNT RATE ELICITATION QUESTIONS Exercise 1. Binary choice series Now we want to do an exercise with you. The purpose of this exercise is to see how you make decisions in the future. I am going to give you a series of choices for receiving money either today, or sometime in the future. To start off with, you can choose between receiving $100 now (show card) or  $105 in one month (show card). Which would you prefer?  Continue down list, marking choices.   142 SERIES 1 TODAY IN 1 MONTH 100 105 100 110 100 115 100 120 100 125 100 130 100 135 100 140 100 145 100 150 100 155 100 160 100 165 100 170  SERIES 2  TODAY IN 6 MONTHS 100 134 100 177 100 231 100 299 100 381 100 483 100 605 100 753 100 929 100 1139 100 1386 100 1677 100 2017 100 2413       143 SERIES 3  1 YEAR 1 YEAR + 1 MONTH 500 525 500 550 500 575 500 600 500 625 500 650 500 675 500 700 500 725 500 750 500 775 500 800 500 825 500 850    144  Appendix C  Interview questionnaire for Fiji  1. PERSONAL/DEMOGRAPHIC  Name:        Date: Age:        Gender: Education: Primary/Secondary/None   Indian/Fijian/Other: Village:       Full-time/ Part-time: 1. How many years have you been fishing? 2. Have you always lived in this village?  If NO, where did you come from? When did you move here? 3. How many people are there in your household? 4. Marital status: single__ married__ 4a. If married, how many children do you have?  2. FISHING ACTIVITY 1. What gear(s) do you use to fish? 2. How often do you use each gear? 3. What are the typical fish species you catch with each gear? 4. How many hours do you spend fishing per trip? 5. How much fish do you catch per trip? 6. Do you glean on the reef? a. How many people in your family glean? b. How often do you go gleaning in a week? c. What do you collect? d. Do you sell the collected products or keep it for your own consumption?  7. Do you fish for the purpose of (a) keeping the fish for own consumption  go to section 2I. (b) selling your catch and/or bartering  go to section 2II (c) both (a) and (b)  Complete sections 2I and 2II     145 2I. Subsistence fishing 1. On average how much fish do you keep for your own consumption? 2. Which species of fish do you keep for your own consumption? a. Has this changed over time? 3. On average how many days per week do you go fishing for food? 4. Has this fishing time increased or decreased from the past? 5. How many people do you feed with your daily catch? 6. Is the fish you catch enough to feed all the people in your household?  2II. Artisanal fishing 1. How long have you been engaged in fishing for the purpose of selling your catch (artisanal fishing)? 2. Compared to 5 years ago, has there been an increase or decrease in artisanal fishing in this village? 10 years ago? 3. What proportion of your catch do you sell? 4. Do you: (i) sell to a fish buyer/middleman  go to 5 (ii) sell the catch yourself?  go to 6 5a.Who do you sell to? 5b. How are the fish brought to market? 5c. What price do you get for each of the species you sell? Fill in table in #9. 6. Where do you bring the fish to sell? (i) municipal market  go to # 8 (ii) sell to other villagers door to door/informally. go to #7. (iii) other 7a. Why do you choose to sell the fish directly to villagers? (i) no fish buyer in this village (ii) fish buyer doesn’ t provide a good price (iii) municipal market is too far away 7b. What price do you get for the fish species you sell? Fill in table in #9 8a. How many times a week do you sell fish at the municipal market? 8b. How far away is the nearest municipal market that you sell your fish at? Do you rent a stall at the market? Y/N   146 If YES: How much is your rental? 8c. How many months per year do you sell fish at the market? 9. What price do you get for each of the species you sell? Species         Price (F$/kg)     10. How much do you earn from fishing per month? (clarify if it’ s revenue or net income) a. Does this income vary throughout the year? 10a. Is fishing your only source of income?    Y/N If NO:   a. What is your other job?   b. Do you work at your other job year round? 11. How much income do you get from that job ? 12. Is the income you earn from fishing enough to live on? Y/N 13. If there were other jobs available that offer the same income as fishing, would you want to switch jobs?  Y / N If NO, why? Enjoy fishing/want to be close to the sea   ____ No other skills      ____ Want to be own boss     ____ Other       ____ 14. Do other members of your household work? Y/N If YES a. How much does each household member contribute to household income?  Response to shifts in export market 1. Do you participate in any of these trades? Live reef food fish  ______    Beche de mer  ______   147 Aquarium fish  ______     Live rock ______ Trochus  ______     Fresh reef fish for sale  ______ 2b. When did you start participating in these trades? 3a. When you first started targeting these species, what were their prices? 3b. What are their prices now? 4. Who do you sell the live fish/beche de mer/live rock/trochus/ fresh fish to? 5. Do you know whether the species are sold domestically or exported? 6. How much income do you make from selling live fish/ beche de mer/ live rock/ trochus/ fresh reef fish trade? 7.  Has participating in the trade increased your daily/monthly fishing income? Y/N If YES By how much has it increased your daily/monthly fishing income? 8a. Do other people in this village participate in the LRFF/aquarium/beche de mer/live rock/trochus/fresh reef fish trade also? 8b. Have you noticed a change in the abundance of the target species since you started fishing for them? 8c. Has participation in the trade caused any changes to socio-economic or marine management aspects in the village? Y/N If YES, please describe the changes 9. Compared to 5 years ago, are you more likely to sell all your fish now rather than keeping a portion for your own consumption? 10 years ago? If MORE LIKELY  go to 10 If LESS LIKELY  go to 11 10. Why are you more likely to sell your fish now? • Able to sell fish for higher price now • Catch too little fish so must sell all that is caught • Fish alternatives are more easily available • Fish alternatives are cheaper to buy than before • Easier to get fish to market  11. Why are you less likely to sell your fish now?     148 3. COST QUANTIFICATION – quantify net income from harvesting species meant for export for each part of the supply chain. [Interviews with fishers as well as with wholesalers/exporters and middlemen]  Boat & Gear costs 1a. Do you own your own boat/canoe? 1b. How many years does a boat last? 1c. How much did your boat cost? 1d. How much did your engine cost (if applicable)? 1e. Where did the money come from? Your own, trader or government? 2a. How much fuel do you use for each fishing trip on average? 2b. How much do you pay for fuel per fishing trip 3a. How much time do you spend repairing/maintaining your boat and gear every month? 3b. On average, how much money do you spend on maintaining your boat and gear every month?  Costs associated with selling fish: 4a. What is the rental per month for your stall at the market? 4b. What is the transportation cost to bring your catch to the market? 4c. Do you hire family or workers to help you sell the fish or transport the fish? Y/N If YES, what wage do you pay per month for your helper?   149  Cost factor F$/unit # of units purchased per year Hook & line:  - Hooks  - Line  - Reel  - Weight  - Bait Gillnet:  - net  - floats Trap - wire - frame Spear and speargun Mask Snorkel Fin  4. DISCOUNT RATE ELICITATION EXERCISE Now we want to do an exercise with you. The purpose of this exercise is to see how you make decisions about money. I am going to give you a series of choices for receiving money either now, or sometime in the future. To start off with, you can choose between receiving $400 now (show card) or  $405 in one month (show card). Which would you prefer?  Continue down list, marking choices.   150 NOW ($) 1 MONTH ($) Selection 400 420 A      B 400 435 A      B 400 450 A      B 400 465 A      B 400 480 A      B 400 495 A      B 400  510 A      B 400 525 A      B 400 540 A      B 400 555 A      B 400 570 A      B 400 585 A      B 400 600 A      B  Now the choice is between receiving $400 now, and $550 in 1 year. Which would you prefer? (Go down the list) NOW ($) 1 YEAR ($) Selection 400 550 A      B 400 600 A      B 400 650 A      B 400 700 A      B 400 750 A      B 400  800 A      B 400 850 A      B 400 900 A      B 400 950 A      B 400 1000 A      B 400 1050 A      B 400 1100 A      B 400 1150 A      B        151 Appendix D  Immediate and future payment amounts with associated discount rates for each choice series.  Appendix D1. Immediate and future payment amounts (RM) and associated discount rate (annual %) for each option in Series As  (Sabah). Payment today (RM) Future payment Delay = 1 month (RM)1 Implied mid-point discount rate    (annual %) 100 105   29 100 110   86 100 115 141 100 120 193 100 125 243 100 130 291 100 135 337 100 140 381 100 145 424 100 150 466 100 155 506 100 160 545 100 165 583 100 170 619 1  The exchange rate on 27 Oct. 2010 is 1 USD = 3.1 RM                 152 Appendix D2. Immediate and future payment amounts (RM) and associated discount rate (annual %) for each option in Series Bs (Sabah). Payment today (RM) Future payment Delay = 6 months (RM) Implied mid-point discount rate    (annual %) 100 134    29 100 177    86 100 231 141 100 299 193 100 381 243 100 483 291 100 605 337 100 753 381 100 929 424 100 1139 466 100 1386 506 100 1677 545 100 2017 583 100 2413 619  Appendix D3. Payment amounts (RM) for delays of 1 year and 1 year + 1 month, and associated discount rate (annual %) for each option in Series Cs (Sabah). Payment in 1 year (RM) Payment in 1 year + 1 month (RM) Implied mid-point discount rate    (annual %) 500 525    29 500 550    86 500 575 141 500 600 193 500 625 243 500 650 291 500 675 337 500 700 381 500 725 424 500 750 466 500 775 506 500 800 545 500 825 583 500 850 619   153  Appendix D4. Immediate and future payment amounts (FJD) and associated discount rate (annual %) for each option in Series AF  (Fiji). Payment today (FJD) Future payment Delay = 1 month (FJD)1 Implied mid-point discount rate    (annual %) 400 420     29 400 435   80 400 450 121 400 465 161 400 480 200 400 495 237 400 510 274 400 525 309 400 540 343 400 555 377 400 570 409 400 585 441 400 600 471 1 The exchange rate on 27 Oct. 2010 is 1 USD = 1.9 FJD  Appendix D5. Current and future payment amounts (FJD) and associated discount rate (annual %) for each option in Series BF  (Fiji). Payment today (FJD) Future payment Delay = 1 year (FJD) Implied mid-point discount rate    (annual %) 400    550     27 400    600    36 400    650    45 400    700    52 400    750    60 400    800    66 400    850    72 400    900    78 400    950    84 400 1000    89 400 1050    94 400 1100    99 400 1150   103    154 Appendix E  Estimated intrinsic growth rates (r) for reef fish species Family Taxon name r Family Taxon name r Acanthuridae    Pseudobalistes flavimarginatus 1.26  Acanthurus achilles 1.54  Rhinecanthus aculeatus 1.22  Acanthurus bahianus 0.84  Rhinecanthus rectangulus 0.65  Acanthurus blochii 0.36  Rhinecanthus verrucosus 1.32  Acanthurus chirurgus 0.54  Sufflamen fraenatum 0.45  Acanthurus coeruleus 0.23  Xanthichthys ringens 1.51  Acanthurus gahhm 0.75 Belonidae  Acanthurus guttatus 1.51  Strongylura incisa 0.59  Acanthurus leucocheilus 0.52  Strongylura leiura 0.67  Acanthurus lineatus 1.00  Tylosurus acus melanotus 0.68  Acanthurus mata 0.29  Tylosurus crocodilus crocodilus 0.43  Acanthurus nigricauda 1.60 Blenniidae  Acanthurus nigrofuscus 0.90  Omobranchus elongatus 2.99  Acanthurus nigroris 1.53 Bothidae  Acanthurus nubilus 1.51  Bothus lunatus 0.27  Acanthurus olivaceus 0.78 Bramidae  Acanthurus pyroferus 0.87  Brama orcini 0.52  Acanthurus tennentii 1.45 Caesionidae  Acanthurus triostegus 2.02  Caesio caerulaurea 1.51  Acanthurus xanthopterus 0.36  Caesio cuning 0.54  Ctenochaetus binotatus 1.59  Caesio lunaris 1.44  Ctenochaetus striatus 3.18  Caesio teres 1.44  Ctenochaetus strigosus 3.20  Caesio xanthonota 1.44  Melichthys niger 0.48  Pterocaesio chrysozona 3.12  Naso annulatus 0.27  Pterocaesio marri 1.51  Naso brachycentron 0.13  Pterocaesio pisang 3.37  Naso brevirostris 0.81  Pterocaesio tesellata 2.97  Naso caesius 0.50  Pterocaesio tile 1.58  Naso fageni 0.23 Callionymidae  Naso hexacanthus 0.28  Synchiropus  corallinus 2.23  Naso lituratus 0.78 Carangidae  Naso minor 0.78  Alectis ciliaris 0.17  Naso tuberosus 0.82  Alectis indicus 0.24  Naso unicornis 0.20  Atule mate 1.79  Naso vlamingii 0.36  Carangoides bartholomaei 0.59  Zebrasoma scopas 0.20  Carangoides coeruleopinnatus 1.43 Albulidae    Carangoides ferdau 0.46  Albula forsteri 0.73  Carangoides fulvoguttatus 0.22  Albula neoguinaica 0.69  Carangoides gymnostethus 0.45  Albula vulpes 0.49  Carangoides hedlandensis 1.53 Atherinidae    Carangoides orthogrammus 0.49  Atherinomorus lacunosus 0.35  Carangoides plagiotaenia 0.75 Aulostomidae    Carangoides ruber 0.30  Aulostomus maculatus 0.25  Caranx crysos 1.36 Balistidae    Caranx hippos 0.33  Abalistes stellaris 0.37  Caranx ignobilis 0.22  Abalistes stellatus 0.36  Caranx latus 0.35  Balistapus undulatus 0.63  Caranx lugubris 0.17  Balistes capriscus 1.33  Caranx melampygus 0.46  Balistes vetula 0.44  Caranx papuensis 0.86  Balistoides conspicillum 0.40  Caranx sexfasciatus 0.49  Balistoides viridescens 0.28  Chloroscombrus chrysurus 1.54  Canthidermis sufflamen 0.30  Decapterus macarellus 2.96  Odonus niger 0.40  Decapterus macrosoma 1.59      155 Family Taxon name r Family Taxon name r Carangidae Elagatis bipinnulata 0.91 Congridae contd. Gnathanodon speciosus 0.60  Conger cinereus 0.17  Megalaspis cordyla 0.74 Coryphaenidae  Scomberoides lysan 0.72  Coryphaena hippurus 3.24  Scomberoides tala 0.50 Dactylopteridae  Selar boops 3.58  Dactylopterus volitans 1.74  Selar crumenophthalmus 3.10 Dasyatidae  Selaroides leptolepis 3.38  Dasyatis akajei 0.04  Seriola dumerili 0.46  Dasyatis americana 0.04  Seriola rivoliana 0.21  Taeniura lymma 0.03  Trachinotus blochii 0.36 Diodontidae  Ulua aurochs 0.75  Chilomycterus antennatus 1.39  Ulua mentalis 0.37  Chilomycterus antillarum 1.45 Carcharhinidae    Diodon holocanthus 0.40  Carcharhinus amblyrhynchos 0.30  Diodon hystrix 0.29  Carcharhinus limbatus 0.10 Echeneidae  Carcharhinus melanopterus 0.06  Echeneis naucrates 0.24  Carcharhinus perezii 0.04  Remora remora 0.46  Galeocerdo cuvier 0.07 Eleotridae Centrarchidae    Eleotris fusca 0.43  Acantharchus pomotis 0.99 Elopidae Centropomidae    Elops saurus 0.66  Centropomus pectinatus 1.50 Engraulidae  Centropomus undecimalis 0.52  Stolephorus commersonni 2.06 Chaetodontidae    Stolephorus indicus 2.04  Chaetodon auriga 3.58 Ephippidae  Chaetodon capistratus 3.58  Chaetodipterus faber 0.72  Chaetodon fasciatus 3.20  Platax orbicularis 0.56  Chaetodon guttatissimus 3.58 Exocoetidae  Chaetodon ocellatus 3.32  Cheilopogon abei 1.94  Chaetodon sedentarius 2.06  Cheilopogon unicolor 1.91  Chaetodon striatus 3.42  Exocoetus volitans 3.21  Heniochus monoceros 3.10 Fistulariidae  Prognathodes aculeatus 3.58  Fistularia commersonii 0.19 Chanidae   Gempylidae  Chanos chanos 0.36  Promethichthys prometheus 0.34 Chimaeridae    Ruvettus pretiosus 0.11  Chimaera monstrosa 0.24 Gerreidae Chirocentridae    Eugerres brasilianus 0.56  Chirocentrus nudus 0.43  Eugerres plumieri 0.57 Cirrhitidae    Gerres acinaces 0.50  Cirrhitus pinnulatus 1.46  Gerres argyreus 1.60  Paracirrhites hemistictus 1.56  Gerres cinereus 0.97 Clupeidae    Gerres erythrourus 1.43  Amblygaster clupeoides 0.92  Gerres filamentosus 3.02  Amblygaster sirm 2.03  Gerres longirostris 0.50  Dussumieria elopsoides 0.94  Gerres oblongus 1.45  Etrumeus teres 1.63  Gerres oyena 1.43  Herklotsichthys quadrimaculatus 2.23 Ginglymostomatidae  Herklotsichthys spilurus 2.12  Ginglymostoma cirratum 0.05  Opisthonema oglinum 0.96 Gobiidae  Sardinella gibbosa 2.23  Acentrogobius  puntang 0.50  Spratelloides delicatulus 2.23 Haemulidae  Spratelloides gracilis 2.23  Anisotremus virginicus 0.49   156 Family Taxon name r Family Taxon name r Haemulidae Diagramma picta 0.47  Sargocentron diadema 3.40 contd. Haemulon album 0.38  Sargocentron praslin 1.53  Haemulon aurolineatum 0.40  Sargocentron rubrum 1.53  Haemulon carbonarium 0.52  Sargocentron spiniferum 0.78  Haemulon chrysargyreum 1.42  Sargocentron tiere 1.53  Haemulon flavolineatum 0.60  Sargocentron vexillarium 3.30  Haemulon melanurum 0.58  Sargocentron violaceum 1.40  Haemulon parra 0.52 Istiophoridae  Haemulon plumierii 0.24  Istiophorus platypterus 0.40  Haemulon sciurus 0.56 Kuhlidae  Haemulon striatum 0.75  Kuhlia marginata 0.59  Haemulopsis leuciscus 0.59  Kuhlia mugil 0.23  Plectorhinchus albovittatus 0.14 Kyphosidae  Plectorhinchus celebicus 0.40  Kyphosus bigibbus 0.19  Plectorhinchus ceylonensis 0.50  Kyphosus cinerascens 0.52  Plectorhinchus chaetodonoides 0.23  Kyphosus saltatrix 0.15  Plectorhinchus chrysotaenia 0.43  Kyphosus vaigiensis 0.20  Plectorhinchus diagrammus 0.53 Labridae  Plectorhinchus flavomaculatus 0.37  Anampses caeruleopunctatus 0.38  Plectorhinchus gibbosus 0.38  Anampses neoguinaicus 0.57  Plectorhinchus lessonii 0.53  Bodianus bilunulatus 0.20  Plectorhinchus lineatus 0.55  Bodianus diana 0.42  Plectorhinchus obscurus 0.15  Bodianus perditio 0.12  Plectorhinchus orientalis 0.19  Bodianus rufus 0.39  Plectorhinchus pictus 0.24  Cheilinus chlorourus 0.37  Plectorhinchus picus 0.37  Cheilinus fasciatus 0.22  Plectorhinchus vittatus 0.17  Cheilinus trilobatus 0.38 Hemiramphidae    Cheilinus undulatus 0.14  Hemiramphus archipelagicus 1.61  Cheilio inermis 0.17  Hemiramphus far 0.86  Choerodon anchorago 0.32  Hyporhamphus dussumierii 0.84  Choerodon schoenleinii 0.12  Zenarchopterus dispar 2.07  Cirrhilabrus exquisitus 1.53 Holocentridae    Cirrhilabrus walindi 3.39  Holocentrus adscensionis 0.55  Clepticus parrae 0.54  Holocentrus ascensionis 0.55  Epibulus insidiator 0.17  Holocentrus rufus 3.18  Halichoeres bivittatus 0.41  Myripristis adusta 1.52  Halichoeres chloropterus 0.57  Myripristis amaena 1.54  Halichoeres garnoti 0.84  Myripristis berndti 1.59  Halichoeres hortulanus 1.67  Myripristis botche 1.51  Halichoeres scapularis 1.61  Myripristis chryseres 2.99  Hemigymnus fasciatus 0.16  Myripristis jacobus 2.90  Iniistius pavo 0.38  Myripristis kuntee 1.62  Lachnolaimus maximus 0.14  Myripristis murdjan 0.72  Novaculichthys taeniourus 0.54  Myripristis pralinia 3.16  Oxycheilinus celebicus 0.59  Myripristis violacea 1.52  Oxycheilinus unifasciatus 0.37  Myripristis vittata 2.99  Thalassoma hebraicum 0.59  Neoniphon opercularis 1.52  Thalassoma lunare 0.58  Neoniphon sammara 1.55  Thalassoma purpureum 0.37  Ostichthys kaianus 0.56  Thalassoma quinquevittatum 1.49  Plectrypops lima 3.37  Xyrichtys martinicensis 1.49  Sargocentron caudimaculatum 2.99 Leiognathidae  Sargocentron cornutum 1.61  Gazza minuta 2.08  Sargocentron coruscum 3.40  Leiognathus equulus 2.03   157 Family Taxon name r Family Taxon name r Lethrinidae    Lutjanus fulvus 1.63  Gnathodentex aureolineatus 0.82  Lutjanus gibbus 0.81  Gymnocranius audleyi 0.92  Lutjanus goldiei 0.22  Gymnocranius elongatus 0.76  Lutjanus griseus 0.50  Gymnocranius euanus 0.40  Lutjanus jocu 0.14  Gymnocranius grandoculis 0.48  Lutjanus kasmira 0.40  Gymnocranius griseus 0.42  Lutjanus mahogoni 0.14  Gymnocranius microdon 0.53  Lutjanus monostigma 0.41  Lethrinus  rubrioperculatus 0.41  Lutjanus notatus 1.63  Lethrinus amboinensis 0.36  Lutjanus purpureus 0.18  Lethrinus atkinsoni 0.54  Lutjanus quinquelineatus 0.40  Lethrinus borbonicus 0.56  Lutjanus rivulatus 0.47  Lethrinus conchyliatus 0.35  Lutjanus russellii 0.55  Lethrinus crocineus 0.39  Lutjanus sebae 0.28  Lethrinus erythracanthus 0.41  Lutjanus semicinctus 0.82  Lethrinus erythropterus 0.51  Lutjanus synagris 0.40  Lethrinus genivittatus 1.84  Lutjanus vitta 0.41  Lethrinus harak 0.83  Lutjanus vivanus 0.14  Lethrinus lentjan 0.57  Macolor macularis 0.50  Lethrinus mahsena 0.21  Macolor niger 0.45  Lethrinus microdon 0.38  Ocyurus chrysurus 0.18  Lethrinus miniatus 0.35  Pristipomoides filamentosus 0.50  Lethrinus nebulosus 0.26  Pristipomoides sieboldii 0.70  Lethrinus obsoletus 0.55  Pristipomoides typus 0.51  Lethrinus olivaceus 0.34  Pristipomoides zonatus 0.47  Lethrinus ornatus 1.50  Rhomboplites aurorubens 0.36  Lethrinus rubrioperculatus 0.41  Symphorichthys spilurus 0.50  Lethrinus semicinctus 1.55  Symphorus nematophorus 0.46  Lethrinus variegatus 0.68 Malacanthidae  Lethrinus xanthochilus 0.20  Malacanthus plumieri 0.19  Monotaxis grandoculis 0.39 Monacanthidae Lutjanidae    Aluterus monoceros 0.05  Aphareus furca 0.46  Aluterus schoepfii 0.06  Aphareus rutilans 0.32  Aluterus scriptus 0.12  Aprion virescens 0.28  Cantherhines macrocerus 0.29  Apsilus dentatus 0.92  Cantherhines pullus 0.61  Apsilus fuscus 0.42  Monacanthus chinensis 0.32  Etelis carbunculus 0.73 Monodactylidae  Etelis coruscans 0.16  Monodactylus argenteus 1.45  Etelis oculatus 1.10 Mugilidae  Lipocheilus carnolabrum 0.52  Crenimugil crenilabis 0.52  Lutjanus adetii 0.51  Liza melinoptera 0.51  Lutjanus analis 0.26  Liza parsia 1.60  Lutjanus apodus 0.59  Liza tade 0.38  Lutjanus argentimaculatus 0.36  Liza vaigiensis 0.21  Lutjanus bengalensis 1.50  Mugil cephalus 0.50  Lutjanus bohar 0.52  Neomyxus chaptalii 0.50  Lutjanus boutton 0.83  Valamugil seheli 0.48  Lutjanus buccanella 0.24 Mullidae  Lutjanus campechanus 0.23  Mulloidichthys flavolineatus 0.42  Lutjanus carponotatus 0.79  Mulloidichthys martinicus 0.76  Lutjanus decussatus 0.83  Mulloidichthys vanicolensis 0.97  Lutjanus ehrenbergii 0.82  Parupeneus barberinoides 1.47  Lutjanus fulviflamma 0.56  Parupeneus barberinus 0.43   158 Family Taxon name r Family Taxon name r Mullidae Parupeneus ciliatus 0.76 Pleuronectidae contd. Parupeneus cyclostomus 0.53  Nematops microstoma 1.71  Parupeneus heptacanthus 0.68 Plotosidae  Parupeneus indicus 0.55  Plotosus lineatus 0.63  Parupeneus macronemus 0.74 Polymixiidae  Parupeneus multifasciatus 0.79  Polymixia japonica 0.23  Parupeneus pleurostigma 1.42 Polynemidae  Parupeneus rubescens 0.74  Polydactylus sexfilis 0.75  Parupeneus trifasciatus 0.78 Pomacanthidae  Parupenues macronemus 0.76  Holacanthus ciliaris 0.38  Pseudupeneus maculatus 1.62  Holacanthus isabelita 0.38  Upeneus taeniopterus 1.51  Holacanthus tricolor 0.54  Upeneus vittatus 1.61  Pomacanthus arcuatus 0.24 Muraenidae    Pomacanthus paru 0.69  Echidna nebulosa 0.37  Pomacanthus sexstriatus 0.40  Echidna polyzona 0.50  Pygoplites diacanthus 0.57  Gymnothorax eurostus 0.37 Pomacentridae  Gymnothorax fimbriatus 0.40  Abudefduf saxatilis 0.44  Gymnothorax flavimarginatus 0.21  Abudefduf septemfasciatus 0.46  Gymnothorax funebris 0.15  Abudefduf sexfasciatus 1.29  Gymnothorax meleagris 0.35  Abudefduf sordidus 0.46  Gymnothorax miliaris 0.32  Abudefduf vaigiensis 2.76  Gymnothorax moringa 0.11  Amphiprion clarkii 0.70  Gymnothorax pictus 0.24  Chromis multilineata 0.47  Gymnothorax vicinus 0.16  Microspathodon chrysurus 0.45 Nemipteridae    Neoglyphidodon melas 0.86  Pentapodus emeryii 1.50  Pomacentrus reidi 1.48  Pentapodus paradiseus 1.50  Stegastes adustus 1.53  Pentapodus trivittatus 1.59  Stegastes leucostictus 1.67  Scolopsis ghanam 1.50  Stegastes planifrons 1.63  Scolopsis lineata 3.43 Priacanthidae  Scolopsis margaritifera 1.53  Heteropriacanthus cruentatus 1.42  Scolopsis monogramma 1.54  Priacanthus arenatus 1.56  Scolopsis temporalis 0.86  Priacanthus blochii 2.81  Scolopsis trilineata 1.68  Priacanthus hamrur 1.59 Ostraciidae   Rajidae  Acanthostracion polygonius 0.39  Dipturus canutus 0.06  Acanthostracion quadricornis 0.67 Rhinobatidae  Lactophrys bicaudalis 0.40  Glaucostegus typus 0.04  Lactophrys trigonus 0.38  Rhynchobatus djiddensis 0.04  Ostracion cubicus 0.98 Scaridae  Ostracion meleagris 0.91  Bolbometopon muricatum 0.14  Rhinesomus triqueter 0.40  Calotomus carolinus 0.50 Paralichthyidae    Calotomus japonicus 1.36  Paralichthys tropicus 0.73  Cetoscarus bicolor 0.27  Pseudorhombus arsius 0.32  Chlorurus bleekeri 0.53 Pempheridae    Chlorurus frontalis 0.52  Pempheris vanicolensis 1.51  Chlorurus japanensis 1.53 Pinguipedidae    Chlorurus microrhinos 0.28  Parapercis clathrata 1.44  Chlorurus sordidus 1.95 Platycephalidae    Chlorurus strongylocephalus 0.50  Cymbacephalus beauforti 0.39  Hipposcarus longiceps 1.50  Papilloculiceps longiceps 0.38  Leptoscarus vaigiensis 1.39  Platycephalus indicus 0.75  Scarus altipinnis 0.52   159 Family Taxon name r Family Taxon name r Scaridae Scarus coeruleus 0.55  Cephalopholis cyanostigma 0.52 contd. Scarus dimidiatus 1.30  Cephalopholis fulva 0.67  Scarus forsteni 0.50  Cephalopholis igarashiensis 0.44  Scarus frenatus 1.02  Cephalopholis miniata 0.15  Scarus ghobban 0.44  Cephalopholis polleni 0.50  Scarus globiceps 1.50  Cephalopholis sexmaculata 0.40  Scarus guacamaia 0.53  Cephalopholis sonnerati 0.36  Scarus iseri 1.43  Cephalopholis spiloparaea 0.58  Scarus niger 1.66  Cephalopholis urodeta 0.59  Scarus oviceps 1.37  Cromileptes altivelis 0.20  Scarus psittacus 1.62  Epinephelus adscensionis 0.21  Scarus rivulatus 0.77  Epinephelus areolatus 0.55  Scarus rubroviolaceus 0.36  Epinephelus bilobatus 0.56  Scarus schlegeli 0.42  Epinephelus chlorostigma 0.56  Scarus spinus 1.45  Epinephelus coioides 0.34  Scarus taeniopterus 1.37  Epinephelus corallicola 0.40  Scarus tricolor 1.53  Epinephelus cyanopodus 0.13  Scarus vetula 1.56  Epinephelus fasciatus 0.23  Scarus viride 1.62  Epinephelus flavocaeruleus 0.16  Sparisoma aurofrenatum 0.41  Epinephelus flavolimbatus 0.13  Sparisoma chrysopterum 2.98  Epinephelus fuscoguttatus 0.38  Sparisoma rubripinne 1.56  Epinephelus guttatus 0.25  Sparisoma viride 0.85  Epinephelus hexagonatus 3.16 Sciaenidae    Epinephelus howlandi 0.38  Argyrosomus hololepidotus 0.13  Epinephelus itajara 0.17  Argyrosomus thorpei 0.46  Epinephelus lanceolatus 0.06  Equetus lanceolatus 1.62  Epinephelus macrospilos 0.39  Equetus punctatus 1.59  Epinephelus maculatus 0.53 Scombridae    Epinephelus malabaricus 0.07  Acanthocybium solandri 0.65  Epinephelus melanostigma 0.55  Euthynnus affinis 1.35  Epinephelus merra 1.61  Euthynnus alletteratus 0.34  Epinephelus miliaris 0.39  Grammatorcynus bilineatus 0.51  Epinephelus morio 0.17  Gymnosarda unicolor 0.15  Epinephelus morrhua 0.16  Katsuwonus pelamis 0.79  Epinephelus multinotatus 0.51  Rastrelliger brachysoma 2.23  Epinephelus niveatus 0.13  Rastrelliger kanagurta 3.58  Epinephelus ongus 0.58  Sarda orientalis 0.52  Epinephelus polyphekadion 0.13  Scomberomorus commerson 0.72  Epinephelus quoyanus 0.52  Thunnus albacares 0.67  Epinephelus retouti 0.40  Thunnus atlanticus 0.69  Epinephelus rivulatus 1.49  Thunnus obesus 0.44  Epinephelus septemfasciatus 0.10  Thunnus orientalis 0.13  Epinephelus socialis 0.39 Scorpaenidae    Epinephelus spilotoceps 0.55  Scorpaena plumieri 0.14  Epinephelus striatus 0.17  Scorpaenodes caribbaeus 0.62  Epinephelus tauvina 0.18 Serranidae    Epinephelus tukula 0.09  Aethaloperca rogaa 0.37  Gracila albomarginata 0.52  Alphestes afer 0.94  Hypoplectrus chlorurus 1.70  Anyperodon leucogrammicus 0.21  Hyporthodus mystacinus 0.09  Cephalopholis argus 0.37  Hyporthodus nigritus 0.18  Cephalopholis aurantia 0.37  Mycteroperca bonaci 0.22  Cephalopholis boenak 0.80  Mycteroperca interstitialis 0.09  Cephalopholis cruentata 0.59  Mycteroperca microlepis 0.17   160 Family Taxon name r Family Taxon name r Serranidae Mycteroperca tigris 0.16 Sphyrnidae  Mycteroperca venenosa 0.24  Sphyrna lewini 0.03  Paranthias furcifer 0.41  Sphyrna zygaena 0.03  Plectropomus areolatus 0.20 Synodontidae  Plectropomus laevis 0.13  Synodus intermedius 1.41  Plectropomus leopardus 0.48  Synodus jaculum 3.32  Plectropomus maculatus 0.39 Terapontidae  Plectropomus oligacanthus 0.19  Pelates quadrilineatus 0.44  Plectropomus pessuliferus 0.13  Terapon jarbua 0.33  Plectropomus punctatus 0.16 Tetraodontidae  Rypticus saponaceus 0.55  Arothron hispidus 0.40  Saloptia powelli 0.50  Arothron stellatus 0.05  Serranus tabacarius 1.42  Canthigaster rostrata 3.29  Variola albimarginata 0.21  Diodon liturosus 0.52  Variola louti 0.37 Torpedinidae Siganidae    Torpedo sinuspersici 0.02  Siganus argenteus 2.99 Triacanthidae  Siganus canaliculatus 3.58  Triacanthus biaculeatus 0.30  Siganus doliatus 1.59 Triakidae  Siganus fuscescens 3.15  Mustelus canis 0.05  Siganus guttatus 3.58 Trichiuridae  Siganus lineatus 1.59  Trichiurus lepturus 0.41  Siganus luridus 0.56 Xiphiidae  Siganus puellus 1.50  Xiphias gladius 0.51  Siganus punctatus 1.49 Zanclidae  Siganus randalli 1.61  Zanclus cornutus 1.68  Siganus rivulatus 0.99  Siganus spinus 3.58  Siganus sutor 3.09  Siganus vermiculatus 2.23  Siganus vulpinus 1.63 Sillaginidae  Sillago ciliata 0.93 Sparidae  Acanthopagrus bifasciatus 0.40  Calamus bajonado 0.18  Calamus calamus 0.61  Calamus penna 0.38  Calamus pennatula 0.41  Calamus proridens 0.39  Cheimerius nufar 0.11  Crenidens crenidens 1.10  Diplodus bermudensis 0.54  Rhabdosargus sarba 0.50 Sphyraenidae  Sphyraena acutipinnis 0.44  Sphyraena barracuda 0.12  Sphyraena flavicauda 0.55  Sphyraena forsteri 0.38  Sphyraena jello 0.15  Sphyraena novaehollandiae 0.19  Sphyraena picudilla 0.46  Sphyraena putnamae 0.73  Sphyraena qenie 0.12   161 Appendix F  Top 5 families making up the catch composition for each fishery case Country Case No. Family % catch  Country Case No. Family % catch Anguilla 2 Lutjanidae 0.38  Marshall Islands  64 Serranidae 0.40 Anguilla 2 Haemulidae 0.11  Marshall Islands  64 Lutjanidae 0.23 Anguilla 2 Acanthuridae 0.10  Marshall Islands  64 Siganidae 0.14 Anguilla 2 Sparidae 0.09  Marshall Islands  64 Lethrinidae 0.11 Anguilla 2 Serranidae 0.07  Marshall Islands  64 Acanthuridae 0.05 Antigua and Barbuda 3 Serranidae 0.22  Marshall Islands  65 Acanthuridae 0.20 Antigua and Barbuda 3 Lutjanidae 0.21  Marshall Islands  65 Lutjanidae 0.18 Antigua and Barbuda 3 Haemulidae 0.16  Marshall Islands  65 Siganidae 0.15 Antigua and Barbuda 3 Scaridae 0.15  Marshall Islands  65 Serranidae 0.13 Antigua and Barbuda 3 Acanthuridae 0.14  Marshall Islands  65 Scaridae 0.13 Bahamas 153 Lutjanidae 0.62  Marshall Islands  66 Serranidae 0.34 Bahamas 153 Serranidae 0.17  Marshall Islands  66 Lutjanidae 0.14 Bahamas 153 Haemulidae 0.10  Marshall Islands  66 Acanthuridae 0.12 Bahamas 153 Balistidae 0.05  Marshall Islands  66 Siganidae 0.10 Bahamas 153 Carangidae 0.03  Marshall Islands  66 Lethrinidae 0.09 Bahamas 153 Sphryaenidae 0.03  Marshall Islands  67 Serranidae 0.30 Barbados 4 Scaridae 0.24  Marshall Islands  67 Lutjanidae 0.20 Barbados 4 Monacanthidae 0.12  Marshall Islands  67 Lethrinidae 0.12 Barbados 4 Acanthuridae 0.12  Marshall Islands  67 Acanthuridae 0.08 Barbados 4 Serranidae 0.11  Marshall Islands  67 Kyphosidae 0.07 Barbados 4 Holocentridae 0.10  Martinique 68 Muraenidae 0.44 Barbados 5 Mullidae 0.15  Martinique 68 Squalidae 0.18 Barbados 5 Ostraciidae 0.10  Martinique 68 Carangidae 0.15 Barbados 5 Pomacanthidae 0.09  Martinique 68 Holocentridae 0.12 Barbados 5 Pomacentridae 0.07  Martinique 68 Lutjanidae 0.11 Barbados 5 Belonidae 0.07  Martinique 69 Scaridae 0.69 Bermuda 6 Serranidae 0.25  Martinique 69 Balistidae 0.39 Bermuda 6 Carangidae 0.20  Martinique 69 Lutjanidae 0.24 Bermuda 6 Scaridae 0.15  Martinique 69 Mullidae 0.19 Bermuda 6 Lutjanidae 0.13  Martinique 69 Serranidae 0.14 Bermuda 6 Acanthuridae 0.04  Mauritius 70 Siganidae 0.28 Bermuda 6 Sparidae 0.04  Mauritius 70 Mullidae 0.21 Bermuda 6 Haemulidae 0.04  Mauritius 70 Mugilidae 0.11 Bermuda 6 Balistidae 0.04  Mauritius 70 Carangidae 0.11 Bermuda 6 Labridae 0.04  Mauritius 70 Lethrinidae 0.10 Bermuda 6 Holocentridae 0.04  Mauritius 136 Siganidae 0.52 Bermuda 6 Kyphosidae 0.04  Mauritius 136 Acanthuridae 0.17 Bermuda 7 Lutjanidae 0.41  Mauritius 136 Scaridae 0.10 Bermuda 7 Kyphosidae 0.28  Mauritius 136 Mullidae 0.08 Bermuda 7 Haemulidae 0.20  Mauritius 136 Serranidae 0.04 Bermuda 7 Sparidae 0.06  Mauritius 137 Serranidae 0.39 Bermuda 7 Holocentridae 0.04  Mauritius 137 Siganidae 0.15 British Virgin Islands 8 Serranidae 0.17  Mauritius 137 Sphyraenidae 0.12 British Virgin Islands 8 Acanthuridae 0.16  Mauritius 137 Mullidae 0.10 British Virgin Islands 8 Balistidae 0.12  Mauritius 137 Acanthuridae 0.08 British Virgin Islands 8 Haemulidae 0.11  Mauritius 138 Signidae 0.40 British Virgin Islands 8 Ostraciidae 0.11  Mauritius 138 Acanthuridae 0.13 Comoros 9 Serranidae 0.18  Mauritius 138 Chanidae 0.10 Comoros 9 Scombridae 0.03  Mauritius 138 Carangidae 0.08 Comoros 9 Carangidae 0.02  Mauritius 138 Mullidae 0.07 Comoros 9 Lutjanidae 0.02  Mayotte 71 Scombridae 0.00 Comoros 9 Lethrinidae 0.01  Mayotte 71 Lutjanidae 0.00 Cook Islands1 10 Kyphosidae 0.39  Mayotte 71 Serranidae 0.00 Cook Islands1 10 Acanthuridae 0.22  Mayotte 71 Sparidae 0.00 Cook Islands1 10 Scaridae 0.12  Mayotte 71 Carangidae 0.00 Cook Islands1 10 Mullidae 0.08  Nauru 72 Acanthuridae 0.37 Cook Islands1 10 Serranidae 0.06  Nauru 72 Holocentridae 0.14 Cook Islands2 11 Serranidae 0.29  Nauru 72 Lutjanidae 0.11 Cook Islands2 11 Kyphosidae 0.26  Nauru 72 Scaridae 0.09 Cook Islands2 11 Holocentridae 0.13  Nauru 72 Serranidae 0.09 Cook Islands2 11 Labridae 0.12  NetherlandAntilles 73 Lutjanidae 0.46 Cook Islands 11 Acanthuridae 0.12  NetherlandAntilles 73 Haemulidae 0.15   162 Country Case No. Family % catch  Country Case No. Family % catch Cook Islands 12 Scaridae 0.75  NetherlandAntilles 73 Serranidae 0.15 Cook Islands 12 Serranidae 0.08  NetherlandAntilles 73 Balistidae 0.10 Cook Islands 12 Lutjanidae 0.07  NetherlandAntilles 73 Scombridae 0.07 Cook Islands 12 Mugilidae 0.04  NetherlandAntilles 143 Scaridae 0.40 Cook Islands 12 Holocentridae 0.03  NetherlandAntilles 143 Haemulidae 0.20 Cuba 13 Lutjanidae 0.62  NetherlandAntilles 143 Acanthuridae 0.13 Cuba 13 Carangidae 0.18  NetherlandAntilles 143 Lutjanidae 0.12 Cuba 13 Sphyraenidae 0.06  NetherlandAntilles 143 Chaetodontidae 0.04 Cuba 13 Centropomidae 0.04  New Caledonia 74 Scaridae 0.23 Cuba 13 Elopidae 0.02  New Caledonia 74 Lethrinidae 0.20 Dominica 14 Monacanthidae 0.25  New Caledonia 74 Acanthuridae 0.17 Dominica 14 Serranidae 0.12  New Caledonia 74 Mullidae 0.13 Dominica 14 Dactylopteridae 0.10  New Caledonia 74 Serranidae 0.12 Dominica 14 Holocentridae 0.10  New Caledonia 75 Mugilidae 0.49 Dominica 14 Balistidae 0.09  New Caledonia 75 Lethrinidae 0.20 Dominica 15 Muraenidae 0.16  New Caledonia 75 Siganidae 0.11 Dominica 15 Lutjanidae 0.15  New Caledonia 75 Mullidae 0.08 Dominica 15 Mullidae 0.13  New Caledonia 75 Serranidae 0.03 Dominica 15 Scaridae 0.09  New Caledonia 76 Scaridae 0.27 Dominica 15 Serranidae 0.08  New Caledonia 76 Acanthuridae 0.25 East Timor 16 Lutjanidae 0.10  New Caledonia 76 Siganidae 0.20 East Timor 16 Caesionidae 0.10  New Caledonia 76 Serranidae 0.14 East Timor 16 Acanthuridae 0.07  New Caledonia 76 Lethrinidae 0.10 East Timor 16 Scombridae 0.06  New Caledonia 77 Mugilidae 0.25 East Timor 16 Hemiramphidae 0.04  New Caledonia 77 Lethrinidae 0.21 East Timor 16 Clupeidae 0.04  New Caledonia 77 Siganidae 0.20 East Timor 16 Belonidae 0.04  New Caledonia 77 Acanthuridae 0.07 FSM1  27 Scaridae 0.36  New Caledonia 77 Hemiramphidae 0.06 FSM 27 Acanthuridae 0.27  New Caledonia 77 Gerreidae 0.06 FSM 27 Serranidae 0.18  New Caledonia 78 Lethrinidae 0.24 FSM 27 Lethrinidae 0.10  New Caledonia 78 Serranidae 0.22 FSM 27 Mullidae 0.03  New Caledonia 78 Mugilidae 0.19 FSM 28 Scaridae 0.52  New Caledonia 78 Acanthuridae 0.11 FSM 28 Serranidae 0.28  New Caledonia 78 Hemiramphidae 0.06 FSM 28 Acanthuridae 0.19  New Caledonia 78 Scaridae 0.06 FSM 29 Acanthuridae 0.44  Niue 79 Kyphosidae 0.24 FSM 29 Scaridae 0.21  Niue 79 Carangidae 0.22 FSM 29 Siganidae 0.13  Niue 79 Holocentridae 0.21 FSM 29 Carangidae 0.08  Niue 79 Cirrhitidae 0.09 FSM 29 Lethrinidae 0.04  Niue 79 Serranidae 0.07 FSM 30 Acanthuridae 0.34  Palau 80 Lethrinidae 0.33 FSM 30 Scaridae 0.23  Palau 80 Siganidae 0.21 FSM 30 Siganidae 0.17  Palau 80 Scaridae 0.14 FSM 30 Carangidae 0.14  Palau 80 Lutjanidae 0.10 FSM 30 Gerreidae 0.11  Palau 80 Mugilidae 0.10 FSM 31 Carangidae 0.47  Palau 81 Lethrinidae 0.28 FSM 31 Siganidae 0.15  Palau 81 Lutjanidae 0.17 FSM 31 Sphyraenidae 0.12  Palau 81 Scaridae 0.13 FSM 31 Scaridae 0.11  Palau 81 Serranidae 0.10 FSM 31 Acanthuridae 0.08  Palau 81 Acanthuridae 0.09 FSM 84 Acanthuridae 0.28  Palau 82 Lethrinidae 0.32 FSM 84 Scaridae 0.15  Palau 82 Siganidae 0.19 FSM 84 Serranidae 0.15  Palau 82 Serranidae 0.15 FSM 84 Lutjanidae 0.10  Palau 82 Lutjanidae 0.10 FSM 84 Carangidae 0.09  Palau 82 Acanthuridae 0.07 FSM 155 Scaridae 0.57  Papua New Guinea 123 Lutjanidae 0.28 FSM 155 Acanthuridae 0.30  Papua New Guinea 123 Lethrinidae 0.26 FSM 155 Lethrinidae 0.08  Papua New Guinea 123 Carangidae 0.11 FSM 155 Kyphosidae 0.05  Papua New Guinea 123 Serranidae 0.07 FSM 156 Acanthuridae 0.36  Papua New Guinea 123 Balistidae 0.05 FSM 156 Scaridae 0.27  Papua New Guinea 124 Lethrinidae 0.23 FSM 156 Carangidae 0.14  Papua New Guinea 124 Mugilidae 0.14 FSM 156 Kyphosidae 0.11  Papua New Guinea 124 Gerreidae 0.10 FSM 156 Lethrinidae 0.07  Papua New Guinea 124 Lutjanidae 0.09   163 Country Case No. Family % catch  Country Case No. Family % catch FSM 157 Scaridae 0.42  Papua New Guinea 124 Acanthuridae 0.09 FSM 157 Acanthuridae 0.30  Papua New Guinea 125 Siganidae 0.30 FSM 157 Kyphosidae 0.07  Papua New Guinea 125 Acanthuridae 0.29 FSM 157 Lethrinidae 0.06  Papua New Guinea 125 Lethrinidae 0.11 FSM 157 Mullidae 0.05  Papua New Guinea 125 Scaridae 0.08 Fiji 17 Lethrinidae 0.21  Papua New Guinea 125 Clupeidae 0.05 Fiji 17 Mugilidae  0.14  Philippines 83 Scombridae  0.63 Fiji 17 Lutjanidae  0.10  Philippines 83 Serranidae  0.14 Fiji 17 Leiognathidae  0.08  Philippines 83 Lethrinidae  0.01 Fiji 17 Balistidae  0.07  Philippines 83 Scaridae  0.01 Fiji 19 Lethrinidae 0.30  Philippines 83 Lutjanidae  0.01 Fiji 19 Mugilidae  0.17  Philippines 130 Carangidae  0.47 Fiji 19 Acanthuridae 0.11  Philippines 130 Acanthuridae  0.25 Fiji 19 Scombridae  0.08  Philippines 130 Scombridae  0.13 Fiji 19 Lutjanidae  0.07  Philippines 130 Scaridae  0.07 Fiji 20 Lethrinidae 0.46  Philippines 130 Caesionidae  0.04 Fiji 20 Serranidae  0.12  Philippines 131 Acanthuridae  0.23 Fiji 20 Carangidae  0.10  Philippines 131 Belonidae 0.21 Fiji 20 Acanthuridae 0.09  Philippines 131 Caesionidae  0.19 Fiji 20 Scaridae  0.04  Philippines 131 Exocoetidae 0.10 Fiji 21 Lethrinidae 0.35  Philippines 131 Scombridae  0.08 Fiji 21 Serranidae  0.14  Philippines 144 Carangidae 0.29 Fiji 21 Haemulidae  0.13  Philippines 144 Scombridae 0.29 Fiji 21 Acanthuridae 0.12  Philippines 144 Acanthuridae 0.19 Fiji 21 Carangidae  0.08  Philippines 144 Caesionidae  0.17 Fiji 22 Lethrinidae 0.57  Philippines 144 Scaridae 0.03 Fiji 22 Gerreidae  0.11  Philippines 145 Scombridae 0.32 Fiji 22 Hemiramphidae  0.08  Philippines 145 Acanthuridae 0.26 Fiji 22 Lutjanidae  0.08  Philippines 145 Carangidae 0.25 Fiji 22 Scaridae  0.07  Philippines 145 Caesionidae  0.10 Fiji 23 Lethrinidae 0.26  Philippines 145 Lutjanidae 0.03 Fiji 23 Lutjanidae  0.16  Philippines 146 Carangidae 0.41 Fiji 23 Serranidae  0.11  Philippines 146 Acanthuridae 0.25 Fiji 23 Mugilidae  0.09  Philippines 146 Scombridae 0.22 Fiji 23 Acanthuridae 0.06  Philippines 146 Lutjanidae 0.06 Fiji 132 Lethrinidae 0.36  Philippines 146 Caesionidae  0.04 Fiji 132 Acanthuridae 0.16  Philippines 147 Scombridae 0.35 Fiji 132 Serranidae 0.12  Philippines 147 Carangidae 0.30 Fiji 132 Carangidae 0.10  Philippines 147 Acanthuridae 0.28 Fiji 132 Balistidae 0.06  Philippines 147 Lutjanidae 0.04 Fiji 133 Lethrinidae 0.39  Philippines 147 Caesionidae  0.02 Fiji 133 Serranidae 0.19  Philippines 148 Carangidae 0.48 Fiji 133 Others 0.09  Philippines 148 Acanthuridae 0.30 Fiji 133 Acanthuridae 0.09  Philippines 148 Caesionidae  0.09 Fiji 133 Lutjanidae 0.07  Philippines 148 Scombridae 0.05 Fiji 134 Serranidae 0.24  Philippines 148 Scaridae 0.04 Fiji 134 Lethrinidae 0.22  Philippines 149 Carangidae 0.48 Fiji 134 Acanthuridae 0.11  Philippines 149 Acanthuridae 0.25 Fiji 134 Others 0.08  Philippines 149 Scombridae 0.13 Fiji 134 Lutjanidae 0.07  Philippines 149 Scaridae 0.07 Fiji 135 Lethrinidae 0.17  Philippines 149 Caesionidae  0.04 Fiji 135 Others 0.16  Philippines 154 Siganidae 0.48 Fiji 135 Serranidae 0.14  Philippines 154 Scaridae 0.19 Fiji 135 Scombridae 0.12  Philippines 154 Labridae 0.11 Fiji 135 Acanthuridae 0.11  Philippines 154 Gobiidae 0.05 French Polynesia 24 Acanthuridae 0.34  Philippines 154 Lethrinidae 0.03 French Polynesia 24 Scaridae 0.26  Philippines 154 Muraenidae 0.03 French Polynesia 24 Serranidae 0.12  Puerto Rico 85 Lutjanidae 0.29 French Polynesia 24 Holocentridae 0.08  Puerto Rico 85 Holocentridae 0.16 French Polynesia 24 Lethrinidae 0.07  Puerto Rico 85 Serranidae 0.13 French Polynesia 25 Acanthuridae 0.22  Puerto Rico 85 Ostraciidae 0.08 French Polynesia 25 Carangidae 0.15  Puerto Rico 85 Mullidae 0.05 French Polynesia 25 Mullidae 0.15  Reunion 86 Lethrinidae 0.67 French Polynesia 25 Serranidae 0.12  Reunion 86 Scaridae 0.22   164 Country Case No. Family % catch  Country Case No. Family % catch French Polynesia 25 Siganidae 0.11  Reunion 86 Acanthuridae 0.14 French Polynesia 26 Carangidae 0.44  Reunion 86 Muraenidae 0.07 French Polynesia 26 Priacanthidae 0.18  Reunion 86 Haemulidae 0.07 French Polynesia 26 Lutjanidae 0.07  Samoa 1 Balistidae 0.13 French Polynesia 26 Acanthuridae 0.07  Samoa 1 Mullidae 0.13 French Polynesia 26 Lethrinidae 0.07  Samoa 1 Scaridae 0.10 Grenada 32 Serranidae 0.82  Samoa 1 Mugilidae 0.08 Grenada 32 Lutjanidae 0.16  Samoa 1 Muraenidae 0.08 Grenada 32 Balistidae 0.00  Samoa 1 Sphyraenidae 0.08 Grenada 32 Malacanthidae 0.00  Samoa 87 Scaridae 0.20 Guadeloupe 33 Dactylopteridae 0.17  Samoa 87 Lethrinidae 0.18 Guadeloupe 33 Muraenidae 0.15  Samoa 87 Acanthuridae 0.14 Guadeloupe 33 Mullidae 0.13  Samoa 87 Holocentridae 0.07 Guadeloupe 33 Haemulidae 0.12  Samoa 87 Lutjanidae 0.06 Guadeloupe 33 Holocentridae 0.12  Samoa 88 Acanthuridae 0.24 Guadeloupe 34 Mullidae 0.22  Samoa 88 Scaridae 0.15 Guadeloupe 34 Muraenidae 0.17  Samoa 88 Lutjanidae 0.09 Guadeloupe 34 Lutjanidae 0.16  Samoa 88 Lethrinidae 0.09 Guadeloupe 34 Carangidae 0.10  Samoa 88 Mugilidae 0.08 Guadeloupe 34 Holocentridae 0.09  Samoa 89 Lethrinidae 0.19 Guam 35 Scaridae 0.41  Samoa 89 Acanthuridae 0.16 Guam 35 Acanthuridae 0.22  Samoa 89 Scaridae 0.15 Guam 35 Labridae 0.08  Samoa 89 Mugilidae 0.12 Guam 35 Serranidae 0.07  Samoa 89 Siganidae 0.10 Guam 35 Kyphosidae 0.06  Samoa 90 Acanthuridae 0.24 Indonesia 36 Lethrinidae 0.34  Samoa 90 Scaridae 0.18 Indonesia 36 Mullidae 0.16  Samoa 90 Lethrinidae 0.11 Indonesia 36 Siganidae 0.12  Samoa 90 Holocentridae 0.11 Indonesia 36 Scaridae 0.12  Samoa 90 Mugilidae 0.09 Indonesia 36 Clupeidae 0.09  Samoa 91 Acanthuridae 0.21 Indonesia 37 Lethrinidae 0.68  Samoa 91 Lethrinidae 0.17 Indonesia 37 Labridae 0.08  Samoa 91 Scaridae 0.14 Indonesia 37 Serranidae 0.08  Samoa 91 Siganidae 0.12 Indonesia 37 Siganidae 0.04  Samoa 91 Serranidae 0.10 Indonesia 37 Gerreidae 0.03  Samoa 92 Acanthuridae 0.28 Indonesia 38 Lethrinidae 0.27  Samoa 92 Scaridae 0.19 Indonesia 38 Serranidae 0.27  Samoa 92 Lethrinidae 0.14 Indonesia 38 Lutjanidae 0.20  Samoa 92 Holocentridae 0.08 Indonesia 38 Carangidae 0.07  Samoa 92 Lutjanidae 0.06 Indonesia 38 Holocentridae 0.07  Samoa 93 Acanthuridae 0.29 Indonesia 39 Scaridae 0.33  Samoa 93 Scaridae 0.17 Indonesia 39 Mullidae 0.31  Samoa 93 Lethrinidae 0.14 Indonesia 39 Labridae 0.07  Samoa 93 Holocentridae 0.14 Indonesia 39 Acanthuridae 0.06  Samoa 93 Muraenidae 0.09 Indonesia 39 Lethrinidae 0.06  Samoa 94 Scaridae 0.49 Indonesia 40 Lethrinidae 0.33  Samoa 94 Acanthuridae 0.19 Indonesia 40 Siganidae 0.21  Samoa 94 Lutjanidae 0.15 Indonesia 40 Labridae 0.12  Samoa 94 Lethrinidae 0.12 Indonesia 40 Scaridae 0.08  Samoa 94 Labridae 0.05 Indonesia 40 Mullidae 0.08  Seychelles 95 Serranidae 0.56 Indonesia 41 Lethrinidae 0.18  Seychelles 95 Lutjanidae 0.24 Indonesia 41 Siganidae 0.17  Seychelles 95 Lethrinidae 0.20 Indonesia 41 Gerreidae 0.16  Solomon Islands 96 Scaridae 0.27 Indonesia 41 Lutjanidae 0.08  Solomon Islands 96 Serranidae 0.15 Indonesia 41 Mullidae 0.07  Solomon Islands 96 Labridae 0.11 Indonesia 42 Hemiramphidae 0.84  Solomon Islands 96 Lutjanidae 0.11 Indonesia 42 Clupeidae 0.09  Solomon Islands 96 Balistidae 0.11 Indonesia 42 Labridae 0.03  Solomon Islands 97 Lutjanidae 0.18 Indonesia 42 Mullidae 0.02  Solomon Islands 97 Carangidae 0.12 Indonesia 42 Nemipteridae 0.02  Solomon Islands 97 Scaridae 0.11 Indonesia 43 Caesionidae 0.66  Solomon Islands 97 Serranidae 0.10 Indonesia 43 Acanthuridae 0.09  Solomon Islands 97 Haemulidae 0.10 Indonesia 43 Serranidae 0.04  Solomon Islands 98 Serranidae 0.25 Indonesia 43 Lutjanidae 0.04  Solomon Islands 98 Carangidae 0.23   165 Country Case No. Family % catch  Country Case No. Family % catch Indonesia 43 Siganidae 0.04  Solomon Islands 98 Holocentridae 0.11 Jamaica 44 Carangidae 0.32  Solomon Islands 98 Scombridae 0.09 Jamaica 44 Serranidae 0.21  Solomon Islands 98 Lethrinidae 0.08 Jamaica 44 Lutjanidae 0.17  Solomon Islands 99 Serranidae 0.25 Jamaica 44 Holocentridae 0.09  Solomon Islands 99 Holocentridae 0.13 Jamaica 44 Sphyraenidae 0.07  Solomon Islands 99 Lutjanidae 0.10 Jamaica 45 Carangidae 0.44  Solomon Islands 99 Carangidae 0.09 Jamaica 45 Haemulidae 0.22  Solomon Islands 99 Lethrinidae 0.06 Jamaica 45 Scaridae 0.14  Solomon Islands 100 Lethrinidae 0.24 Jamaica 45 Lutjanidae 0.11  Solomon Islands 100 Lutjanidae 0.20 Jamaica 45 Acanthuridae 0.04  Solomon Islands 100 Serranidae 0.11 Jamaica 45 Sphyraenidae 0.04  Solomon Islands 100 Scaridae 0.08 Jamaica 46 Sphyraenidae 0.35  Solomon Islands 100 Mugilidae 0.07 Jamaica 46 Scaridae 0.35  Sri Lanka 101 Scaridae 0.30 Jamaica 46 Serranidae 0.08  Sri Lanka 101 Serranidae 0.23 Jamaica 46 Mullidae 0.06  Sri Lanka 101 Monodactylidae 0.15 Jamaica 46 Acanthuridae 0.06  Sri Lanka 101 Labridae 0.15 Jamaica 47 Scaridae 0.42  Sri Lanka 101 Clupeidae 0.10 Jamaica 47 Acanthuridae 0.18  Sri Lanka 102 Mugilidae 0.30 Jamaica 47 Lutjanidae 0.10  Sri Lanka 102 Carangidae 0.30 Jamaica 47 Holocentridae 0.06  Sri Lanka 102 Serranidae 0.10 Jamaica 47 Haemulidae 0.05  Sri Lanka 102 Scaridae 0.10 Jamaica 48 Sphyraenidae 0.74  Sri Lanka 102 Monodactylidae 0.05 Jamaica 48 Scombridae 0.16  Sri Lanka 102 Scrombridae 0.05 Jamaica 48 Belonidae 0.10  Sri Lanka 102 Sphyraenidae 0.05 Jamaica 150 Scaridae 0.42  Sri Lanka 102 Haemulidae 0.05 Jamaica 150 Acanthuridae 0.15  Tonga 103 Lethrinidae 0.67 Jamaica 150 Holocentridae 0.10  Tonga 103 Scaridae 0.07 Jamaica 150 Pomadasyidae  0.06  Tonga 103 Carangidae 0.07 Jamaica 150 Serranidae 0.06  Tonga 103 Gerreidae 0.04 Jamaica 151 Scaridae 0.48  Tonga 103 Lutjanidae 0.04 Jamaica 151 Acanthuridae 0.15  Tonga 104 Acanthuridae 0.22 Jamaica 151 Holocentridae 0.08  Tonga 104 Lethrinidae 0.19 Jamaica 151 Pomadasyidae 0.06  Tonga 104 Scaridae 0.18 Jamaica 151 Serranidae 0.05  Tonga 104 Serranidae 0.16 Jamaica 152 Scaridae 0.34  Tonga 104 Siganidae 0.09 Jamaica 152 Sphyraenidae 0.15  Tonga 105 Lethrinidae 0.30 Jamaica 152 Acanthuridae 0.11  Tonga 105 Serranidae 0.19 Jamaica 152 Lutjanidae 0.07  Tonga 105 Acanthuridae 0.15 Jamaica 152 Serranidae 0.07  Tonga 105 Scaridae 0.13 Jamaica 158 Scaridae 0.41  Tonga 105 Lutjanidae 0.09 Jamaica 158 Mullidae 0.18  Tonga 106 Lethrinidae 0.69 Jamaica 158 Holocentridae 0.09  Tonga 106 Lutjanidae 0.14 Jamaica 158 Acanthuridae 0.08  Tonga 106 Serranidae 0.14 Jamaica 158 Serranidae 0.08  Tonga 106 Carangidae 0.03 Kenya 49 Lethrinidae 0.63  Tonga 106 Holocentridae 0.00 Kenya 49 Labridae 0.14  Tonga 107 Acanthuridae 0.39 Kenya 49 Lutjanidae 0.11  Tonga 107 Scaridae 0.24 Kenya 49 Mullidae 0.02  Tonga 107 Siganidae 0.19 Kenya 49 Scaridae 0.01  Tonga 107 Lethrinidae 0.09 Kenya 50 Siganidae 0.36  Tonga 107 Holocentridae 0.03 Kenya 50 Lethrinidae 0.27  Trinidad & Tobago 120 Lutjanidae 0.95 Kenya 50 Scaridae 0.24  Trinidad & Tobago 120 Haemulidae 0.03 Kenya 50 Mullidae 0.06  Trinidad & Tobago 120 Serranidae 0.02 Kenya 50 Acanthuridae 0.04  Trinidad & Tobago 121 Lutjanidae 0.87 Kenya 159 Scaridae 0.28  Trinidad & Tobago 121 Balistidae 0.05 Kenya 159 Others 0.23  Trinidad & Tobago 121 Haemulidae 0.05 Kenya 159 Scavengers 0.20  Trinidad & Tobago 121 Serranidae 0.03 Kenya 159 Scaridae 0.18  Trinidad & Tobago 122 Lutjanidae 0.66 Kenya 159 Mullidae 0.10  Trinidad & Tobago 122 Sparidae 0.10 Kiribati 51 Lethrinidae 0.35  Trinidad & Tobago 122 Balistidae 0.09 Kiribati 51 Mullidae 0.11  Trinidad & Tobago 122 Haemulidae 0.09 Kiribati 51 Gerreidae 0.08  Trinidad & Tobago 122 Holocentridae 0.03 Kiribati 51 Belonidae 0.07  Turks & Caicos 108 Haemulidae 0.38   166 Country Case No. Family % catch  Country Case No. Family % catch Kiribati 51 Lutjanidae 0.07  Turks & Caicos 108 Lutjanidae 0.26 Kiribati 52 Serranidae 0.41  Turks & Caicos 108 Scaridae 0.24 Kiribati 52 Scaridae 0.15  Turks & Caicos 108 Serranidae 0.06 Kiribati 52 Holocentridae 0.10  Turks & Caicos 108 Mullidae 0.03 Kiribati 52 Labridae 0.09  Turks & Caicos 139 Serranidae 0.33 Kiribati 52 Lethrinidae 0.07  Turks & Caicos 139 Lutjanidae 0.26 Kiribati 53 Mugilidae 0.34  Turks & Caicos 139 Haemulidae 0.16 Kiribati 53 Albulidae 0.28  Turks & Caicos 139 Labridae 0.09 Kiribati 53 Lethrinidae 0.10  Turks & Caicos 139 Carangidae 0.07 Kiribati 53 Mullidae 0.10  Turks & Caicos 140 Lutjanidae 0.50 Kiribati 53 Gerreidae 0.10  Turks & Caicos 140 Haemulidae 0.39 Kiribati 54 Chanidae 0.65  Turks & Caicos 140 Serranidae 0.05 Kiribati 54 Albulidae 0.17  Turks & Caicos 140 Carangidae 0.05 Kiribati 54 Serranidae 0.07  Turks & Caicos 141 Serranidae 0.72 Kiribati 54 Mugilidae 0.05  Turks & Caicos 141 Haemulidae 0.21 Kiribati 54 Acanthuridae 0.03  Turks & Caicos 141 Lutjanidae 0.03 Kiribati 55 Lutjanidae 0.43  Turks & Caicos 141 Scaridae 0.01 Kiribati 55 Carangidae 0.40  Turks & Caicos 141 Holocentridae 0.01 Kiribati 55 Serranidae 0.09  Turks & Caicos 142 Serranidae 0.33 Kiribati 55 Lethrinidae 0.08  Turks & Caicos 142 Lutjanidae 0.25 Madagascar 56 Carangidae 0.39  Turks & Caicos 142 Haemulidae 0.17 Madagascar 56 Sharks 0.21  Turks & Caicos 142 Carangidae 0.09 Madagascar 56 Lutjanidae 0.12  Turks & Caicos 142 Labridae 0.07 Madagascar 56 Gerreidae 0.08  Tuvalu 109 Lethrinidae 0.26 Madagascar 56 Sphyraenidae 0.07  Tuvalu 109 Serranidae 0.25 Madagascar 57 Haemulidae 0.33  Tuvalu 109 Lutjanidae 0.24 Madagascar 57 Serranidae 0.22  Tuvalu 109 Acanthuridae 0.14 Madagascar 57 Scombridae 0.21  Tuvalu 109 Kyphosidae 0.04 Madagascar 57 Nemipteridae 0.14  Tuvalu 110 Acanthuridae 0.27 Madagascar 57 Isiophoridae 0.10  Tuvalu 110 Kyphosidae 0.16 Madagascar 57 Lethrinidae 0.10  Tuvalu 110 Lethrinidae 0.15 Madagascar 58 Scombridae 0.09  Tuvalu 110 Mugilidae 0.10 Madagascar 58 Carangidae 0.06  Tuvalu 110 Lutjanidae 0.08 Madagascar 58 Mugilidae 0.05  Tuvalu 110 Serranidae 0.08 Madagascar 58 Scaridae 0.05  Tuvalu 111 Mugilidae 0.52 Madagascar 58 Gerreidae 0.04  Tuvalu 111 Kyphosidae 0.12 Madagascar 58 Clupeidae 0.04  Tuvalu 111 Lutjanidae 0.09 Madagascar 59 Clupeidae 0.15  Tuvalu 111 Gerreidae 0.07 Madagascar 59 Siganidae 0.14  Tuvalu 111 Acanthuridae 0.06 Madagascar 59 Carangidae 0.13  Tuvalu 112 Lutjanidae 0.14 Madagascar 59 Belonidae 0.13  Tuvalu 112 Serranidae 0.14 Madagascar 59 Gerreidae 0.12  Tuvalu 112 Acanthuridae 0.13 Madagascar 60 Lutjanidae 0.34  Tuvalu 112 Carangidae 0.11 Madagascar 60 Labridae 0.34  Tuvalu 112 Exocoetidae 0.10 Madagascar 60 Lethrinidae 0.10  U.S. Virgin Islands 113 Scaridae 0.28 Madagascar 60 Serranidae 0.08  U.S. Virgin Islands 113 Balistidae 0.19 Madagascar 60 Sparidae 0.07  U.S. Virgin Islands 113 Serranidae 0.17 Madagascar 126 Lethrinidae 0.34  U.S. Virgin Islands 113 Acanthuridae 0.13 Madagascar 126 Siganidae 0.23  U.S. Virgin Islands 113 Holocetridae 0.09 Madagascar 126 Labridae 0.10  Vanuatu 114 Lethrinidae 0.36 Madagascar 126 Acanthuridae 0.08  Vanuatu 114 Siganidae 0.23 Madagascar 126 Gerreidae 0.04  Vanuatu 114 Mullidae 0.17 Madagascar 127 Clupeidae 0.80  Vanuatu 114 Scaridae 0.06 Madagascar 127 Pomacentridae 0.04  Vanuatu 114 Serranidae 0.04 Madagascar 127 Scaridae 0.02  Vanuatu 115 Scaridae 0.48 Madagascar 127 Mullidae 0.02  Vanuatu 115 Acanthuridae 0.29 Madagascar 127 Atherinidae 0.01  Vanuatu 115 Balistidae 0.04 Madagascar 128 Acanthuridae 0.28  Vanuatu 115 Carangidae 0.04 Madagascar 128 Mullidae 0.15  Vanuatu 115 Mullidae 0.03 Madagascar 128 Muraenidae 0.13  Vanuatu 116 Siganidae 0.24 Madagascar 128 Tetraodontidae 0.06  Vanuatu 116 Mugilidae 0.17 Madagascar 128 Balistidae 0.06  Vanuatu 116 Acanthuridae 0.12 Madagascar 129 Scaridae 0.41  Vanuatu 116 Carangidae 0.11 Madagascar 129 Acanthuridae 0.23  Vanuatu 116 Albulidae 0.10   167 Country Case No. Family % catch  Country Case No. Family % catch Madagascar 129 Haemulidae 0.06  Wallis and Futuna 117 Acanthuridae 0.33 Madagascar 129 Holocentridae 0.05  Wallis and Futuna 117 Mugilidae 0.16 Madagascar 129 Siganidae 0.05  Wallis and Futuna 117 Carangidae 0.11 Malaysia  61 Scombridae 0.29  Wallis and Futuna 117 Holocentridae 0.08 Malaysia  61 Carangidae 0.21  Wallis and Futuna 117 Serranidae 0.07 Malaysia  61 Lethrinidae 0.15  Wallis and Futuna 118 Lethrinidae 0.24 Malaysia  61 Serranidae 0.14  Wallis and Futuna 118 Carangidae 0.18 Malaysia  61 Lutjanidae 0.05  Wallis and Futuna 118 Acanthuridae 0.18 Malaysia  62 Serranidae 0.37  Wallis and Futuna 118 Lutjanidae 0.16 Malaysia  62 Lethrinidae 0.16  Wallis and Futuna 118 Scaridae 0.07 Malaysia  62 Nemipteridae 0.10  Wallis and Futuna 119 Acanthuridae 0.33 Malaysia  62 Lutjanidae 0.08  Wallis and Futuna 119 Mugilidae 0.19 Malaysia  62 Carangidae 0.05  Wallis and Futuna 119 Carangidae 0.14 Maldives 63 Holocentridae 0.26  Wallis and Futuna 119 Lethrinidae 0.08 Maldives 63 Lutjanidae 0.22  Wallis and Futuna 119 Scaridae 0.07 Maldives 63 Scombridae 0.15 Maldives 63 Lethrinidae 0.12 Maldives 63 Serranidae 0.07 1 Federated States of Micronesia      


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